Project 3773: M. D. Ezcurra, S. J. Nesbitt, M. Bronzati, F. M. Dalla Vecchia, F. L. Agnolin, R. B. Benson, F. Brissón Egli, S. F. Cabreira, S. W. Evers, A. R. Gentil, R. B. Irmis, A. G. Martinelli, F. E. Novas, L. Roberto da Silva, N. D. Smith, M. R. Stocker, A. H. Turner, M. C. Langer. 2020. Enigmatic dinosaur precursors bridge the gap to the origin of Pterosauria. Nature. 588 (7838):445-449.
Abstract
Pterosaurs −the flying reptiles− were the first vertebrates to evolve powered flight1 and one of the main evolutionary radiations in Mesozoic terrestrial ecosystems (c. 252−66 Ma), but their origin has remained one of the most outstanding enigmas in palaeontology since the 19th Century2−4. They have been hypothesised as close relatives of a wide variety of reptilian clades, including dinosaur relatives2−8, and there is still a major morphological gap between those forms and the oldest, unambiguous pterosaurs from the Upper Triassic. Driven by recent discoveries of well-preserved cranial remains, μCT scans of fragile skull bones (jaws, braincases), and reliably associated postcrania, we demonstrate that lagerpetids, a group of cursorial, non-volant dinosaur precursors, are the sister group of pterosaurs, sharing numerous synapomorphies across the entire skeleton. This finding substantially shortens the temporal and morphological gap between the oldest pterosaurs and their closest relatives, simultaneously strengthening the evidence that pterosaurs belong to the avian-line of archosaurs. Neuroanatomical features related to the enhanced sensory capabilities of pterosaurs9 are already present in lagerpetids, indicating that they evolved before flight. Our new evidence illuminates the first steps of the assembly of the unique pterosaur body plan, whose conquest of aerial space represents one of the most astonishing morphofunctional innovations in vertebrate evolution.Read the article »
Article DOI: 10.1038/s41586-020-3011-4
Project DOI: 10.7934/P3773, http://dx.doi.org/10.7934/P3773
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 64847 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 823 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 1907 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:716/107 |
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MorphoBank Project 3773
MorphoBank Project 3773
- Creation Date:
19 July 2020 - Publication Date:
16 September 2020 - Project views: 16877
- Media downloads: 5
- Matrix downloads: 119
Authors' Institutions
- Humboldt-Universitaet zu Berlin (Humboldt University Berlin)
- University of Oxford
- University of Birmingham
- Université de Fribourg (University of Fribourg)
- University of Goettingen
- University of Utah
- Virginia Tech (Virginia Polytechnic Institute and State University)
- Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET)
- Museo Argentino de Ciencias Naturales Bernardino Rivadavia
