Project 4871: D. T. Ksepka, D. J. Field, T. A. Heath, W. Pett, D. B. Thomas, S. Giovanardi, A. J. Tennyson. 2023. Largest-known fossil penguin provides insight into the early evolution of sphenisciform body size and flipper anatomy. Journal of Paleontology. 97 (2):434-453.