Project 2606: J. C. Lamsdell. 2021. A new method for quantifying heterochrony in evolutionary lineages. Paleobiology. 47:363-384.
Abstract
The occupation of new environments by evolutionary lineages is frequently associated with morphological changes. This covariation of ecotype and phenotype is expected due to the process of natural selection, whereby environmental pressures lead to the proliferation of morphological variants that are a better fit for the prevailing abiotic conditions. One primary mechanism by which phenotypic variants are known to arise is through changes in the timing or duration of organismal development resulting in alterations to adult morphology, a process known as heterochrony. While numerous studies have demonstrated heterochronic trends in association with environmental gradients, few have done so within a phylogenetic context. Understanding species interrelationships is necessary to determine whether morphological change is due to heterochronic processes; however, research is hampered by the lack of a quantitative metric with which to assess the degree of heterochronic traits expressed within and among species. Here I present a new metric for quantifying heterochronic change, expressed as a heterochronic weighting, and apply it to xiphosuran chelicerates within a phylogenetic context to reveal concerted independent heterochronic trends. These trends correlate with shifts in environmental occupation from marine to nonmarine habitats, resulting in a macroevolutionary ratchet. Critically, the distribution of heterochronic weightings among species shows evidence of being influenced by both historical, phylogenetic processes and external ecological pressures. Heterochronic weighting proves to be an effective method to quantify heterochronic trends within a phylogenetic framework and is readily applicable to any group of organisms that have well-defined morphological characteristics, ontogenetic information, and resolved internal relationships.Read the article »
Article DOI: 10.1017/pab.2020.17
Project DOI: 10.7934/P2606, http://dx.doi.org/10.7934/P2606
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 26774 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 256 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 612 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:256/0 |
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MorphoBank Project 2606
MorphoBank Project 2606
- Creation Date:
13 November 2020 - Publication Date:
04 June 2021 - Project views: 12156
- Matrix downloads: 1
Authors' Institutions
- West Virginia University
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
James Lamsdell Project Administrator | 153 | 3 | 1 | 256 | 0 | 0 | 26774 (20760, 0, 6014) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M24451' (153 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Fuxianhuia protensa Last Modified in 07/02/14 | 35 | 148 | 148 | 0 | 73 | 0 | 0 | 1 |