- Universidade de Sao Paulo (USP, University of Sao Paulo)
- Institut Catala de Paleontologia
- Natural History Museum of Los Angeles County
- Natural History Museum of Utah
- Stony Brook University
- Universidad Maimonides
- Museo Friulano di Storia Naturale
- Associação Sul Brasileira de Paleontologia
Members
member name | taxa | specimens | media | media notes | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||||||||||
Martin Ezcurra Project Administrator | 194 | 1 | 1 | 0 | 823 | 0 | 0 | 0 | 0 | 64847 (58059, 0, 6788) | 0 | 686 | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M26833' (194 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Petrolacosaurus kansensis Last Modified in 09/16/20 | 132 | 594 | 576 | 0 | 99 | 0 | 0 | 1 |
[2] † Acerosodontosaurus piveteaui Last Modified in 09/16/20 | 541 | 221 | 214 | 0 | 62 | 0 | 0 | 1 |
[3] † Youngina capensis Last Modified in 09/16/20 | 128 | 611 | 568 | 0 | 89 | 0 | 0 | 1 |
[4] † Paliguana whitei Last Modified in 09/16/20 | 693 | 102 | 100 | 0 | 28 | 0 | 0 | 1 |
[5] † Planocephalosaurus robinsonae Last Modified in 09/16/20 | 291 | 440 | 427 | 0 | 94 | 0 | 0 | 1 |
[6] † Gephyrosaurus bridensis Last Modified in 09/16/20 | 274 | 475 | 447 | 0 | 79 | 0 | 0 | 1 |
[7] † Cteniogenys sp Last Modified in 09/16/20 | 475 | 297 | 290 | 0 | 56 | 0 | 0 | 1 |
[8] † Simoedosaurus lemoinei Last Modified in 09/16/20 | 205 | 540 | 534 | 0 | 83 | 0 | 0 | 1 |
[9] † Aenigmastropheus parringtoni Last Modified in 09/16/20 | 780 | 43 | 41 | 0 | 1 | 0 | 0 | 1 |
[10] † Protorosaurus speneri Last Modified in 09/16/20 | 242 | 516 | 469 | 0 | 78 | 0 | 0 | 1 |
[11] † Dinocephalosaurus orientalis Last Modified in 09/16/20 | 416 | 313 | 310 | 0 | 97 | 0 | 0 | 1 |
[12] † Amotosaurus rotfeldensis Last Modified in 09/16/20 | 441 | 328 | 320 | 0 | 60 | 0 | 0 | 1 |
[13] † Macrocnemus bassanii Last Modified in 09/16/20 | 289 | 458 | 428 | 0 | 81 | 0 | 0 | 1 |
[14] † Macrocnemus obristi Last Modified in 09/16/20 | 734 | 71 | 70 | 0 | 19 | 0 | 0 | 1 |
[15] † Macrocnemus fuyuanensis Last Modified in 09/16/20 | 359 | 391 | 385 | 0 | 79 | 0 | 0 | 1 |
[16] † Fuyuansaurus acutirostris Last Modified in 09/16/20 | 665 | 135 | 131 | 0 | 26 | 0 | 0 | 1 |
[17] † Pectodens zhenyuensis Last Modified in 09/16/20 | 511 | 249 | 242 | 0 | 70 | 0 | 0 | 1 |
[18] † Augustaburiania vatagini Last Modified in 09/16/20 | 743 | 73 | 73 | 0 | 7 | 0 | 0 | 1 |
[19] † Protanystropheus antiquus Last Modified in 09/16/20 | 802 | 22 | 20 | 0 | 1 | 0 | 0 | 1 |
[20] † Trachelosaurus fischeri Last Modified in 09/16/20 | 730 | 87 | 78 | 0 | 15 | 0 | 0 | 1 |
[21] † Tanystropheus longobardicus Last Modified in 09/16/20 | 131 | 603 | 546 | 0 | 106 | 0 | 0 | 1 |
[22] † Tanystropheus haasi Last Modified in 09/16/20 | 812 | 10 | 10 | 0 | 1 | 0 | 0 | 1 |
[23] † Langobadisaurus pandolfii Last Modified in 09/16/20 | 481 | 284 | 274 | 0 | 67 | 0 | 0 | 1 |
