Project 2086: D. W. Krause, S. Hoffmann, J. R. Wible, E. C. Kirk, J. A. Schultz, W. von Koenigswald, J. R. Groenke, J. B. Rossie, P. M. O’Connor, E. R. Seiffert, E. R. Dumont, W. L. Holloway, R. R. Rogers, L. J. Rahantarisoa, A. D. Kemp, H. Andriamialison. 2014. First cranial remains of a gondwanatherian mammal reveal remarkable mosaicism. Nature. 515 (7528):512-517.
Abstract
Previously known only from isolated teeth and lower jaw fragments recovered from the Cretaceous and Palaeogene of the Southern Hemisphere, the Gondwanatheria constitute the most poorly known of all major mammaliaform radiations. Here we report the discovery of the first skull material of a gondwanatherian, a complete and well-preserved cranium from Upper Cretaceous strata in Madagascar that we assign to a new genus and species. Phylogenetic analysis strongly supports its placement within Gondwanatheria, which are recognized as monophyletic and closely related to multituberculates, an evolutionarily successful clade of Mesozoic mammals known almost exclusively from the Northern Hemisphere. The new taxon is the largest known mammaliaform from the Mesozoic of Gondwana. Its craniofacial anatomy reveals that it was herbivorous, large-eyed and agile, with well-developed high-frequency hearing and a keen sense of smell. The cranium exhibits a mosaic of primitive and derived features, the disparity of which is extreme and probably reflective of a long evolutionary history in geographic isolation.Read the article »
Article DOI: 10.1038/nature13922
Project DOI: 10.7934/P2086, http://dx.doi.org/10.7934/P2086
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 22667 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 453 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 1088 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:453/0 |
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MorphoBank Project 2086
MorphoBank Project 2086
- Creation Date:
18 December 2014 - Publication Date:
18 December 2014 - Project views: 42164
- Matrix downloads: 29
This research
supported by
Authors' Institutions
- Ohio University
- University of Texas at Austin
- Rheinische Friedrich-Wilhelms-Universität Bonn (University of Bonn)
- Carnegie Museum of Natural History
- Macalester College
- New York Institute of Technology
- Stony Brook University
- University of Massachusetts Amherst
- Université d’Antananarivo
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 92 | 0 | 1 | 453 | 0 | 0 | 22667 (16446, 0, 6221) | 0 | 0 | 0 | ||||
Maureen Admin Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Simone Hoffmann Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
David Krause Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
John Wible Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M23163' (91 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Thrinaxodon Last Modified in 12/22/14 | 39 | 263 | 256 | 0 | 158 | 0 | 0 | 5 |
