Project 2804: J. D. Beech, J. C. Lamsdell, J. Álvaro. 2021. Phylogeny, disparity and mass extinction response in the trilobite order Harpetida. Papers in Palaeontology. 7 (4):2205-2225.

Specimen: † Bohemoharpes ungula Ebach and McNamara, 2002 (YPM/IP:559330)
View: dorsal thorax
View: dorsal thorax
Abstract
The trilobite order Harpetida has long been easily recognized but many unanswered evolutionary questions about the group remain. This work explores the phylogenetic relationships within Harpetida, and studies the harpetid response to the Late Ordovician mass extinction to better understand the relationship between extinction events and disparity. A discrete morphological character matrix was assembled from published descriptions and refined through first-hand observations. This matrix is the first attempt of its kind to characterize the overall morphology of Harpetida, rather than focusing on individual harpetid genera. Phylogenetic analyses under both maximum parsimony and Bayesian inference optimality criteria retrieve tree topologies that support harpetid monophyly but throw doubt onto previous hypotheses of the internal relationships of the order. Harpetid disparity proves remarkably stable over time. A modest peak in the Ordovician is followed by a slow decline throughout the Silurian and Devonian. After the Ordovician, harpetids demonstrate little or no ability to colonize new areas of morphospace. This may represent a fundamental failure to recover, where the lasting impacts of Late Ordovician mass extinction continue to suppress morphological innovation. These findings demonstrate that mass extinction events may have complex impacts that play out over many millions of years.Read the article »
Article DOI: 10.1002/spp2.1399
Project DOI: 10.7934/P2804, http://dx.doi.org/10.7934/P2804
This project contains | Matrices |
---|---|
Download Project SDD File ![]() | Total scored cells: 13218 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 255 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 239 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:255/0 |
Currently Viewing:
MorphoBank Project 2804
MorphoBank Project 2804
- Creation Date:
09 January 2018 - Publication Date:
25 August 2021 - Project views: 29822
- Media downloads: 43
- Matrix downloads: 22
This research
supported by
Authors' Institutions
- West Virginia University
Members
member name | taxa ![]() |
specimens ![]() |
media ![]() | media notes | chars ![]() | character
| cell scorings (scored, NPA, "-") | cell
| rules ![]() | ||||||||||||||
James Beech Project Administrator | 188 | 38 | 45 | 4 | 244 | 0 | 132 | 0 | 0 | 8066 (7135, 5, 926) | 0 | 127 | 0 | 0 | 40 | ||||||||
James Lamsdell Full membership ![]() | 2 | 1 | 0 | 0 | 11 | 0 | 6 | 0 | 0 | 5152 (4263, 0, 889) | 0 | 82 | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M24940' (47 Taxa)
taxon | unscored cells |
scored cells ![]() |