[24] † Tanytrachelos ahynis Last Modified in 09/16/20 | 459 | 322 | 302 | 0 | 62 | 0 | 0 | 1 |
[25] † Jesairosaurus lehmani Last Modified in 09/16/20 | 419 | 336 | 320 | 0 | 78 | 0 | 0 | 1 |
[26] † Pamelaria dolichotrachela Last Modified in 09/16/20 | 218 | 548 | 543 | 0 | 62 | 0 | 0 | 1 |
[27] † Azendohsaurus madagaskarensis Last Modified in 09/16/20 | 110 | 647 | 636 | 0 | 72 | 0 | 0 | 1 |
[28] † Azendohsaurus laaroussi Last Modified in 09/16/20 | 748 | 65 | 64 | 0 | 11 | 0 | 0 | 1 |
[29] † Shringasaurus indicus Last Modified in 09/16/20 | 392 | 385 | 378 | 0 | 53 | 0 | 0 | 1 |
[30] † Malerisaurus robinsonae Last Modified in 09/16/20 | 501 | 298 | 287 | 0 | 35 | 0 | 0 | 1 |
[31] † Trilophosaurus buettneri Last Modified in 09/16/20 | 127 | 609 | 577 | 0 | 92 | 0 | 0 | 1 |
[32] † Spinosuchus caseanus Last Modified in 09/16/20 | 433 | 327 | 323 | 0 | 67 | 0 | 0 | 1 |
[33] † Teraterpeton hrynewichorum Last Modified in 09/16/20 | 391 | 359 | 355 | 0 | 77 | 0 | 0 | 1 |
[34] † Arctosaurus osborni Last Modified in 09/16/20 | 809 | 13 | 13 | 0 | 1 | 0 | 0 | 1 |
[35] † Noteosuchus colletti Last Modified in 09/16/20 | 617 | 189 | 178 | 0 | 20 | 0 | 0 | 1 |
[36] † Mesosuchus browni Last Modified in 09/16/20 | 109 | 639 | 608 | 0 | 79 | 0 | 0 | 1 |
[37] † Howesia browni Last Modified in 09/16/20 | 416 | 369 | 344 | 0 | 43 | 0 | 0 | 1 |
[38] † Eohyosaurus wolvaardti Last Modified in 09/16/20 | 647 | 160 | 154 | 0 | 21 | 0 | 0 | 1 |
[39] † Rhynchosaurus articeps Last Modified in 09/16/20 | 229 | 517 | 502 | 0 | 81 | 0 | 0 | 1 |
[40] † Bentonyx sidensis Last Modified in 09/16/20 | 541 | 227 | 226 | 0 | 56 | 0 | 0 | 1 |
[41] † Mesodapedon kuttyi Last Modified in 09/16/20 | 776 | 37 | 37 | 0 | 10 | 0 | 0 | 1 |
[42] † Ammorhynchus navajoi Last Modified in 09/16/20 | 753 | 47 | 46 | 0 | 24 | 0 | 0 | 1 |
[43] † Stenaulorhynchus stockleyi Last Modified in 09/16/20 | 168 | 576 | 574 | 0 | 81 | 0 | 0 | 1 |
[44] † Brasinorhynchus mariantensis Last Modified in 09/16/20 | 497 | 275 | 273 | 0 | 53 | 0 | 0 | 1 |
[45] † Chanares rhynchosaur Last Modified in 09/16/20 | 767 | 48 | 45 | 0 | 11 | 0 | 0 | 1 |
[46] † Langeronyx brodiei Last Modified in 09/16/20 | 638 | 142 | 141 | 0 | 44 | 0 | 0 | 1 |
[47] † Fodonyx spenceri Last Modified in 09/16/20 | 668 | 134 | 134 | 0 | 21 | 0 | 0 | 1 |
[48] † Isalorhynchus genovefae Last Modified in 09/16/20 | 250 | 504 | 500 | 0 | 71 | 0 | 0 | 1 |
[49] † Hyperodapedon gordoni Last Modified in 09/16/20 | 147 | 587 | 582 | 0 | 94 | 0 | 0 | 1 |
[50] † Hyperodapedon huxleyi Last Modified in 09/16/20 | 144 | 587 | 585 | 0 | 94 | 0 | 0 | 1 |
[51] † Teyumbaita sulcognathus Last Modified in 09/16/20 | 264 | 479 | 476 | 0 | 83 | 0 | 0 | 1 |
[52] † Eorasaurus olsoni Last Modified in 09/16/20 | 795 | 28 | 25 | 0 | 2 | 0 | 0 | 1 |
[53] † Prolacertoides jimusarensis Last Modified in 09/16/20 | 722 | 70 | 69 | 0 | 31 | 0 | 0 | 1 |