[2] † Leanchoilia illecebrosa Last Modified in 07/02/14 | 36 | 162 | 162 | 0 | 58 | 0 | 0 | 1 |
[3] † Alalcomenaeus cambricus Last Modified in 07/02/14 | 28 | 169 | 169 | 0 | 59 | 0 | 0 | 1 |
[4] † Yohoia tenuis Last Modified in 07/02/14 | 70 | 132 | 132 | 0 | 54 | 0 | 0 | 1 |
[5] † Emeraldella brocki Last Modified in 07/02/14 | 21 | 164 | 164 | 0 | 71 | 0 | 0 | 1 |
[6] † Sydneyia inexpectans Last Modified in 07/02/14 | 42 | 146 | 146 | 0 | 68 | 0 | 0 | 1 |
[7] † Olenoides serratus Last Modified in 07/02/14 | 13 | 186 | 186 | 0 | 57 | 0 | 0 | 1 |
[8] Pycnogonum litorale Last Modified in 07/02/14 | 8 | 102 | 102 | 0 | 146 | 0 | 0 | 1 |
[9] † Haliestes dasos Last Modified in 07/02/14 | 39 | 85 | 85 | 0 | 132 | 0 | 0 | 1 |
[10] † Palaeoisopus problematicus Last Modified in 07/02/14 | 22 | 116 | 116 | 0 | 118 | 0 | 0 | 1 |
[11] † Offacolus kingi Last Modified in 07/02/14 | 87 | 130 | 130 | 0 | 39 | 0 | 0 | 1 |
[12] † Dibasterium durgae Last Modified in 07/02/14 | 57 | 148 | 148 | 0 | 51 | 0 | 0 | 1 |
[13] † Anderella parva Last Modified in 08/20/14 | 155 | 86 | 86 | 0 | 15 | 0 | 0 | 1 |
[14] † Weinbergina opitzi Last Modified in 07/02/14 | 67 | 160 | 160 | 0 | 29 | 0 | 0 | 1 |
[15] † Camanchia grovensis Last Modified in 07/02/14 | 146 | 94 | 94 | 0 | 16 | 0 | 0 | 1 |
[16] † Venustulus waukeshaensis Last Modified in 07/02/14 | 127 | 110 | 110 | 0 | 19 | 0 | 0 | 1 |
[17] † Legrandella lombardi Last Modified in 07/02/14 | 137 | 105 | 105 | 0 | 14 | 0 | 0 | 1 |
[18] † Bembicosoma pomphicus Last Modified in 07/02/14 | 158 | 84 | 84 | 0 | 14 | 0 | 0 | 1 |
[19] † Bunodes lunula Last Modified in 07/02/14 | 155 | 85 | 85 | 0 | 16 | 0 | 0 | 1 |
[20] † Limuloides limuloides Last Modified in 07/02/14 | 148 | 95 | 95 | 0 | 13 | 0 | 0 | 1 |
[21] † Pasternakevia podolica Last Modified in 07/02/14 | 156 | 86 | 86 | 0 | 14 | 0 | 0 | 1 |
[22] † Cyamocephalus loganensis Last Modified in 07/02/14 | 150 | 92 | 92 | 0 | 14 | 0 | 0 | 1 |
[23] † Pseudoniscus roosevelti Last Modified in 07/02/14 | 155 | 87 | 87 | 0 | 14 | 0 | 0 | 1 |
[24] † Winneshiekia youngae Last Modified in 07/16/18 | 156 | 87 | 87 | 0 | 13 | 0 | 0 | 1 |
[25] † Euproops anthrax Last Modified in 07/02/14 | 98 | 135 | 135 | 0 | 23 | 0 | 0 | 1 |
[26] † Euproops mariae Last Modified in 07/17/14 | 168 | 77 | 77 | 0 | 11 | 0 | 0 | 1 |
[27] † Euproops danae Last Modified in 07/02/14 | 93 | 140 | 140 | 0 | 23 | 0 | 0 | 1 |
[28] † Euproops rotundatus Last Modified in 07/17/14 | 146 | 98 | 98 | 0 | 12 | 0 | 0 | 1 |
[29] † Euproops sp Last Modified in 07/02/14 | 91 | 138 | 138 | 0 | 27 | 0 | 0 | 1 |
[30] † Bellinurus lunatus Last Modified in 07/18/14 | 148 | 95 | 95 | 0 | 13 | 0 | 0 | 1 |
[31] † Bellinurus arcuatus Last Modified in 07/18/14 | 149 | 94 | 94 | 0 | 13 | 0 | 0 | 1 |
[32] † Bellinurus reginae Last Modified in 07/18/14 | 149 | 93 | 93 | 0 | 14 | 0 | 0 | 1 |
[33] † Bellinurus truemanii Last Modified in 07/18/14 | 150 | 92 | 92 | 0 | 14 | 0 | 0 | 1 |