no cell support ![]() |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Eoredlichia intermediata Last Modified in 06/28/19 | 0 | 56 | 55 | 0 | 20 | 0 | 0 | 2 ![]() |
[2] † Baikadamaspis jikdongensis Last Modified in 06/28/19 | 3 | 53 | 52 | 0 | 20 | 0 | 0 | 2 ![]() |
[3] † Bohemoharpes naumanni Last Modified in 06/28/19 | 7 | 63 | 62 | 0 | 6 | 0 | 0 | 2 ![]() |
[4] † Bohemoharpes ungula Last Modified in 06/28/19 | 2 | 66 | 66 | 0 | 8 | 0 | 0 | 2 ![]() |
[5] † Bowmania lassieae Last Modified in 06/28/19 | 3 | 53 | 53 | 0 | 20 | 0 | 0 | 2 ![]() |
[6] † Brachyhipposiderus secundus Last Modified in 06/28/19 | 8 | 63 | 61 | 1 | 5 | 0 | 0 | 2 ![]() |
[7] † Conococheaguea ovata Last Modified in 06/28/19 | 11 | 46 | 45 | 0 | 19 | 0 | 0 | 2 ![]() |
[8] † Dolichoharpes dentoni Last Modified in 06/28/19 | 1 | 71 | 70 | 0 | 4 | 0 | 0 | 2 ![]() |
[9] † Dubhglasina parvula Last Modified in 06/28/19 | 8 | 59 | 58 | 0 | 9 | 0 | 0 | 2 ![]() |
[10] † Entomaspis radiatus Last Modified in 06/28/19 | 3 | 61 | 60 | 0 | 12 | 0 | 0 | 2 ![]() |
[11] † Eoharpes benignensis Last Modified in 06/28/19 | 1 | 70 | 64 | 0 | 5 | 0 | 0 | 2 ![]() |
[12] † Eoharpes primus Last Modified in 06/28/19 | 2 | 69 | 69 | 0 | 5 | 0 | 0 | 2 ![]() |
[13] † Eskoharpes boltoni Last Modified in 06/28/19 | 8 | 62 | 59 | 0 | 6 | 0 | 0 | 2 ![]() |
[14] † Eskoharpes guthae Last Modified in 06/28/19 | 12 | 58 | 58 | 0 | 6 | 0 | 0 | 2 ![]() |
[15] † Eskoharpes palanasus Last Modified in 06/28/19 | 2 | 68 | 66 | 0 | 6 | 0 | 0 | 2 ![]() |
[16] † Eskoharpes wandjina Last Modified in 06/28/19 | 9 | 61 | 61 | 0 | 6 | 0 | 0 | 2 ![]() |
[17] † Globoharpes friendi Last Modified in 06/28/19 | 8 | 62 | 59 | 0 | 6 | 0 | 0 | 2 ![]() |
[18] † Globoharpes teicherti Last Modified in 06/28/19 | 8 | 62 | 61 | 0 | 6 | 0 | 0 | 2 ![]() |
[19] † Harpes gracilis Last Modified in 06/28/19 | 32 | 42 | 40 | 0 | 2 | 0 | 0 | 2 ![]() |
[20] † Harpes macrocephalus Last Modified in 10/31/19 | 0 | 71 | 66 | 0 | 5 | 0 | 0 | 2 ![]() |
[21] † Harpes neogracilis Last Modified in 06/28/19 | 11 | 59 | 59 | 0 | 6 | 0 | 0 | 2 ![]() |
[22] † Harpes whidbournei Last Modified in 06/28/19 | 10 | 61 | 60 | 0 | 5 | 0 | 0 | 2 ![]() |
[23] † Harpides grimmi Last Modified in 06/28/19 | 5 | 44 | 44 | 2 | 25 | 0 | 0 | 2 ![]() |
[24] † Helioharpes perradiatus Last Modified in 06/28/19 | 2 | 68 | 68 | 0 | 6 | 0 | 0 | 2 ![]() |
[25] † Helioharpes radians Last Modified in 06/28/19 | 12 | 58 | 57 | 0 | 6 | 0 | 0 | 2 ![]() |
[26] † Heterocaryon vargum Last Modified in 06/28/19 | 5 | 51 | 51 | 0 | 20 | 0 | 0 | 2 ![]() |
[27] † Hibbertia balclatchiensis Last Modified in 06/28/19 | 9 | 62 | 62 | 0 | 5 | 0 | 0 | 2 ![]() |
[28] † Hibbertia wegelini Last Modified in 06/28/19 | 9 | 61 | 61 | 0 | 6 | 0 | 0 | 2 ![]() |
[29] † Kielania dorbignyana Last Modified in 06/28/19 | 11 | 57 | 57 | 0 | 8 | 0 | 0 | 2 ![]() |
[30] † Lioharpes montagnei Last Modified in 06/28/19 | 12 | 56 | 51 | 0 | 8 | 0 | 0 | 2 ![]() |