[54] † Prolacerta broomi Last Modified in 09/16/20 | 69 | 694 | 650 | 0 | 73 | 0 | 0 | 1 |
[55] † K australiensis combined Last Modified in 09/16/20 | 693 | 103 | 98 | 0 | 32 | 0 | 0 | 1 |
[56] † Boreopricea funerea Last Modified in 09/16/20 | 461 | 319 | 295 | 0 | 50 | 0 | 0 | 1 |
[57] † Teyujagua paradoxa Last Modified in 09/16/20 | 513 | 256 | 255 | 0 | 55 | 0 | 0 | 1 |
[58] † Antarctanax shackletoni Last Modified in 09/16/20 | 752 | 66 | 61 | 0 | 6 | 0 | 0 | 1 |
[59] † Archosaurus rossicus holotype Last Modified in 09/16/20 | 803 | 19 | 19 | 0 | 1 | 0 | 0 | 1 |
[60] † Archosaurus complete Last Modified in 09/16/20 | 725 | 100 | 96 | 0 | 2 | 0 | 0 | 1 |
[61] † Proterosuchus fergusi Last Modified in 09/16/20 | 207 | 592 | 577 | 0 | 37 | 0 | 0 | 1 |
[62] † Proterosuchus goweri Last Modified in 09/16/20 | 541 | 269 | 266 | 0 | 16 | 0 | 0 | 1 |
[63] † Proterosuchus alexanderi Last Modified in 09/16/20 | 274 | 509 | 506 | 0 | 42 | 0 | 0 | 1 |
[64] † Chasmatosaurus yuani Last Modified in 09/16/20 | 216 | 569 | 556 | 0 | 50 | 0 | 0 | 1 |
[65] † Panchet proterosuchid Last Modified in 09/16/20 | 639 | 180 | 170 | 0 | 14 | 0 | 0 | 1 |
[66] † Tasmaniosaurus triassicus Last Modified in 09/16/20 | 695 | 121 | 114 | 0 | 14 | 0 | 0 | 1 |
[67] † Vonhuenia fredericki Last Modified in 09/16/20 | 800 | 20 | 20 | 0 | 3 | 0 | 0 | 1 |
[68] † C rossicus combined Last Modified in 09/16/20 | 776 | 45 | 44 | 0 | 3 | 0 | 0 | 1 |
[69] † C magnus combined Last Modified in 09/16/20 | 793 | 29 | 28 | 0 | 2 | 0 | 0 | 1 |
[70] † Chasmatosuchus vjushkovi Last Modified in 09/16/20 | 798 | 26 | 24 | 0 | 1 | 0 | 0 | 1 |
[71] † Koilamasuchus gonzalezdiazi Last Modified in 09/16/20 | 790 | 32 | 29 | 0 | 4 | 0 | 0 | 1 |
[72] † Kalisuchus rewanensis holotype Last Modified in 09/16/20 | 771 | 44 | 40 | 0 | 12 | 0 | 0 | 1 |
[73] † NMQR 3570 Last Modified in 09/16/20 | 779 | 33 | 33 | 0 | 11 | 0 | 0 | 1 |
[74] † Fugusuchus hejiapanensis Last Modified in 09/16/20 | 585 | 219 | 216 | 0 | 21 | 0 | 0 | 1 |
[75] † Sarmatosuchus otschevi Last Modified in 09/16/20 | 568 | 253 | 245 | 0 | 10 | 0 | 0 | 1 |
[76] † Guchengosuchus shiguaiensis Last Modified in 09/16/20 | 608 | 197 | 189 | 0 | 26 | 0 | 0 | 1 |
[77] † Cuyosuchus huenei Last Modified in 09/16/20 | 621 | 200 | 190 | 0 | 12 | 0 | 0 | 1 |
[78] † Garjainia prima Last Modified in 09/16/20 | 184 | 604 | 598 | 0 | 40 | 0 | 0 | 1 |
[79] † Garjainia madiba combined Last Modified in 09/16/20 | 450 | 362 | 346 | 0 | 22 | 0 | 0 | 1 |
[80] † Erythrosuchus africanus Last Modified in 09/16/20 | 125 | 655 | 642 | 0 | 48 | 0 | 0 | 1 |
[81] † Shansisuchus shansisuchus Last Modified in 09/16/20 | 245 | 539 | 532 | 0 | 45 | 0 | 0 | 1 |
[82] † Shansisuchus kuyeheensis Last Modified in 09/16/20 | 726 | 89 | 85 | 0 | 12 | 0 | 0 | 1 |
[83] † Chalishevia cothurnata Last Modified in 09/16/20 | 716 | 98 | 96 | 0 | 11 | 0 | 0 | 1 |
[84] † Youngosuchus sinensis Last Modified in 09/16/20 | 456 | 351 | 346 | 0 | 21 | 0 | 0 | 1 |