[34] † Bellinurus bellulus Last Modified in 07/18/14 | 152 | 90 | 90 | 0 | 14 | 0 | 0 | 1 |
[35] † Bellinurus trilobitoides Last Modified in 07/02/14 | 149 | 93 | 93 | 0 | 14 | 0 | 0 | 1 |
[36] † Anacontium carpenteri Last Modified in 07/16/14 | 218 | 36 | 36 | 0 | 2 | 0 | 0 | 1 |
[37] † Liomesaspis laevis Last Modified in 07/16/14 | 150 | 88 | 88 | 0 | 18 | 0 | 0 | 1 |
[38] † Alanops magnifica Last Modified in 07/16/14 | 58 | 155 | 155 | 0 | 43 | 0 | 0 | 1 |
[39] † Pringlia birtwelli Last Modified in 07/17/14 | 161 | 73 | 73 | 0 | 22 | 0 | 0 | 1 |
[40] † Kasibelinurus amoricum Last Modified in 07/02/14 | 153 | 91 | 91 | 0 | 12 | 0 | 0 | 1 |
[41] † Houia yueya Last Modified in 05/25/15 | 144 | 88 | 88 | 0 | 24 | 0 | 0 | 1 |
[42] † Kasibelinurus randalli Last Modified in 07/02/14 | 164 | 81 | 81 | 0 | 11 | 0 | 0 | 1 |
[43] † Lunataspis aurora Last Modified in 07/02/14 | 142 | 96 | 96 | 0 | 18 | 0 | 0 | 1 |
[44] Limulus polyphemus Last Modified in 07/02/14 | 0 | 216 | 216 | 0 | 40 | 0 | 0 | 1 |
[45] † Limulus ? darwini Last Modified in 07/16/18 | 162 | 77 | 77 | 0 | 17 | 0 | 0 | 1 |
[46] † Limulus coffini Last Modified in 07/14/14 | 198 | 44 | 44 | 0 | 14 | 0 | 0 | 1 |
[47] † Crenatolimulus paluxyensis Last Modified in 07/14/14 | 159 | 81 | 81 | 0 | 16 | 0 | 0 | 1 |
[48] Carcinoscorpius rotundicauda Last Modified in 07/03/14 | 0 | 217 | 217 | 0 | 39 | 0 | 0 | 1 |
[49] † Heterolimulus gadeai Last Modified in 07/16/18 | 149 | 91 | 91 | 0 | 16 | 0 | 0 | 1 |
[50] Tachypleus gigas Last Modified in 07/03/14 | 0 | 217 | 217 | 0 | 39 | 0 | 0 | 1 |
[51] † Tachypleus decheni Last Modified in 07/16/18 | 148 | 92 | 92 | 0 | 16 | 0 | 0 | 1 |
[52] † Tachypleus syriacus Last Modified in 07/16/18 | 143 | 98 | 97 | 0 | 16 | 0 | 0 | 1 |
[53] Tachypleus tridentatus Last Modified in 07/02/14 | 0 | 217 | 217 | 0 | 39 | 0 | 0 | 1 |
[54] † Yunnanolimulus luopingensis Last Modified in 07/11/14 | 30 | 186 | 186 | 0 | 40 | 0 | 0 | 1 |
[55] † Xaniopyramis linseyi Last Modified in 07/10/14 | 187 | 64 | 64 | 0 | 5 | 0 | 0 | 1 |
[56] † Paleolimulus woodae Last Modified in 07/11/18 | 157 | 86 | 86 | 0 | 13 | 0 | 0 | 1 |
[57] † Paleolimulus signatus Last Modified in 07/02/14 | 46 | 180 | 180 | 0 | 30 | 0 | 0 | 1 |
[58] † Paleolimulus kunguricus Last Modified in 07/16/18 | 141 | 101 | 101 | 0 | 14 | 0 | 0 | 1 |
[59] † Bellinuroopsis rossicus Last Modified in 07/18/14 | 152 | 90 | 90 | 0 | 14 | 0 | 0 | 1 |
[60] † Rolfeia fouldenensis Last Modified in 07/10/14 | 143 | 99 | 99 | 0 | 14 | 0 | 0 | 1 |
[61] † Tarracolimulus rieki Last Modified in 07/14/14 | 154 | 87 | 87 | 0 | 15 | 0 | 0 | 1 |
[62] † Casterolimulus kletti Last Modified in 07/16/14 | 219 | 35 | 35 | 0 | 2 | 0 | 0 | 1 |
[63] † Limulitella vicensis Last Modified in 07/21/14 | 175 | 69 | 69 | 0 | 12 | 0 | 0 | 1 |
[64] † Victalimulus mcqueeni Last Modified in 07/16/14 | 116 | 122 | 122 | 0 | 18 | 0 | 0 | 1 |
[65] † Limulus woodwardi Last Modified in 07/13/18 | 220 | 34 | 34 | 0 | 2 | 0 | 0 | 1 |