[31] † Lioharpes venulosus Last Modified in 06/28/19 | 8 | 60 | 60 | 0 | 8 | 0 | 0 | 2 ![]() |
[32] † Loganopeltis depressa Last Modified in 06/28/19 | 5 | 46 | 46 | 0 | 25 | 0 | 0 | 2 ![]() |
[33] † Loganopeltoides zenkeri Last Modified in 06/28/19 | 6 | 47 | 47 | 0 | 23 | 0 | 0 | 2 ![]() |
[34] † Notchpeakia milleri Last Modified in 01/22/19 | 2 | 54 | 54 | 0 | 20 | 0 | 0 | 2 ![]() |
[35] † Scotoharpes consuetus Last Modified in 06/28/19 | 10 | 59 | 59 | 0 | 7 | 0 | 0 | 2 ![]() |
[36] † Scotoharpes domina Last Modified in 06/28/19 | 11 | 59 | 59 | 0 | 6 | 0 | 0 | 2 ![]() |
[37] † Scotoharpes latior Last Modified in 06/28/19 | 11 | 58 | 55 | 0 | 7 | 0 | 0 | 2 ![]() |
[38] † Scotoharpes loma Last Modified in 06/28/19 | 8 | 62 | 61 | 0 | 6 | 0 | 0 | 2 ![]() |
[39] † Scotoharpes raaschi Last Modified in 06/28/19 | 8 | 62 | 61 | 0 | 6 | 0 | 0 | 2 ![]() |
[40] † Scotoharpes spasskii Last Modified in 06/28/19 | 8 | 61 | 58 | 0 | 7 | 0 | 0 | 2 ![]() |
[41] † Scotoharpes tatouyangensis Last Modified in 06/28/19 | 16 | 53 | 50 | 1 | 6 | 0 | 0 | 2 ![]() |
[42] † Cedaria minor Last Modified in 06/28/19 | 0 | 55 | 55 | 0 | 21 | 0 | 0 | 2 ![]() |
[43] † Coosina kindlei Last Modified in 06/28/19 | 10 | 47 | 46 | 0 | 19 | 0 | 0 | 2 ![]() |
[44] † Crepicephalus iowensis Last Modified in 06/28/19 | 9 | 45 | 45 | 0 | 22 | 0 | 0 | 2 ![]() |
[45] † Modocia whiteleyi Last Modified in 06/28/19 | 1 | 54 | 54 | 0 | 21 | 0 | 0 | 2 ![]() |
[46] † Ptychoparia striata Last Modified in 06/28/19 | 1 | 54 | 54 | 1 | 20 | 0 | 0 | 2 ![]() |
[47] † Tricrepicephalus coria Last Modified in 06/28/19 | 4 | 49 | 48 | 0 | 23 | 0 | 0 | 2 ![]() |
Taxonomic Overview for Matrix 'M25898' (53 Taxa)
taxon | unscored cells |
scored cells ![]() |
no cell support ![]() |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Eoredlichia intermediata Last Modified in 06/28/19 | 9 | 73 | 71 | 0 | 22 | 0 | 0 | 2 ![]() |
[2] † Baikadamaspis jikdongensis Last Modified in 06/28/19 | 14 | 70 | 67 | 0 | 20 | 0 | 0 | 2 ![]() |
[3] † Bohemoharpes naumanni Last Modified in 06/28/19 | 34 | 57 | 55 | 0 | 12 | 0 | 0 | 2 ![]() |
[4] † Bohemoharpes ungula Last Modified in 06/28/19 | 7 | 86 | 82 | 0 | 12 | 0 | 0 | 2 ![]() |
[5] † Bowmania lassieae Last Modified in 06/28/19 | 46 | 45 | 45 | 0 | 12 | 0 | 0 | 2 ![]() |
[6] † Brachyhipposiderus secundus Last Modified in 06/28/19 | 47 | 46 | 45 | 0 | 10 | 0 | 0 | 2 ![]() |
[7] † Chencunia tripleura Last Modified in 02/22/19 | 96 | 7 | 7 | 0 | 0 | 0 | 0 | 2 ![]() |
[8] † Conococheaguea ovata Last Modified in 06/28/19 | 25 | 55 | 54 | 0 | 23 | 0 | 0 | 2 ![]() |
[9] † Dolichoharpes dentoni Last Modified in 06/28/19 | 21 | 78 | 77 | 0 | 4 | 0 | 0 | 2 ![]() |
[10] † Dubhglasina parvula Last Modified in 06/28/19 | 52 | 41 | 40 | 0 | 11 | 0 | 0 | 2 ![]() |
[11] † Entomaspis radiatus Last Modified in 06/28/19 | 20 | 64 | 62 | 0 | 20 | 0 | 0 | 2 ![]() |