[85] † Uralosaurus combined Last Modified in 09/16/20 | 787 | 30 | 29 | 0 | 7 | 0 | 0 | 1 |
[86] † Vancleavea campi Last Modified in 09/16/20 | 286 | 470 | 465 | 0 | 70 | 0 | 0 | 1 |
[87] † Litorosuchus somnii Last Modified in 09/16/20 | 357 | 440 | 434 | 0 | 32 | 0 | 0 | 1 |
[88] † Asperoris mnyama Last Modified in 09/16/20 | 705 | 110 | 95 | 0 | 17 | 0 | 0 | 1 |
[89] † Euparkeria capensis Last Modified in 09/16/20 | 80 | 720 | 709 | 0 | 34 | 0 | 0 | 1 |
[90] † Dorosuchus neoetus Last Modified in 09/16/20 | 706 | 114 | 114 | 0 | 3 | 0 | 0 | 1 |
[91] † Halazhaisuchus qiaoensis Last Modified in 09/16/20 | 721 | 94 | 93 | 0 | 9 | 0 | 0 | 1 |
[92] † Osmolskina czatkoviensis Last Modified in 09/16/20 | 793 | 23 | 23 | 0 | 7 | 0 | 0 | 1 |
[93] † Osmolskina complete Last Modified in 09/16/20 | 354 | 451 | 441 | 0 | 28 | 0 | 0 | 1 |
[94] † Yarasuchus deccanensis Last Modified in 09/16/20 | 602 | 211 | 208 | 0 | 13 | 0 | 0 | 1 |
[95] † Dongusuchus efremovi Last Modified in 09/16/20 | 786 | 35 | 35 | 0 | 2 | 0 | 0 | 1 |
[96] † Teleocrater rhadinus Last Modified in 09/16/20 | 504 | 301 | 300 | 0 | 19 | 0 | 0 | 1 |
[97] † Spondylosoma absconditum Last Modified in 09/16/20 | 738 | 81 | 81 | 0 | 4 | 0 | 0 | 1 |
[98] † Proterochampsa barrionuevoi Last Modified in 09/16/20 | 292 | 485 | 476 | 0 | 55 | 0 | 0 | 1 |
[99] † Proterochampsa nodosa Last Modified in 09/16/20 | 582 | 222 | 216 | 0 | 20 | 0 | 0 | 1 |
[100] † Tropidosuchus romeri Last Modified in 09/16/20 | 201 | 587 | 575 | 0 | 40 | 0 | 0 | 1 |
[101] † Cerritosaurus binsfeldi Last Modified in 09/16/20 | 575 | 226 | 223 | 0 | 25 | 0 | 0 | 1 |
[102] † Gualosuchus reigi Last Modified in 09/16/20 | 284 | 501 | 492 | 0 | 45 | 0 | 0 | 1 |
[103] † Chanaresuchus bonapartei Last Modified in 09/16/20 | 127 | 657 | 643 | 0 | 49 | 0 | 0 | 1 |
[104] † Pseudochampsa ischigualastensis Last Modified in 09/16/20 | 375 | 418 | 412 | 0 | 36 | 0 | 0 | 1 |
[105] † Rhadinosuchus gracilis Last Modified in 09/16/20 | 697 | 119 | 113 | 0 | 13 | 0 | 0 | 1 |
[106] † Archeopelta arborensis Last Modified in 09/16/20 | 712 | 107 | 107 | 0 | 4 | 0 | 0 | 1 |
[107] † Tarjadia ruthae Last Modified in 09/16/20 | 324 | 472 | 469 | 0 | 30 | 0 | 0 | 1 |
[108] † Pagosvenator candelariensis Last Modified in 09/16/20 | 655 | 166 | 165 | 0 | 3 | 0 | 0 | 1 |
[109] † Erpetosuchus granti Last Modified in 09/16/20 | 464 | 334 | 330 | 0 | 27 | 0 | 0 | 1 |
[110] † Erpetosuchus sp Last Modified in 09/16/20 | 719 | 94 | 93 | 0 | 11 | 0 | 0 | 1 |
[111] † Dyoplax arenaceus Last Modified in 09/16/20 | 765 | 57 | 55 | 0 | 3 | 0 | 0 | 1 |
[112] † Jaxtasuchus salomoni Last Modified in 09/16/20 | 604 | 208 | 200 | 0 | 19 | 0 | 0 | 1 |
[113] † Doswellia kaltenbachi Last Modified in 09/16/20 | 416 | 359 | 353 | 0 | 54 | 0 | 0 | 1 |
[114] † Triopticus primus Last Modified in 09/16/20 | 720 | 100 | 92 | 0 | 7 | 0 | 0 | 1 |