[66] † Mesolimulus crespelli Last Modified in 07/15/14 | 163 | 77 | 77 | 0 | 16 | 0 | 0 | 1 |
[67] † Mesolimulus sp. Last Modified in 07/13/18 | 155 | 85 | 85 | 0 | 16 | 0 | 0 | 1 |
[68] † Mesolimulus walchi Last Modified in 07/02/14 | 57 | 170 | 170 | 0 | 29 | 0 | 0 | 1 |
[69] † Panduralimulus babcocki Last Modified in 07/11/14 | 142 | 95 | 95 | 0 | 19 | 0 | 0 | 1 |
[70] † Austrolimulus fletcheri Last Modified in 07/11/14 | 149 | 88 | 88 | 0 | 19 | 0 | 0 | 1 |
[71] † Vaderlimulus tricki Last Modified in 07/11/18 | 147 | 89 | 89 | 0 | 20 | 0 | 0 | 1 |
[72] † Psammolimulus gottingensis Last Modified in 07/11/14 | 113 | 119 | 119 | 0 | 24 | 0 | 0 | 1 |
[73] † Limulitella bronni Last Modified in 07/17/14 | 149 | 87 | 87 | 0 | 20 | 0 | 0 | 1 |
[74] † Limulitella tejraensis Last Modified in 07/11/18 | 159 | 77 | 77 | 0 | 20 | 0 | 0 | 1 |
[75] † Limulitella henkeli Last Modified in 07/17/14 | 168 | 73 | 73 | 0 | 15 | 0 | 0 | 1 |
[76] † Paleolimulus fuchsbergensis Last Modified in 07/11/14 | 156 | 81 | 81 | 0 | 19 | 0 | 0 | 1 |
[77] † Dubbolimulus peetae Last Modified in 07/16/18 | 145 | 91 | 91 | 0 | 20 | 0 | 0 | 1 |
[78] † Paleolimulus longispinus Last Modified in 07/15/14 | 136 | 98 | 98 | 0 | 22 | 0 | 0 | 1 |
[79] † Valloisella lievinensis Last Modified in 07/16/14 | 146 | 94 | 94 | 0 | 16 | 0 | 0 | 1 |
[80] † Stoermeropterus conicus Last Modified in 07/02/14 | 25 | 198 | 198 | 0 | 33 | 0 | 0 | 1 |
[81] † Moselopterus ancylotelson Last Modified in 07/02/14 | 24 | 194 | 194 | 0 | 38 | 0 | 0 | 1 |
[82] † Onychopterella kokomoensis Last Modified in 07/02/14 | 33 | 189 | 189 | 0 | 34 | 0 | 0 | 1 |
[83] † Eurypterus tetragonophthalmus Last Modified in 07/02/14 | 10 | 213 | 213 | 0 | 33 | 0 | 0 | 1 |
[84] † Erieopterus macrophthalmus Last Modified in 07/02/14 | 33 | 190 | 190 | 0 | 33 | 0 | 0 | 1 |
[85] † Strobilopterus proteus Last Modified in 07/02/14 | 47 | 186 | 186 | 0 | 23 | 0 | 0 | 1 |
[86] † Dolichopterus macrocheirus Last Modified in 07/02/14 | 22 | 201 | 201 | 0 | 33 | 0 | 0 | 1 |
[87] † Carcinosoma newlini Last Modified in 07/02/14 | 31 | 188 | 188 | 0 | 37 | 0 | 0 | 1 |
[88] † Megalograptus ohioensis Last Modified in 07/02/14 | 22 | 201 | 201 | 0 | 33 | 0 | 0 | 1 |
[89] † Mixopterus kiaeri Last Modified in 07/02/14 | 32 | 187 | 187 | 0 | 37 | 0 | 0 | 1 |
[90] † Orcanopterus manitoulinensis Last Modified in 07/02/14 | 73 | 145 | 145 | 0 | 38 | 0 | 0 | 1 |
[91] † Adelophthalmus sievertsi Last Modified in 07/02/14 | 46 | 174 | 174 | 0 | 36 | 0 | 0 | 1 |
[92] † Nanahughmilleria norvegica Last Modified in 07/02/14 | 24 | 198 | 198 | 0 | 34 | 0 | 0 | 1 |
[93] † Hughmilleria socialis Last Modified in 07/02/14 | 23 | 195 | 195 | 0 | 38 | 0 | 0 | 1 |
[94] † Slimonia acuminata Last Modified in 07/02/14 | 25 | 193 | 193 | 0 | 38 | 0 | 0 | 1 |
[95] † Erettopterus bilobus Last Modified in 07/02/14 | 25 | 191 | 191 | 0 | 40 | 0 | 0 | 1 |