[12] † Eoharpes antiquatus Last Modified in 02/22/19 | 21 | 76 | 66 | 0 | 7 | 0 | 0 | 2 ![]() |
[13] † Eoharpes benignensis Last Modified in 06/28/19 | 24 | 70 | 65 | 0 | 10 | 0 | 0 | 2 ![]() |
[14] † Eoharpes primus Last Modified in 06/28/19 | 38 | 56 | 56 | 0 | 9 | 0 | 0 | 2 ![]() |
[15] † Eskoharpes boltoni Last Modified in 06/28/19 | 28 | 71 | 64 | 0 | 7 | 0 | 0 | 2 ![]() |
[16] † Eskoharpes guthae Last Modified in 06/28/19 | 35 | 60 | 58 | 0 | 9 | 0 | 0 | 2 ![]() |
[17] † Eskoharpes palanasus Last Modified in 06/28/19 | 19 | 78 | 73 | 0 | 9 | 0 | 0 | 2 ![]() |
[18] † Eskoharpes wandjina Last Modified in 06/28/19 | 22 | 62 | 60 | 0 | 21 | 0 | 0 | 2 ![]() |
[19] † Globoharpes friendi Last Modified in 06/28/19 | 27 | 67 | 65 | 0 | 9 | 0 | 0 | 2 ![]() |
[20] † Globoharpes teicherti Last Modified in 06/28/19 | 30 | 69 | 66 | 0 | 6 | 0 | 0 | 2 ![]() |
[21] † Harpes gracilis Last Modified in 06/28/19 | 56 | 32 | 29 | 0 | 15 | 0 | 0 | 2 ![]() |
[22] † Harpes macrocephalus Last Modified in 10/31/19 | 26 | 71 | 64 | 0 | 6 | 0 | 0 | 2 ![]() |
[23] † Harpes neogracilis Last Modified in 06/28/19 | 26 | 61 | 59 | 0 | 18 | 0 | 0 | 2 ![]() |
[24] † Harpes whidbournei Last Modified in 06/28/19 | 27 | 57 | 56 | 0 | 19 | 0 | 0 | 2 ![]() |
[25] † Harpides grimmi Last Modified in 06/28/19 | 72 | 28 | 27 | 0 | 3 | 0 | 0 | 2 ![]() |
[26] † Helioharpes perradiatus Last Modified in 06/28/19 | 28 | 61 | 61 | 0 | 14 | 0 | 0 | 2 ![]() |
[27] † Helioharpes radians Last Modified in 06/28/19 | 29 | 56 | 55 | 0 | 18 | 0 | 0 | 2 ![]() |
[28] † Heterocaryon vargum Last Modified in 06/28/19 | 18 | 62 | 60 | 0 | 25 | 0 | 0 | 2 ![]() |
[29] † Hibbertia balclatchiensis Last Modified in 06/28/19 | 46 | 51 | 51 | 0 | 6 | 0 | 0 | 2 ![]() |
[30] † Hibbertia ottawaensis Last Modified in 02/22/19 | 84 | 17 | 17 | 0 | 2 | 0 | 0 | 2 ![]() |
[31] † Hibbertia wegelini Last Modified in 06/28/19 | 28 | 68 | 68 | 0 | 7 | 0 | 0 | 2 ![]() |
[32] † Kielania dorbignyana Last Modified in 06/28/19 | 37 | 55 | 55 | 0 | 11 | 0 | 0 | 2 ![]() |
[33] † Lioharpes montagnei Last Modified in 06/28/19 | 22 | 68 | 61 | 0 | 13 | 0 | 0 | 2 ![]() |
[34] † Lioharpes venulosus Last Modified in 06/28/19 | 21 | 74 | 74 | 0 | 8 | 0 | 0 | 2 ![]() |
[35] † Loganopeltis depressa Last Modified in 06/28/19 | 20 | 77 | 75 | 0 | 8 | 0 | 0 | 2 ![]() |
[36] † Loganopeltoides zenkeri Last Modified in 06/28/19 | 41 | 49 | 46 | 0 | 15 | 0 | 0 | 2 ![]() |
[37] Notchpeakia milleri Last Modified in 06/28/19 | 15 | 72 | 71 | 0 | 16 | 0 | 0 | 2 ![]() |
[38] † Scotoharpes consuetus Last Modified in 06/28/19 | 44 | 46 | 46 | 0 | 13 | 0 | 0 | 2 ![]() |
[39] † Scotoharpes domina Last Modified in 06/28/19 | 39 | 52 | 52 | 0 | 12 | 0 | 0 | 2 ![]() |
[40] † Scotoharpes judex Last Modified in 02/22/19 | 23 | 72 | 72 | 0 | 8 | 0 | 0 | 2 ![]() |
[41] † Scotoharpes latior Last Modified in 06/28/19 | 31 | 64 | 60 | 0 | 9 | 0 | 0 | 2 ![]() |
[42] † Scotoharpes loma Last Modified in 06/28/19 | 22 | 75 | 75 | 0 | 6 | 0 | 0 | 2 ![]() |