[115] † Parasuchus angustifrons Last Modified in 09/16/20 | 554 | 239 | 236 | 0 | 33 | 0 | 0 | 1 |
[116] † Parasuchus hislopi Last Modified in 09/16/20 | 138 | 652 | 643 | 0 | 40 | 0 | 0 | 1 |
[117] † Nicrosaurus kapffi Last Modified in 09/16/20 | 291 | 493 | 485 | 0 | 46 | 0 | 0 | 1 |
[118] † Smilosuchus spp. Last Modified in 09/16/20 | 165 | 612 | 598 | 0 | 54 | 0 | 0 | 1 |
[119] † Diandongosuchus fuyuanensis Last Modified in 09/16/20 | 364 | 436 | 434 | 0 | 25 | 0 | 0 | 1 |
[120] † Wannia scurriensis Last Modified in 09/16/20 | 627 | 171 | 161 | 0 | 32 | 0 | 0 | 1 |
[121] † Ebrachosuchus neukami Last Modified in 09/16/20 | 516 | 274 | 271 | 0 | 35 | 0 | 0 | 1 |
[122] † Paleorhinus sawini Last Modified in 09/16/20 | 551 | 247 | 243 | 0 | 29 | 0 | 0 | 1 |
[123] † Angistorhinus talainti Last Modified in 09/16/20 | 485 | 304 | 301 | 0 | 37 | 0 | 0 | 1 |
[124] † Mystriosuchus planirostris Last Modified in 09/16/20 | 339 | 462 | 443 | 0 | 41 | 0 | 0 | 1 |
[125] † Machaeroprosopus pristinus Last Modified in 09/16/20 | 362 | 431 | 413 | 0 | 48 | 0 | 0 | 1 |
[126] † Redondasaurus spp Last Modified in 09/16/20 | 482 | 313 | 302 | 0 | 39 | 0 | 0 | 1 |
[127] † Ornithosuchus woodwardi Last Modified in 09/16/20 | 281 | 511 | 504 | 0 | 34 | 0 | 0 | 1 |
[128] † Riojasuchus tenuisceps Last Modified in 09/16/20 | 163 | 624 | 613 | 0 | 41 | 0 | 0 | 1 |
[129] † Venaticosuchus rusconii Last Modified in 09/16/20 | 635 | 171 | 169 | 0 | 19 | 0 | 0 | 1 |
[130] † Dynamosuchus collisensis Last Modified in 09/16/20 | 551 | 272 | 269 | 0 | 3 | 0 | 0 | 1 |
[131] † Nundasuchus songeaensis Last Modified in 09/16/20 | 543 | 272 | 267 | 0 | 13 | 0 | 0 | 1 |
[132] † Turfanosuchus dabanensis Last Modified in 09/16/20 | 291 | 508 | 502 | 0 | 28 | 0 | 0 | 1 |
[133] † Gracilisuchus stipanicicorum Last Modified in 09/16/20 | 183 | 610 | 600 | 0 | 40 | 0 | 0 | 1 |
[134] † Yonghesuchus sangbiensis Last Modified in 09/16/20 | 621 | 185 | 182 | 0 | 20 | 0 | 0 | 1 |
[135] † Parringtonia gracilis Last Modified in 09/16/20 | 365 | 435 | 430 | 0 | 28 | 0 | 0 | 1 |
[136] † Otter sandstone archosaur Last Modified in 09/16/20 | 775 | 50 | 45 | 0 | 3 | 0 | 0 | 1 |
[137] † Aetosauroides scagliai Last Modified in 09/16/20 | 290 | 513 | 503 | 0 | 29 | 0 | 0 | 1 |
[138] † Aetosaurus ferratus Last Modified in 09/16/20 | 242 | 547 | 542 | 0 | 39 | 0 | 0 | 1 |
[139] † Stagonolepis robertsoni Last Modified in 09/16/20 | 124 | 658 | 658 | 0 | 41 | 0 | 0 | 1 |
[140] † Ticinosuchus ferox Last Modified in 09/16/20 | 504 | 303 | 296 | 0 | 22 | 0 | 0 | 1 |
[141] † Prestosuchus nyassicus Last Modified in 09/16/20 | 659 | 162 | 156 | 0 | 8 | 0 | 0 | 1 |
[142] † Dagasuchus santacruzensis Last Modified in 09/16/20 | 792 | 29 | 29 | 0 | 2 | 0 | 0 | 1 |
[143] † Qianosuchus mixtus Last Modified in 09/16/20 | 343 | 450 | 448 | 0 | 32 | 0 | 0 | 1 |
[144] † Arizonasaurus babbitti Last Modified in 09/16/20 | 391 | 398 | 393 | 0 | 39 | 0 | 0 | 1 |
