Project 3200: M. R. Chani-Posse, A. J. Brunke, S. Chatzimanolis, H. Schillhammer, A. Solodovnikov. 2018. Phylogeny of the hyper-diverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics. 34 (1):1-40.
Abstract
With 71 genera and over 2700 described species, Philonthina is the most speciose subtribe of rove beetle tribe Staphylinini and forms a major component of the largest remaining higher systematics challenge in Staphylinini, the 'Staphylinini propria' clade. A related systematics issue concerns the position of the genus Holisus (Hyptiomina), which was recovered within the Neotropical philonthine lineage in several recent analyses of morphology. With the aims of resolving the phylogeny of Philonthina and the position and, thus, validity of Hyptiomina, we performed phylogenetic analyses of the tribe Staphylinini based on molecular (six genes, 4471 bp) and morphological (113 characters) data including 138 taxa from all relevant lineages of Staphylinini. We found that 'Staphylinini propria' is a monophylum consisting of six lineages: current subtribes Anisolinina, Philonthina, Staphylinina and Xanthopygina; and two new subtribes, Algonina Schillhammer and Brunke and Philothalpina Chatzimanolis and Brunke. While the previously hypothesized Neotropical lineage of Philonthina was corroborated, Holisus was recovered as a separate subtribe, outside of Philonthina, within an informal 'Southern Hemisphere clade'. Based on our analyses, we propose tentative new concepts of the polyphyletic genera Belonuchus and Philonthus. We propose the following taxonomic changes: synonymy of the subtribes Staphylinina Latreille (valid name) and Eucibdelina Sharp; resurrection of genera Barypalpus Cameron and Trapeziderus Motschulsky from synonymy with Rientis Sharp and Belonuchus Nordmann, respectively; transfer of 38 Belonuchus species, 16 Hesperus Fauvel species and one Philonthus Stephens species to Trapeziderus as new combinations; transfer of two Hesperus species to Eccoptolonthus Bernhauer as new combinations; transfer of one Belonuchus species to Paederomimus Sharp as a new combination; and transfer of Pridonius Blackwelder new status from its position as a subgenus of Quedius (subtribe Quediina) to Philonthina as a genus, and new combinations for its two described species.Read the article »
Article DOI: 10.1111/cla.12188
Project DOI: 10.7934/P3200, http://dx.doi.org/10.7934/P3200
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 14934 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 113 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 269 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:113/0 |
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MorphoBank Project 3200
MorphoBank Project 3200
- Creation Date:
01 June 2018 - Publication Date:
12 June 2018 - Project views: 19803
- Matrix downloads: 5
Authors' Institutions
- University of Tennessee
- Consejo Nacional de Investigaciones CientÃficas y Técnicas (CONICET)
- Natural History Museum, Vienna (Naturhistorisches Museum, Wien)
- Natural History Museum of Denmark
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 138 | 1 | 1 | 113 | 0 | 0 | 14934 (14054, 0, 880) | 0 | 0 | 0 | ||||
MorphoBank Curator Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M25487' (138 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Gabrius nr. endeius nitidipennis Last Modified in 06/12/18 | 1 | 105 | 105 | 0 | 7 | 0 | 0 | 2 |
[2] Gabrius sp. Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[3] Gabrius picipennis Last Modified in 06/12/18 | 2 | 104 | 104 | 0 | 7 | 0 | 0 | 2 |
[4] Gabrius keysianus Last Modified in 06/12/18 | 11 | 98 | 98 | 0 | 4 | 0 | 0 | 2 |
[5] Gabrius ovaliceps Last Modified in 06/12/18 | 0 | 107 | 107 | 0 | 6 | 0 | 0 | 2 |
[6] Bisnius blandus Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[7] Gabrius brevipennis Last Modified in 06/12/18 | 2 | 106 | 106 | 0 | 5 | 0 | 0 | 2 |
[8] Bisnius picicornis Last Modified in 06/12/18 | 1 | 106 | 106 | 0 | 6 | 0 | 0 | 2 |
[9] Gabrius trossuloides Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[10] Belonuchus pollens Last Modified in 06/12/18 | 10 | 99 | 99 | 0 | 4 | 0 | 0 | 2 |
[11] Belonuchus rufipennis Last Modified in 06/12/18 | 0 | 109 | 109 | 0 | 4 | 0 | 0 | 2 |
[12] Belonuchus rufipennis group Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[13] Belonuchus sp. 4 Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[14] Genus nr. linoderus Last Modified in 06/12/18 | 0 | 110 | 110 | 0 | 3 | 0 | 0 | 2 |
[15] Belonunuchus subaeneus Last Modified in 06/12/18 | 1 | 106 | 106 | 0 | 6 | 0 | 0 | 2 |
[16] Linoderus sp. undescribed Last Modified in 06/12/18 | 0 | 109 | 109 | 0 | 4 | 0 | 0 | 2 |
[17] Paederomimus nobilis Last Modified in 06/12/18 | 0 | 107 | 107 | 0 | 6 | 0 | 0 | 2 |