[43] † Scotoharpes raaschi Last Modified in 06/28/19 | 38 | 61 | 60 | 0 | 4 | 0 | 0 | 2 ![]() |
[44] † Scotoharpes singularis Last Modified in 02/22/19 | 47 | 40 | 40 | 0 | 16 | 0 | 0 | 2 ![]() |
[45] † Scotoharpes spasskii Last Modified in 06/28/19 | 49 | 44 | 41 | 0 | 11 | 0 | 0 | 2 ![]() |
[46] † Scotoharpes tatouyangensis Last Modified in 06/28/19 | 43 | 53 | 51 | 0 | 7 | 0 | 0 | 2 ![]() |
[47] † Scotoharpes telleri Last Modified in 02/22/19 | 44 | 50 | 50 | 0 | 9 | 0 | 0 | 2 ![]() |
[48] † Cedaria minor Last Modified in 06/28/19 | 14 | 63 | 63 | 0 | 26 | 0 | 0 | 2 ![]() |
[49] † Coosina kindlei Last Modified in 06/28/19 | 27 | 56 | 53 | 0 | 21 | 0 | 0 | 2 ![]() |
[50] † Crepicephalus iowensis Last Modified in 06/28/19 | 26 | 53 | 52 | 0 | 25 | 0 | 0 | 2 ![]() |
[51] † Modocia whiteleyi Last Modified in 06/28/19 | 9 | 64 | 64 | 0 | 30 | 0 | 0 | 2 ![]() |
[52] † Ptychoparia striata Last Modified in 06/28/19 | 9 | 77 | 73 | 0 | 18 | 0 | 0 | 2 ![]() |
[53] † Tricrepicephalus coria Last Modified in 06/28/19 | 37 | 44 | 42 | 0 | 23 | 0 | 0 | 2 ![]() |
Taxonomic Overview for Matrix 'M26129' (86 Taxa)
taxon | unscored cells |
scored cells ![]() |
no cell support ![]() |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Baikadamaspis jikdongensis Last Modified in 06/28/19 | 3 | 53 | 52 | 0 | 20 | 0 | 0 | 2 ![]() |
[2] † Bohemoharpes naumanni Last Modified in 06/28/19 | 7 | 63 | 62 | 0 | 6 | 0 | 0 | 2 ![]() |
[3] † Bohemoharpes ungula Last Modified in 06/28/19 | 2 | 66 | 66 | 0 | 8 | 0 | 0 | 2 ![]() |
[4] † Bowmania lassieae Last Modified in 06/28/19 | 3 | 53 | 53 | 0 | 20 | 0 | 0 | 2 ![]() |
[5] † Brachyhipposiderus secundus Last Modified in 06/28/19 | 9 | 63 | 61 | 0 | 5 | 0 | 0 | 2 ![]() |
[6] † Conococheaguea ovata Last Modified in 06/28/19 | 11 | 46 | 45 | 0 | 19 | 0 | 0 | 2 ![]() |
[7] † Dolichoharpes dentoni Last Modified in 06/28/19 | 1 | 71 | 70 | 0 | 4 | 0 | 0 | 2 ![]() |
[8] † Dubhglasina parvula Last Modified in 06/28/19 | 8 | 59 | 58 | 0 | 9 | 0 | 0 | 2 ![]() |
[9] † Entomaspis radiatus Last Modified in 06/28/19 | 3 | 61 | 60 | 0 | 12 | 0 | 0 | 2 ![]() |
[10] † Eoharpes benignensis Last Modified in 06/28/19 | 1 | 70 | 64 | 0 | 5 | 0 | 0 | 2 ![]() |
[11] † Eoharpes primus Last Modified in 06/28/19 | 2 | 69 | 69 | 0 | 5 | 0 | 0 | 2 ![]() |
[12] † Eskoharpes boltoni Last Modified in 06/28/19 | 8 | 62 | 59 | 0 | 6 | 0 | 0 | 2 ![]() |
[13] † Eskoharpes guthae Last Modified in 06/28/19 | 12 | 58 | 58 | 0 | 6 | 0 | 0 | 2 ![]() |
[14] † Eskoharpes palanasus Last Modified in 06/28/19 | 2 | 68 | 66 | 0 | 6 | 0 | 0 | 2 ![]() |
[15] † Eskoharpes wandjina Last Modified in 06/28/19 | 9 | 61 | 61 | 0 | 6 | 0 | 0 | 2 ![]() |
[16] † Globoharpes friendi Last Modified in 06/28/19 | 8 | 62 | 59 | 0 | 6 | 0 | 0 | 2 ![]() |
[17] † Globoharpes teicherti Last Modified in 06/28/19 | 8 | 62 | 61 | 0 | 6 | 0 | 0 | 2 ![]() |