[145] † Lotosaurus adentus Last Modified in 09/16/20 | 335 | 414 | 408 | 0 | 80 | 0 | 0 | 1 |
[146] † Ctenosauriscus koeneni Last Modified in 09/16/20 | 773 | 34 | 34 | 0 | 16 | 0 | 0 | 1 |
[147] † Hypselorhachis mirabilis Last Modified in 09/16/20 | 796 | 26 | 25 | 0 | 2 | 0 | 0 | 1 |
[148] † Waldhaus poposauroid Last Modified in 09/16/20 | 789 | 34 | 33 | 0 | 1 | 0 | 0 | 1 |
[149] † Vytshegdosuchus zbeshartensis Last Modified in 09/16/20 | 813 | 9 | 9 | 0 | 1 | 0 | 0 | 1 |
[150] † Bystrowisuchus flerovi Last Modified in 09/16/20 | 805 | 19 | 16 | 0 | 2 | 0 | 0 | 1 |
[151] † Bromsgroveia walkeri Last Modified in 09/16/20 | 782 | 40 | 40 | 0 | 1 | 0 | 0 | 1 |
[152] † Xilousuchus sapingensis Last Modified in 09/16/20 | 589 | 216 | 215 | 0 | 18 | 0 | 0 | 1 |
[153] † Moenkopi poposauroid Last Modified in 09/16/20 | 782 | 39 | 35 | 0 | 6 | 0 | 0 | 1 |
[154] † Mandasuchus tanyauchen Last Modified in 09/16/20 | 541 | 268 | 264 | 0 | 18 | 0 | 0 | 1 |
[155] † Luperosuchus fractus Last Modified in 09/16/20 | 704 | 117 | 116 | 0 | 3 | 0 | 0 | 1 |
[156] † Decuriasuchus quartacolonia Last Modified in 09/16/20 | 402 | 394 | 389 | 0 | 32 | 0 | 0 | 1 |
[157] † Batrachotomus kupferzellensis Last Modified in 09/16/20 | 194 | 600 | 592 | 0 | 34 | 0 | 0 | 1 |
[158] † Prestosuchus chiniquensis Last Modified in 09/16/20 | 121 | 672 | 667 | 0 | 35 | 0 | 0 | 1 |
[159] † Scleromochlus taylori Last Modified in 09/16/20 | 636 | 167 | 158 | 0 | 29 | 0 | 0 | 1 |
[160] † Austriadraco dallavecchiai Last Modified in 09/16/20 | 646 | 177 | 165 | 0 | 12 | 0 | 0 | 1 |
[161] † Preondactylus buffarinii Last Modified in 09/16/20 | 578 | 207 | 202 | 0 | 41 | 0 | 0 | 1 |
[162] † Austriadactylus cristatus Last Modified in 09/16/20 | 602 | 196 | 191 | 0 | 30 | 0 | 0 | 1 |
[163] † Seazzadactylus venieri Last Modified in 09/16/20 | 563 | 223 | 215 | 0 | 45 | 0 | 0 | 1 |
[164] † Raeticodactylus filisurensis Last Modified in 09/16/20 | 583 | 216 | 211 | 0 | 29 | 0 | 0 | 1 |
[165] † Carniadactylus rosenfeldi Last Modified in 09/16/20 | 489 | 274 | 269 | 0 | 65 | 0 | 0 | 1 |
[166] † Peteinosaurus zambellii Last Modified in 09/16/20 | 589 | 202 | 197 | 0 | 37 | 0 | 0 | 1 |
[167] † Caelestiventus hanseni Last Modified in 09/16/20 | 647 | 144 | 140 | 0 | 36 | 0 | 0 | 1 |
[168] † Dimorphodon macronyx Last Modified in 09/16/20 | 371 | 389 | 385 | 0 | 67 | 0 | 0 | 1 |
[169] † Eudimorphodon ranzii Last Modified in 09/16/20 | 494 | 291 | 288 | 0 | 41 | 0 | 0 | 1 |
[170] † Rhamphorhynchus muensteri Last Modified in 09/16/20 | 189 | 517 | 510 | 0 | 124 | 0 | 0 | 1 |
[171] † Cacibupteryx caribensis Last Modified in 09/16/20 | 665 | 132 | 130 | 0 | 27 | 0 | 0 | 1 |
[172] † Allkaruen koi Last Modified in 09/16/20 | 694 | 119 | 117 | 0 | 12 | 0 | 0 | 1 |
[173] † Lagerpeton chanarensis Last Modified in 09/16/20 | 545 | 247 | 242 | 0 | 34 | 0 | 0 | 1 |