[18] Paederomimus pectoralis Last Modified in 06/12/18 | 3 | 106 | 106 | 0 | 4 | 0 | 0 | 2 |
[19] Paderomimus angularius group Last Modified in 06/12/18 | 11 | 97 | 97 | 0 | 5 | 0 | 0 | 2 |
[20] Paederomimus cribratus Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[21] Paederomimus lustrator group Last Modified in 06/12/18 | 3 | 105 | 105 | 0 | 5 | 0 | 0 | 2 |
[22] Odontolinus fasciatus Last Modified in 06/12/18 | 0 | 110 | 110 | 0 | 3 | 0 | 0 | 2 |
[23] Paederomimus capitalis group Last Modified in 06/12/18 | 10 | 99 | 99 | 0 | 4 | 0 | 0 | 2 |
[24] Belonuchus mimeticus Last Modified in 06/12/18 | 10 | 98 | 98 | 0 | 5 | 0 | 0 | 2 |
[25] Chroaptomus flagrans Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[26] Chroaptomus centralis Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[27] Cafius seriatus group Last Modified in 06/12/18 | 3 | 102 | 102 | 0 | 8 | 0 | 0 | 2 |
[28] Remus serriceus Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[29] Cafius xantholoma Last Modified in 06/12/18 | 0 | 107 | 107 | 0 | 6 | 0 | 0 | 2 |
[30] Cafius canescens Last Modified in 06/12/18 | 10 | 97 | 97 | 0 | 6 | 0 | 0 | 2 |
[31] Cafius seminitens Last Modified in 06/12/18 | 10 | 97 | 97 | 0 | 6 | 0 | 0 | 2 |
[32] Cafius quadriimpressus Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[33] Thinocafius insularis Last Modified in 06/12/18 | 1 | 104 | 104 | 0 | 8 | 0 | 0 | 2 |
[34] Cafius sabulosus Last Modified in 06/12/18 | 3 | 103 | 103 | 0 | 7 | 0 | 0 | 2 |
[35] Cafius sulcatus Last Modified in 06/12/18 | 10 | 97 | 97 | 0 | 6 | 0 | 0 | 2 |
[36] Philonthus marginatus Last Modified in 06/12/18 | 3 | 105 | 105 | 0 | 5 | 0 | 0 | 2 |
[37] Philonthus nitidus Last Modified in 06/12/18 | 0 | 107 | 107 | 0 | 6 | 0 | 0 | 2 |
[38] Philonthus splendens Last Modified in 06/12/18 | 0 | 107 | 107 | 0 | 6 | 0 | 0 | 2 |
[39] Philonthus caeruleipennis Last Modified in 06/12/18 | 0 | 107 | 107 | 0 | 6 | 0 | 0 | 2 |
[40] Philonthus figulus Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[41] Philonthus longicornis Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[42] Belonuchus godmani group Last Modified in 06/12/18 | 13 | 96 | 96 | 0 | 4 | 0 | 0 | 2 |
[43] Philonthus laetipennis Last Modified in 06/12/18 | 1 | 108 | 108 | 0 | 4 | 0 | 0 | 2 |
[44] Philonthus fulgipennis Last Modified in 06/12/18 | 0 | 109 | 109 | 0 | 4 | 0 | 0 | 2 |
[45] Belonuchus laevipennis group Last Modified in 06/12/18 | 0 | 105 | 105 | 0 | 8 | 0 | 0 | 2 |
[46] Belonuchus sp. 6 Last Modified in 06/12/18 | 1 | 105 | 105 | 0 | 7 | 0 | 0 | 2 |
[47] Belonuchus sp. 2 Last Modified in 06/12/18 | 7 | 103 | 103 | 0 | 3 | 0 | 0 | 2 |
[48] Proxenobius borgmeieri Last Modified in 06/12/18 | 1 | 106 | 106 | 0 | 6 | 0 | 0 | 2 |
[49] Termitoquedius sp. undescribed Last Modified in 06/12/18 | 2 | 105 | 105 | 0 | 6 | 0 | 0 | 2 |
[50] Bisnius sordidus Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[51] Gabronthus flavicollis Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[52] Neobisnius senilis Last Modified in 06/12/18 | 13 | 95 | 95 | 0 | 5 | 0 | 0 | 2 |
[53] Neobisnius occidentoides Last Modified in 06/12/18 | 113 | 0 | 0 | 0 | 0 | 0 | 0 | 2 |
[54] Neobisnius terminalis Last Modified in 06/12/18 | 3 | 103 | 103 | 0 | 7 | 0 | 0 | 2 |
[55] Philonthus atratus Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[56] Philonthus carbonarius Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[57] Philonthus sp. Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[58] Rabigus laxellus Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[59] Philonthus fuscatus Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[60] Philonthus paederoides Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[61] Philonthus turbidus Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[62] Philonthus quadricollis Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[63] Philonthus rectangulus Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[64] Philonthus rufotibialis Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[65] Taxiplagus sp. nov. Last Modified in 06/12/18 | 10 | 97 | 97 | 0 | 6 | 0 | 0 | 2 |
[66] Philonthus sericans Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[67] Philonthus cunctans Last Modified in 06/12/18 | 1 | 107 | 107 | 0 | 5 | 0 | 0 | 2 |
[68] Belonuchus sp. 5 Last Modified in 06/12/18 | 21 | 86 | 86 | 0 | 6 | 0 | 0 | 2 |
[69] Belonuchus imitator Last Modified in 06/12/18 | 0 | 105 | 105 | 0 | 8 | 0 | 0 | 2 |
[70] Hesperus apicialis Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[71] Hesperus baltimorensis Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[72] Hesperus sp. nov. Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[73] Hesperus borneensis Last Modified in 06/12/18 | 3 | 103 | 103 | 0 | 7 | 0 | 0 | 2 |
[74] Hybridolinus sp. undescribed Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[75] