Project 496: Williamson, T. E., A. Weil, and B. Standhardt. 2011. Cimolestids (Mammalia) from the Early Paleocene (Puercan) of New Mexico. Journal of Vertebrate Paleontology. 31 (1):162-180.
Abstract
Cimolestids are a minor component of early Paleocene (Puercan) faunas of western North America. Here we report on cimolestids from the early Paleocene (middle and late Puercan; Pu2–3) of the Nacimiento Formation, San Juan Basin, New Mexico. New specimens of Puercolestes simpsoni include portions of the deciduous and lower dentition not previously known. We conclude that “Genus B” of Van Valen is probably conspecific with this taxon. Two new genera and species, Chacopterygus minutus and Betonnia tsosia, are from the Pu2 and Pu2–Pu3, respectively. One additional unnamed taxon is based on isolated teeth from Pu2–3 localities. A phylogenetic analysis indicates that Puercolestes simpsoni is sister to Cimolestes. Chacopterygus minutus, Betonnia tsosia, and the unnamed cimolestid form a polytomy that is sister to Procerberus. This resulting clade is sister to Batodon + Maelestes. The results of this analysis suggest that P. simpsoni is descended from Cretaceous taxa of western North America. The origins of other early Paleocene cimolestids from either Asia or North America is equivocal. The species diversity of Cimolestidae in the Puercan of New Mexico and other Puercan localities of western North America is similar to that of latest Cretaceous (Lancian) faunas of the northern Rocky Mountain Region.Read the article »
| This project contains | Matrices |
|---|---|
| Total scored cells: 16213 Total media associated with cells: 0 Total labels associated with cell media: 0 |
| Characters | |
| Total characters: 408 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 1019 Total character states with associated media: 0 Total character states with media with labels:0 |
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MorphoBank Project 496
MorphoBank Project 496
- Creation Date:
05 April 2011 - Publication Date:
16 April 2011 - Project views: 3329

- Project downloads: 43

Members
| member name | taxa |
specimens |
media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||||||||
| Natalie Milbrodt Project Administrator Last logged in 06/27/11 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | 0 | 0 | |||||||
| Maureen Admin Full membership Last logged in 04/15/11 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | 0 | 0 | |||||||
| Thomas Williamson Full membership Last logged in 04/06/11 | 74 | 0 | 0 | 408 | 0 | 0 | 0 | 16213 (16212, 0, 1) | 0 | 0 | 0 | 0 | 0 | |||||||
Taxonomic Overview for Matrix 'M1106' (74 Taxa)
| taxon | unscored cells |
scored cells |
cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
| [1] †Nanolestes Last Modified in 04/15/11 | 296 | 113 | 0 | 0 | 0 | 0 | 0 | 3 |
| [2] †Peramus Last Modified in 04/15/11 | 285 | 127 | 0 | 0 | 0 | 0 | 0 | 3 |
| [3] †Vincelestes Last Modified in 04/15/11 | 37 | 372 | 0 | 0 | 0 | 0 | 0 | 3 |
| [4] †Kielantherium Last Modified in 04/15/11 | 332 | 78 | 0 | 0 | 0 | 0 | 0 | 3 |
| [5] †Deltatheridium Last Modified in 04/15/11 | 145 | 265 | 0 | 0 | 0 | 0 | 0 | 3 |
| [6] †Mayulestes Last Modified in 04/15/11 | 67 | 341 | 0 | 0 | 0 | 0 | 0 | 3 |