[18] † Harpes gracilis Last Modified in 06/28/19 | 32 | 42 | 40 | 0 | 2 | 0 | 0 | 2 ![]() |
[19] † Harpes macrocephalus Last Modified in 10/31/19 | 0 | 71 | 66 | 0 | 5 | 0 | 0 | 2 ![]() |
[20] † Harpes neogracilis Last Modified in 06/28/19 | 11 | 59 | 59 | 0 | 6 | 0 | 0 | 2 ![]() |
[21] † Harpes whidbournei Last Modified in 06/28/19 | 10 | 61 | 60 | 0 | 5 | 0 | 0 | 2 ![]() |
[22] † Harpides grimmi Last Modified in 06/28/19 | 7 | 44 | 44 | 0 | 25 | 0 | 0 | 2 ![]() |
[23] † Helioharpes perradiatus Last Modified in 06/28/19 | 2 | 68 | 68 | 0 | 6 | 0 | 0 | 2 ![]() |
[24] † Helioharpes radians Last Modified in 06/28/19 | 12 | 58 | 57 | 0 | 6 | 0 | 0 | 2 ![]() |
[25] † Heterocaryon vargum Last Modified in 06/28/19 | 5 | 51 | 51 | 0 | 20 | 0 | 0 | 2 ![]() |
[26] † Hibbertia balclatchiensis Last Modified in 06/28/19 | 9 | 62 | 62 | 0 | 5 | 0 | 0 | 2 ![]() |
[27] † Hibbertia wegelini Last Modified in 06/28/19 | 9 | 61 | 61 | 0 | 6 | 0 | 0 | 2 ![]() |
[28] † Kielania dorbignyana Last Modified in 06/28/19 | 11 | 57 | 57 | 0 | 8 | 0 | 0 | 2 ![]() |
[29] † Lioharpes montagnei Last Modified in 06/28/19 | 12 | 56 | 51 | 0 | 8 | 0 | 0 | 2 ![]() |
[30] † Lioharpes venulosus Last Modified in 06/28/19 | 8 | 60 | 60 | 0 | 8 | 0 | 0 | 2 ![]() |
[31] † Loganopeltis depressa Last Modified in 06/28/19 | 5 | 46 | 46 | 0 | 25 | 0 | 0 | 2 ![]() |
[32] † Loganopeltoides zenkeri Last Modified in 06/28/19 | 6 | 47 | 47 | 0 | 23 | 0 | 0 | 2 ![]() |
[33] Notchpeakia milleri Last Modified in 06/28/19 | 2 | 54 | 54 | 0 | 20 | 0 | 0 | 2 ![]() |
[34] † Scotoharpes consuetus Last Modified in 06/28/19 | 10 | 59 | 59 | 0 | 7 | 0 | 0 | 2 ![]() |
[35] † Scotoharpes domina Last Modified in 06/28/19 | 11 | 59 | 59 | 0 | 6 | 0 | 0 | 2 ![]() |
[36] † Scotoharpes latior Last Modified in 06/28/19 | 11 | 58 | 55 | 0 | 7 | 0 | 0 | 2 ![]() |
[37] † Scotoharpes loma Last Modified in 06/28/19 | 8 | 62 | 61 | 0 | 6 | 0 | 0 | 2 ![]() |
[38] † Scotoharpes raaschi Last Modified in 06/28/19 | 8 | 62 | 61 | 0 | 6 | 0 | 0 | 2 ![]() |
[39] † Scotoharpes spasskii Last Modified in 06/28/19 | 8 | 61 | 58 | 0 | 7 | 0 | 0 | 2 ![]() |
[40] † Scotoharpes tatouyangensis Last Modified in 06/28/19 | 17 | 53 | 50 | 0 | 6 | 0 | 0 | 2 ![]() |
[41] † Cedaria minor Last Modified in 06/28/19 | 0 | 55 | 55 | 0 | 21 | 0 | 0 | 2 ![]() |
[42] † Coosina kindlei Last Modified in 06/28/19 | 10 | 47 | 46 | 0 | 19 | 0 | 0 | 2 ![]() |
[43] † Crepicephalus iowensis Last Modified in 06/28/19 | 9 | 45 | 45 | 0 | 22 | 0 | 0 | 2 ![]() |
[44] † Modocia whiteleyi Last Modified in 06/28/19 | 1 | 54 | 54 | 0 | 21 | 0 | 0 | 2 ![]() |
[45] † Ptychoparia striata Last Modified in 06/28/19 | 2 | 54 | 54 | 0 | 20 | 0 | 0 | 2 ![]() |
[46] † Tricrepicephalus coria Last Modified in 06/28/19 | 4 | 49 | 48 | 0 | 23 | 0 | 0 | 2 ![]() |
[47] † Ancestor_1 Last Modified in 06/28/19 | 0 | 68 | 68 | 0 | 8 | 0 | 0 | 2 ![]() |
[48] † Ancestor_2 Last Modified in 06/28/19 | 0 | 71 | 71 | 0 | 5 | 0 | 0 | 2 ![]() |