[174] † Ixalerpeton polesinensis Last Modified in 09/16/20 | 463 | 332 | 326 | 0 | 34 | 0 | 0 | 1 |
[175] † Kongonaphon kely Last Modified in 09/16/20 | 749 | 63 | 62 | 0 | 12 | 0 | 0 | 1 |
[176] † PVSJ 883 Last Modified in 09/16/20 | 813 | 9 | 9 | 0 | 1 | 0 | 0 | 1 |
[177] † Dromomeron romeri Last Modified in 09/16/20 | 660 | 140 | 134 | 0 | 29 | 0 | 0 | 1 |
[178] † Dromomeron gregorii Last Modified in 09/16/20 | 694 | 125 | 125 | 0 | 4 | 0 | 0 | 1 |
[179] † Dromomeron gigas Last Modified in 09/16/20 | 791 | 27 | 27 | 0 | 5 | 0 | 0 | 1 |
[180] † Lagosuchus talampayensis Last Modified in 09/16/20 | 426 | 368 | 362 | 0 | 35 | 0 | 0 | 1 |
[181] † Lutungutali sitwensis Last Modified in 09/16/20 | 787 | 37 | 33 | 0 | 3 | 0 | 0 | 1 |
[182] † Asilisaurus kongwe Last Modified in 09/16/20 | 333 | 461 | 455 | 0 | 35 | 0 | 0 | 1 |
[183] † Nyasasaurus parringtoni Last Modified in 09/16/20 | 772 | 49 | 48 | 0 | 3 | 0 | 0 | 1 |
[184] † Lewisuchus admixtus Last Modified in 09/16/20 | 301 | 500 | 488 | 0 | 29 | 0 | 0 | 1 |
[185] † Silesaurus opolensis Last Modified in 09/16/20 | 199 | 585 | 578 | 0 | 45 | 0 | 0 | 1 |
[186] † Heterodontosaurus tucki Last Modified in 09/16/20 | 112 | 629 | 624 | 0 | 83 | 0 | 0 | 1 |
[187] † Lesothosaurus diagnosticus Last Modified in 09/16/20 | 126 | 629 | 621 | 0 | 75 | 0 | 0 | 1 |
[188] † Herrerasaurus ischigualastensis Last Modified in 09/16/20 | 99 | 678 | 668 | 0 | 56 | 0 | 0 | 1 |
[189] † Eoraptor lunensis Last Modified in 09/16/20 | 191 | 590 | 586 | 0 | 46 | 0 | 0 | 1 |
[190] † Buriolestes schultzi Last Modified in 09/16/20 | 130 | 641 | 635 | 0 | 56 | 0 | 0 | 1 |
[191] † Saturnalia tupiniquim Last Modified in 09/16/20 | 320 | 459 | 453 | 0 | 49 | 0 | 0 | 1 |
[192] † Tawa hallae Last Modified in 09/16/20 | 234 | 531 | 531 | 0 | 58 | 0 | 0 | 1 |
[193] † Coelophysis bauri Last Modified in 09/16/20 | 89 | 669 | 663 | 0 | 71 | 0 | 0 | 1 |
[194] † Megapnosaurus rhodesiensis Last Modified in 09/16/20 | 107 | 650 | 644 | 0 | 72 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 16877 | |
Project overview | 1836 | |
Matrix views | 944 | Matrix landing page (817 views); Cladistic matrix discrete characters TNT (127 views); |
Media views | 2587 | Media search (1913 views); M693845 (674 views); |
Documents list | 630 | |
Bibliography | 572 | |
Specimen list | 1131 | |
Views for media list | 764 | |
Taxon list | 8413 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 291 | |
Matrix downloads | 119 | Cladistic matrix discrete characters TNT (119 downloads); |
Project downloads | 148 | |
Media downloads | 5 | M693845 (5 downloads); |
Document downloads | 19 | Supplementary Data for inner ear characters (3 downloads); Bayesian matrix discrete characters constraint NEXUS (4 downloads); Bayesian matrix discrete characters unconstraint NEXUS (2 downloads); Cladistic matrix discrete and MG characters TNT (4 downloads); Cladistic matrix discrete characters NEXUS (6 downloads); |