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Yavorskaya, M. I., Anton, E., Jałoszyński, P., Polilov, A. and Beutel, R. G. 2018. Cephalic anatomy of Sphaeriusidae and a morphology-based phylogeny of the suborder Myxophaga (Coleoptera). Systematic Entomology. Vol. 43(4), pp. 777-797.