[49] † Ancestor_3 Last Modified in 06/28/19 | 0 | 70 | 70 | 0 | 6 | 0 | 0 | 2 ![]() |
[50] Ancestor_4 Last Modified in 06/28/19 | 0 | 72 | 72 | 0 | 4 | 0 | 0 | 2 ![]() |
[51] † Ancestor_5 Last Modified in 06/28/19 | 0 | 72 | 72 | 0 | 4 | 0 | 0 | 2 ![]() |
[52] † Ancestor_6 Last Modified in 06/28/19 | 0 | 71 | 71 | 0 | 5 | 0 | 0 | 2 ![]() |
[53] † Ancestor_7 Last Modified in 06/28/19 | 0 | 73 | 73 | 0 | 3 | 0 | 0 | 2 ![]() |
[54] † Ancestor_8 Last Modified in 06/28/19 | 0 | 74 | 74 | 0 | 2 | 0 | 0 | 2 ![]() |
[55] † Ancestor_9 Last Modified in 06/28/19 | 0 | 73 | 73 | 0 | 3 | 0 | 0 | 2 ![]() |
[56] † Ancestor_10 Last Modified in 06/28/19 | 0 | 74 | 74 | 0 | 2 | 0 | 0 | 2 ![]() |
[57] † Ancestor_11 Last Modified in 06/28/19 | 0 | 73 | 73 | 0 | 3 | 0 | 0 | 2 ![]() |
[58] † Ancestor_12 Last Modified in 06/28/19 | 0 | 74 | 74 | 0 | 2 | 0 | 0 | 2 ![]() |
[59] † Ancestor_13 Last Modified in 06/28/19 | 0 | 73 | 73 | 0 | 3 | 0 | 0 | 2 ![]() |
[60] † Ancestor_14 Last Modified in 06/28/19 | 0 | 74 | 74 | 0 | 2 | 0 | 0 | 2 ![]() |
[61] † Ancestor_15 Last Modified in 06/28/19 | 0 | 73 | 73 | 0 | 3 | 0 | 0 | 2 ![]() |
[62] † Ancestor_16 Last Modified in 06/28/19 | 0 | 73 | 73 | 0 | 3 | 0 | 0 | 2 ![]() |
[63] † Ancestor_17 Last Modified in 06/28/19 | 0 | 74 | 74 | 0 | 2 | 0 | 0 | 2 ![]() |
[64] † Ancestor_18 Last Modified in 06/28/19 | 0 | 73 | 73 | 0 | 3 | 0 | 0 | 2 ![]() |
[65] † Ancestor_19 Last Modified in 06/28/19 | 0 | 74 | 74 | 0 | 2 | 0 | 0 | 2 ![]() |
[66] † Ancestor_20 Last Modified in 06/28/19 | 0 | 75 | 75 | 0 | 1 | 0 | 0 | 2 ![]() |
[67] † Ancestor_21 Last Modified in 06/28/19 | 0 | 75 | 75 | 0 | 1 | 0 | 0 | 2 ![]() |
[68] † Ancestor_22 Last Modified in 06/28/19 | 0 | 75 | 75 | 0 | 1 | 0 | 0 | 2 ![]() |
[69] † Ancestor_23 Last Modified in 06/28/19 | 0 | 75 | 75 | 0 | 1 | 0 | 0 | 2 ![]() |
[70] † Ancestor_24 Last Modified in 06/28/19 | 0 | 75 | 75 | 0 | 1 | 0 | 0 | 2 ![]() |
[71] † Ancestor_25 Last Modified in 06/28/19 | 0 | 73 | 73 | 0 | 3 | 0 | 0 | 2 ![]() |
[72] † Ancestor_26 Last Modified in 06/28/19 | 0 | 74 | 74 | 0 | 2 | 0 | 0 | 2 ![]() |
[73] † Ancestor_27 Last Modified in 06/28/19 | 0 | 75 | 75 | 0 | 1 | 0 | 0 | 2 ![]() |
[74] † Ancestor_28 Last Modified in 06/28/19 | 0 | 75 | 75 | 0 | 1 | 0 | 0 | 2 ![]() |
[75] † Ancestor_29 Last Modified in 06/28/19 | 0 | 75 | 75 | 0 | 1 | 0 | 0 | 2 ![]() |
[76] † Ancestor_30 Last Modified in 06/28/19 | 0 | 76 | 76 | 0 | 0 | 0 | 0 | 2 ![]() |
[77] † Ancestor_31 Last Modified in 06/28/19 | 0 | 76 | 76 | 0 | 0 | 0 | 0 | 2 ![]() |
[78] † Ancestor_32 Last Modified in 06/28/19 | 0 | 76 | 76 | 0 | 0 | 0 | 0 | 2 ![]() |
[79] † Ancestor_33 Last Modified in 06/28/19 | 0 | 72 | 72 | 0 | 4 | 0 | 0 | 2 ![]() |
[80] † Ancestor_34 Last Modified in 06/28/19 | 0 | 72 | 72 | 0 | 4 | 0 | 0 | 2 ![]() |
[81] † Ancestor_35 Last Modified in 06/28/19 | 0 | 72 | 72 | 0 | 4 | 0 | 0 | 2 ![]() |
[82] † Ancestor_36 Last Modified in 06/28/19 | 0 | 72 | 72 | 0 | 4 | 0 | 0 | 2 ![]() |