| [7] †Pucadelphys Last Modified in 04/15/11 | 42 | 366 | 0 | 0 | 0 | 0 | 0 | 3 |
| [8] †Eomaia Last Modified in 04/15/11 | 224 | 186 | 0 | 0 | 0 | 0 | 0 | 3 |
| [9] †Prokennalestes Last Modified in 04/15/11 | 220 | 194 | 0 | 0 | 0 | 0 | 0 | 3 |
| [10] †Murtoilestes Last Modified in 04/15/11 | 349 | 59 | 0 | 0 | 0 | 0 | 0 | 3 |
| [11] †Bobolestes Last Modified in 04/15/11 | 300 | 110 | 0 | 0 | 0 | 0 | 0 | 3 |
| [12] †Montanalestes Last Modified in 04/15/11 | 338 | 76 | 0 | 0 | 0 | 0 | 0 | 3 |
| [13] †Paranyctoides Last Modified in 04/15/11 | 301 | 113 | 0 | 0 | 0 | 0 | 0 | 3 |
| [14] †Eozhelestes Last Modified in 04/15/11 | 367 | 43 | 0 | 0 | 0 | 0 | 0 | 3 |
| [15] †Sheikhdzheilia Last Modified in 04/15/11 | 337 | 74 | 0 | 0 | 0 | 0 | 0 | 3 |
| [16] †Alostera Last Modified in 04/15/11 | 350 | 58 | 0 | 0 | 0 | 0 | 0 | 3 |
| [17] †Lainodon Last Modified in 04/15/11 | 385 | 23 | 0 | 0 | 0 | 0 | 0 | 3 |
| [18] †Avitotherium Last Modified in 04/15/11 | 338 | 72 | 0 | 0 | 0 | 0 | 0 | 3 |
| [19] †Gallolestes Last Modified in 04/15/11 | 365 | 44 | 0 | 0 | 0 | 0 | 0 | 3 |
| [20] †Parazhelestes Last Modified in 04/15/11 | 255 | 157 | 0 | 0 | 0 | 0 | 0 | 3 |
| [21] †Aspanlestes Last Modified in 04/15/11 | 260 | 154 | 0 | 0 | 0 | 0 | 0 | 3 |
| [22] †Zhelestes Last Modified in 04/15/11 | 244 | 171 | 0 | 0 | 0 | 0 | 0 | 3 |
| [23] †Cimolestes Last Modified in 04/15/11 | 251 | 167 | 0 | 0 | 1 | 0 | 0 | 3 |
| [24] †Maelestes Last Modified in 04/15/11 | 103 | 314 | 0 | 0 | 0 | 0 | 0 | 3 |
| [25] †Batodon Last Modified in 04/15/11 | 305 | 105 | 0 | 0 | 0 | 0 | 0 | 3 |
| [26] †Bulaklestes Last Modified in 04/15/11 | 309 | 101 | 0 | 0 | 0 | 0 | 0 | 3 |
| [27] †Daulestes Last Modified in 04/15/11 | 311 | 100 | 0 | 0 | 0 | 0 | 0 | 3 |
| [28] †Uchkudukodon Last Modified in 04/15/11 | 160 | 258 | 0 | 0 | 0 | 0 | 0 | 3 |
| [29] †Kennalestes Last Modified in 04/15/11 | 112 | 299 | 0 | 0 | 0 | 0 | 0 | 3 |
| [30] †Asioryctes Last Modified in 04/15/11 | 75 | 336 | 0 | 0 | 0 | 0 | 0 | 3 |
| [31] †Ukhaatherium Last Modified in 04/15/11 | 61 | 349 | 0 | 0 | 0 | 0 | 0 | 3 |
| [32] †Deccanolestes Last Modified in 04/15/11 | 295 | 115 | 0 | 0 | 0 | 0 | 0 | 3 |
| [33] †Kulbeckia Last Modified in 04/15/11 | 214 | 204 | 0 | 0 | 0 | 0 | 0 | 3 |
| [34] †Zhangolestes Last Modified in 04/15/11 | 357 | 52 | 0 | 0 | 0 | 0 | 0 | 3 |
| [35] †Alymlestes Last Modified in 04/15/11 | 387 | 21 | 0 | 0 | 0 | 0 | 0 | 3 |
| [36] †Zalambdalestes Last Modified in 04/15/11 | 50 | 370 | 0 | 0 | 0 | 0 | 0 | 3 |
| [37] †Barunlestes Last Modified in 04/15/11 | 127 | 283 | 0 | 0 | 0 | 0 | 0 | 3 |
| [38] †Gypsonictops Last Modified in 04/15/11 | 282 | 135 | 0 | 0 | 0 | 0 | 0 | 3 |
| [39] †Leptictis Last Modified in 04/15/11 | 21 | 388 | 0 | 0 | 0 | 0 | 0 | 3 |
| [40] †Purgatorius Last Modified in 04/15/11 | 305 | 106 | 0 | 0 | 0 | 0 | 0 | 3 |
| [41] †Protungulatum Last Modified in 04/15/11 | 202 | 214 | 0 | 0 | 0 | 0 | 0 | 3 |
| [42] †Oxyprimus Last Modified in 04/15/11 | 308 | 102 | 0 | 0 | 0 | 0 | 0 | 3 |
| [43] †Vulpavus Last Modified in 04/15/11 | 86 | 336 | 0 | 0 | 0 | 0 | 0 | 3 |
| [44] †Miacis Last Modified in 04/15/11 | 131 | 285 | 0 | 0 | 0 | 0 | 0 | 3 |
| [45] †Diacodexis Last Modified in 04/15/11 | 92 | 319 | 0 | 0 | 0 | 0 | 0 | 3 |