[2] † Probainognathus Last Modified in 12/22/14 | 50 | 248 | 248 | 0 | 155 | 0 | 0 | 5 |
[3] † Exaeretodon Last Modified in 12/22/14 | 65 | 245 | 238 | 0 | 150 | 0 | 0 | 5 |
[4] † Diademodon Last Modified in 12/22/14 | 53 | 245 | 244 | 0 | 156 | 0 | 0 | 5 |
[5] † Cynognathus Last Modified in 12/22/14 | 81 | 217 | 217 | 0 | 155 | 0 | 0 | 5 |
[6] † Oligokyphus Last Modified in 12/22/14 | 63 | 252 | 251 | 0 | 139 | 0 | 0 | 5 |
[7] † Bienotherium Last Modified in 12/22/14 | 122 | 191 | 191 | 0 | 140 | 0 | 0 | 5 |
[8] † Kayentatherium Last Modified in 12/22/14 | 84 | 224 | 223 | 0 | 146 | 0 | 0 | 5 |
[9] † Yunnanodon Last Modified in 12/22/14 | 190 | 129 | 129 | 0 | 134 | 0 | 0 | 5 |
[10] † Bocatherium Last Modified in 12/22/14 | 229 | 94 | 94 | 0 | 130 | 0 | 0 | 5 |
[11] † Bienotheroides Last Modified in 12/22/14 | 93 | 225 | 225 | 0 | 135 | 0 | 0 | 5 |
[12] † Tritheledontidae Last Modified in 01/21/15 | 65 | 250 | 234 | 0 | 154 | 0 | 0 | 5 |
[13] † Adelobasileus Last Modified in 12/22/14 | 385 | 61 | 61 | 0 | 7 | 0 | 0 | 5 |
[14] † Sinoconodon Last Modified in 12/22/14 | 67 | 332 | 330 | 0 | 56 | 0 | 0 | 5 |
[15] † Morganucodon Last Modified in 12/22/14 | 33 | 373 | 369 | 0 | 51 | 0 | 0 | 5 |
[16] † Megazostrodon Last Modified in 12/22/14 | 133 | 273 | 273 | 0 | 47 | 0 | 0 | 5 |
[17] † Haldanodon Last Modified in 12/22/14 | 83 | 326 | 324 | 0 | 46 | 0 | 0 | 5 |
[18] † Hadrocodium Last Modified in 12/22/14 | 181 | 223 | 223 | 0 | 49 | 0 | 0 | 5 |
[19] † Pseudotribos Last Modified in 12/22/14 | 294 | 132 | 132 | 0 | 27 | 0 | 0 | 5 |
[20] † Ambondro Last Modified in 12/22/14 | 379 | 55 | 54 | 0 | 20 | 0 | 0 | 5 |
[21] † Ausktribosphenos Last Modified in 12/22/14 | 352 | 83 | 83 | 0 | 18 | 0 | 0 | 5 |
[22] † Henosferus Last Modified in 12/22/14 | 332 | 102 | 101 | 0 | 20 | 0 | 0 | 5 |
[23] † Asfaltomylos Last Modified in 12/22/14 | 362 | 72 | 72 | 0 | 19 | 0 | 0 | 5 |
[24] † Bishops Last Modified in 12/22/14 | 341 | 92 | 91 | 0 | 21 | 0 | 0 | 5 |
[25] † Steropodon Last Modified in 12/22/14 | 388 | 46 | 46 | 0 | 19 | 0 | 0 | 5 |
[26] † Teinolophos Last Modified in 12/22/14 | 366 | 69 | 68 | 0 | 19 | 0 | 0 | 5 |
[27] † Obdurodon Last Modified in 12/22/14 | 148 | 264 | 263 | 0 | 42 | 0 | 0 | 5 |
[28] Ornithorhynchus Last Modified in 12/18/14 | 46 | 333 | 331 | 0 | 76 | 0 | 0 | 5 |
[29] † Gobiconodon Last Modified in 12/22/14 | 127 | 280 | 272 | 0 | 54 | 0 | 0 | 5 |
[30] † Repenomamus Last Modified in 12/22/14 | 120 | 282 | 281 | 0 | 52 | 0 | 0 | 5 |
[31] † Yanoconodon Last Modified in 12/22/14 | 222 | 181 | 181 | 0 | 50 | 0 | 0 | 5 |
[32] † Jeholodens Last Modified in 12/22/14 | 229 | 178 | 176 | 0 | 48 | 0 | 0 | 5 |
[33] † Amphilestes Last Modified in 12/22/14 | 342 | 74 | 74 | 0 | 37 | 0 | 0 | 5 |