[83] † Ancestor_37 Last Modified in 06/28/19 | 1 | 72 | 72 | 0 | 3 | 0 | 0 | 2 ![]() |
[84] † Ancestor_38 Last Modified in 06/28/19 | 0 | 73 | 73 | 0 | 3 | 0 | 0 | 2 ![]() |
[85] † Ancestor_39 Last Modified in 06/28/19 | 1 | 71 | 71 | 0 | 4 | 0 | 0 | 2 ![]() |
[86] † Eoredlichia intermediata Last Modified in 06/28/19 | 0 | 56 | 55 | 0 | 20 | 0 | 0 | 2 ![]() |
Project views 
type | number of views | Individual items viewed (where applicable) |
Total project views | 29822 | |
Project overview | 733 | |
Media views | 13130 | Media search (1546 views); M565506 (219 views); M565507 (215 views); M565508 (229 views); M565509 (220 views); M565510 (217 views); M565511 (250 views); M565512 (216 views); M565513 (234 views); M565514 (404 views); M565515 (247 views); M565516 (281 views); M565517 (220 views); M565518 (234 views); M565519 (214 views); M565520 (214 views); M565521 (238 views); M565522 (227 views); M565523 (233 views); M565524 (221 views); M565525 (214 views); M565526 (228 views); M565527 (228 views); M565528 (239 views); M565529 (222 views); M565530 (226 views); M565531 (230 views); M565532 (217 views); M565533 (215 views); M565534 (236 views); M565535 (234 views); M586603 (236 views); M595166 (232 views); M595167 (281 views); M595168 (323 views); M595169 (226 views); M595170 (215 views); M595171 (202 views); M595172 (414 views); M595173 (527 views); M595174 (383 views); M595181 (236 views); M595186 (224 views); M595188 (307 views); M651731 (369 views); M651735 (387 views); |
Matrix views | 652 | Matrix landing page (467 views); Character Matrix 1 (66 views); Backup matrix (57 views); Appendix S3. Character-taxon matrix. (62 views); |
Views for media list | 871 | |
Documents list | 443 | |
Bibliography | 861 | |
Taxon list | 3807 | |
Specimen list | 9325 |
Project downloads 
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 133 | |
Project downloads | 57 | |
Document downloads | 11 | Appendix S2. Illustrated character list. (9 downloads); Appendix S1: Characters for Phylogenetic Analysis (2 downloads); |
Matrix downloads | 22 | Appendix S3. Character-taxon matrix. (8 downloads); Character Matrix 1 (10 downloads); Backup matrix (4 downloads); |
Media downloads | 43 | M565519 (1 download); M565520 (1 download); M565521 (3 downloads); M565523 (3 downloads); M565524 (3 downloads); M565525 (1 download); M565528 (2 downloads); M565527 (1 download); M565530 (1 download); M565531 (1 download); M565533 (1 download); M565534 (1 download); M595169 (2 downloads); M595186 (1 download); M595172 (1 download); M595173 (1 download); M595188 (3 downloads); M595174 (1 download); M565506 (1 download); M565508 (2 downloads); M565509 (1 download); M565510 (1 download); M565514 (1 download); M565515 (1 download); M565522 (1 download); M565526 (1 download); M586603 (1 download); M595166 (1 download); M595167 (1 download); M595170 (1 download); M651735 (2 downloads); |