| [46] †Hyopsodus Last Modified in 04/15/11 | 41 | 377 | 0 | 0 | 0 | 0 | 0 | 3 |
| [47] †Meniscotherium Last Modified in 04/15/11 | 38 | 375 | 0 | 0 | 0 | 0 | 0 | 3 |
| [48] †Phenacodus Last Modified in 04/15/11 | 59 | 359 | 0 | 0 | 0 | 0 | 0 | 3 |
| [49] Ptilocercus Last Modified in 04/15/11 | 14 | 396 | 0 | 0 | 0 | 0 | 0 | 3 |
| [50] †Plesiadapis Last Modified in 04/15/11 | 52 | 373 | 0 | 0 | 0 | 0 | 0 | 3 |
| [51] †Notharctus Last Modified in 04/15/11 | 44 | 372 | 0 | 0 | 0 | 0 | 0 | 3 |
| [52] †Adapis Last Modified in 04/15/11 | 66 | 352 | 0 | 0 | 0 | 0 | 0 | 3 |
| [53] †Tribosphenomys Last Modified in 04/15/11 | 259 | 149 | 0 | 0 | 0 | 0 | 0 | 3 |
| [54] †Paramys Last Modified in 04/15/11 | 69 | 344 | 0 | 0 | 0 | 0 | 0 | 3 |
| [55] †Rhombomylus Last Modified in 04/15/11 | 61 | 354 | 0 | 0 | 0 | 0 | 0 | 3 |
| [56] †Gomphos Last Modified in 04/15/11 | 134 | 276 | 0 | 0 | 0 | 0 | 0 | 3 |
| [57] †Mimotona Last Modified in 04/15/11 | 286 | 126 | 0 | 0 | 0 | 0 | 0 | 3 |
| [58] Blarina Last Modified in 04/15/11 | 49 | 366 | 0 | 0 | 0 | 0 | 0 | 3 |
| [59] Erinaceus Last Modified in 04/15/11 | 23 | 395 | 0 | 0 | 0 | 0 | 0 | 3 |
| [60] Solenodon Last Modified in 04/15/11 | 33 | 376 | 0 | 0 | 0 | 0 | 0 | 3 |
| [61] †Eoryctes Last Modified in 04/15/11 | 205 | 208 | 0 | 0 | 0 | 0 | 0 | 3 |
| [62] Potamogale Last Modified in 04/15/11 | 32 | 376 | 0 | 0 | 0 | 0 | 0 | 3 |
| [63] Orycteropus Last Modified in 04/15/11 | 132 | 283 | 0 | 0 | 0 | 0 | 0 | 3 |
| [64] Rhynchocyon Last Modified in 04/15/11 | 40 | 370 | 0 | 0 | 0 | 0 | 0 | 3 |
| [65] Procavia Last Modified in 04/15/11 | 31 | 384 | 0 | 0 | 0 | 0 | 0 | 3 |
| [66] †Moeritherium Last Modified in 04/15/11 | 142 | 268 | 0 | 0 | 0 | 0 | 0 | 3 |
| [67] Chaetophractus Last Modified in 04/15/11 | 126 | 282 | 0 | 0 | 0 | 0 | 0 | 3 |
| [68] Bradypus Last Modified in 04/15/11 | 134 | 277 | 0 | 0 | 0 | 0 | 0 | 3 |
| [69] Tamandua Last Modified in 04/15/11 | 163 | 247 | 0 | 0 | 0 | 0 | 0 | 3 |
| [70] †Puercolestes simpsoni Last Modified in 04/06/11 | 287 | 128 | 0 | 0 | 0 | 0 | 0 | 3 |
| [71] †Chacopterygus minutus Last Modified in 04/06/11 | 359 | 50 | 0 | 0 | 0 | 0 | 0 | 3 |
| [72] †Betonnia tsosia Last Modified in 04/11/11 | 343 | 65 | 0 | 0 | 0 | 0 | 0 | 3 |
| [73] †Gen. undet. sp. undet. Last Modified in 04/15/11 | 386 | 24 | 0 | 0 | 0 | 0 | 0 | 3 |
| [74] †Procerberus Last Modified in 04/15/11 | 306 | 105 | 0 | 0 | 0 | 0 | 0 | 3 |
Project views 
| type | number of views | Individual items viewed (where applicable) |
| Total project views | 3329 | |
| Project overview | 281 | |
| Taxon list | 1536 | |
| Matrix views | 172 | Matrix landing page (159 views); Puercan New Mexico Cimolestids (13 views); |
| Media views | 601 | Media search (506 views); M68063 (95 views); |
| Views for media list | 153 | |
| Specimen list | 279 | |
| Bibliography | 141 | |
| Documents list | 166 |
Project downloads 
| type | number of downloads | Individual items downloaded (where applicable) |
| Total downloads from project | 43 | |
| Media downloads | 16 | M68063 (16 downloads); |
| Document downloads | 2 | Copyright information for exemplar media (2 downloads); |
| Project downloads | 24 | |
| Matrix downloads | 1 | Puercan New Mexico Cimolestids (1 download); |