[34] † Priacodon Last Modified in 12/22/14 | 207 | 197 | 197 | 0 | 49 | 0 | 0 | 5 |
[35] † Trioracodon Last Modified in 12/22/14 | 237 | 170 | 169 | 0 | 47 | 0 | 0 | 5 |
[36] † Tinodon Last Modified in 12/22/14 | 304 | 110 | 110 | 0 | 39 | 0 | 0 | 5 |
[37] † Akidolestes Last Modified in 12/22/14 | 271 | 138 | 138 | 0 | 44 | 0 | 0 | 5 |
[38] † Spalacotherium Last Modified in 12/22/14 | 311 | 99 | 99 | 0 | 43 | 0 | 0 | 5 |
[39] † Maotherium Last Modified in 12/22/14 | 147 | 260 | 257 | 0 | 49 | 0 | 0 | 5 |
[40] † Zhangheotherium Last Modified in 12/22/14 | 199 | 217 | 216 | 0 | 38 | 0 | 0 | 5 |
[41] † Amphitherium Last Modified in 12/22/14 | 333 | 98 | 98 | 0 | 22 | 0 | 0 | 5 |
[42] † Cronopio Last Modified in 12/22/14 | 207 | 208 | 206 | 0 | 40 | 0 | 0 | 5 |
[43] † Henkelotherium Last Modified in 12/22/14 | 199 | 222 | 222 | 0 | 32 | 0 | 0 | 5 |
[44] † Dryolestes Last Modified in 12/22/14 | 215 | 205 | 205 | 0 | 33 | 0 | 0 | 5 |
[45] † Peramus Last Modified in 12/22/14 | 285 | 146 | 141 | 0 | 27 | 0 | 0 | 5 |
[46] † Vincelestes Last Modified in 12/22/14 | 39 | 380 | 379 | 0 | 35 | 0 | 0 | 5 |
[47] † Kielantherium Last Modified in 12/22/14 | 339 | 94 | 92 | 0 | 22 | 0 | 0 | 5 |
[48] † Sinodelphys Last Modified in 12/22/14 | 265 | 163 | 163 | 0 | 25 | 0 | 0 | 5 |
[49] † Pappotherium Last Modified in 12/22/14 | 369 | 64 | 64 | 0 | 20 | 0 | 0 | 5 |
[50] † Deltatheridium Last Modified in 12/22/14 | 166 | 254 | 252 | 0 | 35 | 0 | 0 | 5 |
[51] † Asiatherium Last Modified in 12/22/14 | 179 | 248 | 248 | 0 | 26 | 0 | 0 | 5 |
[52] † Kokopellia Last Modified in 12/22/14 | 318 | 113 | 113 | 0 | 22 | 0 | 0 | 5 |
[53] † Pucadelphys Last Modified in 12/22/14 | 59 | 358 | 354 | 0 | 40 | 0 | 0 | 5 |
[54] Didelphis Last Modified in 12/18/14 | 15 | 376 | 375 | 0 | 63 | 0 | 0 | 5 |
[55] † Juramaia Last Modified in 12/22/14 | 287 | 146 | 146 | 0 | 20 | 0 | 0 | 5 |
[56] † Eomaia Last Modified in 12/22/14 | 190 | 226 | 224 | 0 | 39 | 0 | 0 | 5 |
[57] † Prokennalestes Last Modified in 12/22/14 | 239 | 194 | 192 | 0 | 22 | 0 | 0 | 5 |
[58] † Asioryctes Last Modified in 12/22/14 | 101 | 316 | 313 | 0 | 39 | 0 | 0 | 5 |
[59] † Zalambdalestes Last Modified in 12/22/14 | 51 | 365 | 358 | 0 | 44 | 0 | 0 | 5 |
[60] † Ukhaatherium Last Modified in 12/22/14 | 123 | 290 | 289 | 0 | 41 | 0 | 0 | 5 |
[61] † Gomphos Last Modified in 12/22/14 | 114 | 287 | 287 | 0 | 52 | 0 | 0 | 5 |
[62] † Tribosphenomys Last Modified in 12/22/14 | 255 | 154 | 150 | 0 | 48 | 0 | 0 | 5 |
[63] Erinaceus Last Modified in 12/18/14 | 5 | 398 | 397 | 0 | 51 | 0 | 0 | 5 |
[64] Dasypus Last Modified in 12/18/14 | 97 | 288 | 284 | 0 | 72 | 0 | 0 | 5 |
[65] † Ferugliotherium a Last Modified in 12/22/14 | 360 | 12 | 12 | 0 | 81 | 0 | 0 | 5 |
[66] † Ferugliotherium b Last Modified in 12/22/14 | 328 | 32 | 32 | 0 | 93 | 0 | 0 | 5 |
