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Brower, A. V. Z., Wahlberg, N., Ogawa, J. R., Boppre, M. and Vane-Wright, R. I. 2010. Phylogenetic relationships among genera of danaine butterflies (Lepidoptera: Nymphalidae) as implied by morphology and DNA sequences. Systematics and Biodiversity. Vol. 8(1), pp. 75-89.
Brower, A. V. Z., Wahlberg, N., Ogawaa, J. R., Boppréc, M. and Vane-Wright, R. I. 2010. Phylogenetic relationships among genera of danaine butterflies (Lepidoptera: Nymphalidae) as implied by morphology and DNA sequences. Systematics and Biodiversity. Vol. 8(1), pp. 75-89.