[96] † Pterygotus anglicus Last Modified in 07/02/14 | 31 | 184 | 184 | 0 | 41 | 0 | 0 | 1 |
[97] † Brachyopterus stubblefieldi Last Modified in 07/02/14 | 71 | 150 | 150 | 0 | 35 | 0 | 0 | 1 |
[98] † Rhenopterus diensti Last Modified in 07/02/14 | 24 | 198 | 198 | 0 | 34 | 0 | 0 | 1 |
[99] † Parastylonurus ornatus Last Modified in 07/02/14 | 34 | 188 | 188 | 0 | 34 | 0 | 0 | 1 |
[100] † Laurieipterus elegans Last Modified in 07/02/14 | 94 | 128 | 128 | 0 | 34 | 0 | 0 | 1 |
[101] † Kokomopterus longicaudatus Last Modified in 07/02/14 | 44 | 177 | 177 | 0 | 35 | 0 | 0 | 1 |
[102] † Hardieopterus macrophthalmus Last Modified in 07/02/14 | 74 | 150 | 150 | 0 | 32 | 0 | 0 | 1 |
[103] † Drepanopterus pentlandicus Last Modified in 07/02/14 | 28 | 198 | 197 | 0 | 31 | 0 | 0 | 1 |
[104] † Hibbertopterus scouleri Last Modified in 07/02/14 | 30 | 195 | 195 | 0 | 31 | 0 | 0 | 1 |
[105] † Megarachne servinei Last Modified in 07/02/14 | 173 | 62 | 61 | 0 | 22 | 0 | 0 | 1 |
[106] † Chasmataspis laurencii Last Modified in 07/02/14 | 112 | 121 | 121 | 0 | 23 | 0 | 0 | 1 |
[107] † Loganamaraspis dunlopi Last Modified in 07/02/14 | 96 | 129 | 129 | 0 | 31 | 0 | 0 | 1 |
[108] † Achanarraspis reedi Last Modified in 07/02/14 | 153 | 85 | 85 | 0 | 18 | 0 | 0 | 1 |
[109] † Heteroaspis stoermeri Last Modified in 07/02/14 | 144 | 90 | 90 | 0 | 22 | 0 | 0 | 1 |
[110] † Diploaspis casteri Last Modified in 07/02/14 | 106 | 125 | 125 | 0 | 25 | 0 | 0 | 1 |
[111] † Diploaspis muelleri Last Modified in 07/02/14 | 148 | 86 | 86 | 0 | 22 | 0 | 0 | 1 |
[112] † Dvulikiaspis menneri Last Modified in 07/02/14 | 122 | 105 | 105 | 0 | 29 | 0 | 0 | 1 |
[113] † Octoberaspis ushakovi Last Modified in 07/02/14 | 64 | 160 | 160 | 0 | 32 | 0 | 0 | 1 |
[114] Centruroides vittatus Last Modified in 07/02/14 | 0 | 198 | 198 | 0 | 58 | 0 | 0 | 1 |
[115] Androctonus australis Last Modified in 07/02/14 | 1 | 198 | 198 | 0 | 57 | 0 | 0 | 1 |
[116] † Proscorpius osborni Last Modified in 07/02/14 | 29 | 168 | 168 | 0 | 59 | 0 | 0 | 1 |
[117] † Palaeophonus nuncius Last Modified in 07/02/14 | 54 | 145 | 145 | 0 | 57 | 0 | 0 | 1 |
[118] Caddo agilis Last Modified in 07/02/14 | 7 | 177 | 177 | 0 | 72 | 0 | 0 | 1 |
[119] Chileogovea oedipus Last Modified in 07/02/14 | 14 | 173 | 173 | 0 | 69 | 0 | 0 | 1 |
[120] Cyphophthalmus duricorius Last Modified in 07/02/14 | 15 | 173 | 173 | 0 | 68 | 0 | 0 | 1 |
[121] Gonyleptes horridus Last Modified in 07/02/14 | 8 | 178 | 178 | 0 | 70 | 0 | 0 | 1 |
[122] Leiobunum rotundum Last Modified in 07/02/14 | 8 | 177 | 177 | 0 | 71 | 0 | 0 | 1 |
[123] Sclerobunus robustus Last Modified in 07/02/14 | 8 | 178 | 178 | 0 | 70 | 0 | 0 | 1 |
[124] Allothrombium simoni Last Modified in 07/02/14 | 17 | 160 | 159 | 0 | 80 | 0 | 0 | 1 |
[125] Amblyomma americanum Last Modified in 07/02/14 | 15 | 158 | 157 | 0 | 84 | 0 | 0 | 1 |
[126] Neocarus texanus Last Modified in 07/02/14 | 9 | 167 | 167 | 0 | 80 | 0 | 0 | 1 |
[127] Cryptocellus goodnighti Last Modified in 07/02/14 | 5 | 180 | 180 | 0 | 71 | 0 | 0 | 1 |