[67] † Trapalcotherium Last Modified in 12/22/14 | 368 | 8 | 8 | 0 | 77 | 0 | 0 | 5 |
[68] † Bharattherium Last Modified in 12/22/14 | 364 | 13 | 13 | 0 | 76 | 0 | 0 | 5 |
[69] † Gondwanatherium Last Modified in 12/22/14 | 325 | 30 | 30 | 0 | 98 | 0 | 0 | 5 |
[70] † Lavanify Last Modified in 12/22/14 | 364 | 13 | 13 | 0 | 76 | 0 | 0 | 5 |
[71] † Sudamerica a Last Modified in 12/22/14 | 304 | 45 | 44 | 0 | 105 | 0 | 0 | 5 |
[72] † Sudamerica b Last Modified in 12/22/14 | 307 | 54 | 53 | 0 | 93 | 0 | 0 | 5 |
[73] † Tanzanian taxon A Last Modified in 01/21/15 | 337 | 26 | 26 | 0 | 90 | 0 | 0 | 5 |
[74] † Tanzanian taxon B Last Modified in 01/21/15 | 338 | 27 | 27 | 0 | 88 | 0 | 0 | 5 |
[75] † Greniodon Last Modified in 01/05/15 | 360 | 18 | 17 | 0 | 76 | 0 | 0 | 5 |
[76] † Vintana a Last Modified in 01/05/15 | 171 | 198 | 198 | 0 | 84 | 0 | 0 | 5 |
[77] † Vintana b Last Modified in 01/05/15 | 171 | 199 | 199 | 0 | 83 | 0 | 0 | 5 |
[78] † Paulchoffatiidae Last Modified in 01/21/15 | 168 | 198 | 187 | 0 | 98 | 0 | 0 | 5 |
[79] † Plagiaulacidae Last Modified in 01/21/15 | 262 | 101 | 99 | 0 | 92 | 0 | 0 | 5 |
[80] † Ptilodus Last Modified in 12/22/14 | 84 | 265 | 265 | 0 | 104 | 0 | 0 | 5 |
[81] † Taeniolabis Last Modified in 12/22/14 | 184 | 160 | 159 | 0 | 110 | 0 | 0 | 5 |
[82] † Lambdopsalis Last Modified in 12/22/14 | 83 | 256 | 256 | 0 | 114 | 0 | 0 | 5 |
[83] † Kryptobaatar Last Modified in 12/22/14 | 35 | 313 | 312 | 0 | 106 | 0 | 0 | 5 |
[84] † Catopsbaatar Last Modified in 12/22/14 | 122 | 228 | 227 | 0 | 104 | 0 | 0 | 5 |
[85] † Chulsanbaatar Last Modified in 12/22/14 | 87 | 264 | 263 | 0 | 103 | 0 | 0 | 5 |
[86] † Nemegtbaatar Last Modified in 12/22/14 | 87 | 263 | 263 | 0 | 103 | 0 | 0 | 5 |
[87] † Thomasia Last Modified in 12/22/14 | 328 | 39 | 38 | 0 | 87 | 0 | 0 | 5 |
[88] † Haramiyavia Last Modified in 12/22/14 | 295 | 72 | 72 | 0 | 86 | 0 | 0 | 5 |
[89] † Arboroharamiya Last Modified in 12/22/14 | 263 | 94 | 93 | 0 | 97 | 0 | 0 | 5 |
[90] † Rugosodon Last Modified in 12/22/14 | 218 | 143 | 143 | 0 | 92 | 0 | 0 | 5 |
[91] † Sinobaatar Last Modified in 12/22/14 | 183 | 177 | 174 | 0 | 96 | 0 | 0 | 5 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 42164 | |
Project overview | 4170 | |
Matrix views | 2724 | Matrix landing page (2356 views); First cranial remains of a gondwanatherian mammal reveal remarkable mosaicism (368 views); |
Taxon list | 24486 | |
Media views | 6547 | Media search (5608 views); M348988 (939 views); |
Bibliography | 4233 | |
Documents list | 3 | |
Specimen list | 1 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 381 | |
Matrix downloads | 29 | First cranial remains of a gondwanatherian mammal reveal remarkable mosaicism (29 downloads); |
Document downloads | 2 | Acyon Ventral Vector File (.ai) (1 download); Cladosictis Ventral Vector File (.eps) (1 download); |
Project downloads | 350 |