[128] Poliochera punctulata Last Modified in 07/02/14 | 60 | 126 | 126 | 0 | 70 | 0 | 0 | 1 |
[129] Ricinoides atewa Last Modified in 07/02/14 | 5 | 180 | 180 | 0 | 71 | 0 | 0 | 1 |
[130] † Anthracomartus hindi Last Modified in 07/02/14 | 32 | 157 | 157 | 0 | 67 | 0 | 0 | 1 |
[131] † Eophrynus prestvicii Last Modified in 07/02/14 | 33 | 159 | 159 | 0 | 64 | 0 | 0 | 1 |
[132] † Gilboarachne griersoni Last Modified in 07/02/14 | 21 | 167 | 167 | 0 | 68 | 0 | 0 | 1 |
[133] † Palaeocharinus sp. Last Modified in 07/02/14 | 9 | 179 | 179 | 0 | 68 | 0 | 0 | 1 |
[134] † Mesotarbus peteri Last Modified in 07/02/14 | 52 | 136 | 136 | 0 | 68 | 0 | 0 | 1 |
[135] † Plesiosiro madeleyi Last Modified in 07/02/14 | 44 | 145 | 145 | 0 | 67 | 0 | 0 | 1 |
[136] Eukoenenia mirabilis Last Modified in 07/02/14 | 4 | 181 | 180 | 0 | 72 | 0 | 0 | 1 |
[137] Prokoenenia wheeleri Last Modified in 07/02/14 | 4 | 181 | 180 | 0 | 72 | 0 | 0 | 1 |
[138] Mastigoproctus giganteus Last Modified in 07/02/14 | 4 | 190 | 190 | 0 | 62 | 0 | 0 | 1 |
[139] Thelyphonus caudatus Last Modified in 07/02/14 | 4 | 186 | 186 | 0 | 66 | 0 | 0 | 1 |
[140] Hubbardia pentapeltis Last Modified in 07/02/14 | 4 | 186 | 185 | 0 | 67 | 0 | 0 | 1 |
[141] Schizomus crassicaudatus Last Modified in 07/02/14 | 4 | 186 | 185 | 0 | 67 | 0 | 0 | 1 |
[142] † Permarachne novokshonovi Last Modified in 07/02/14 | 116 | 101 | 101 | 0 | 39 | 0 | 0 | 1 |
[143] † Attercopus filmbriunguis Last Modified in 07/02/14 | 170 | 68 | 68 | 0 | 18 | 0 | 0 | 1 |
[144] Liphistius malayanus Last Modified in 07/02/14 | 4 | 175 | 175 | 0 | 77 | 0 | 0 | 1 |
[145] Aphonopelma hentzi Last Modified in 07/02/14 | 4 | 166 | 166 | 0 | 86 | 0 | 0 | 1 |
[146] Hypochilus pococki Last Modified in 07/02/14 | 4 | 166 | 166 | 0 | 86 | 0 | 0 | 1 |
[147] Cupienius getazi Last Modified in 07/02/14 | 4 | 165 | 165 | 0 | 87 | 0 | 0 | 1 |
[148] Neobisium maritimum Last Modified in 07/02/14 | 11 | 166 | 165 | 0 | 80 | 0 | 0 | 1 |
[149] Chelifer cancroides Last Modified in 07/02/14 | 11 | 165 | 164 | 0 | 81 | 0 | 0 | 1 |
[150] Eremocosta striata Last Modified in 07/02/14 | 11 | 162 | 161 | 0 | 84 | 0 | 0 | 1 |
[151] Galeodes armeniacus Last Modified in 07/02/14 | 13 | 161 | 160 | 0 | 83 | 0 | 0 | 1 |
[152] Phrynus marginemaculatus Last Modified in 07/02/14 | 5 | 182 | 182 | 0 | 69 | 0 | 0 | 1 |
[153] Charinus victori Last Modified in 07/02/14 | 6 | 181 | 181 | 0 | 69 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 12156 | |
Project overview | 1067 | |
Media views | 1874 | Media search (1624 views); M782936 (250 views); |
Matrix views | 506 | Xiph focus (79 views); Matrix landing page (427 views); |
Views for media list | 572 | |
Specimen list | 1607 | |
Taxon list | 6129 | |
Bibliography | 397 | |
Documents list | 4 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 57 | |
Project downloads | 56 | |
Matrix downloads | 1 | Xiph focus (1 download); |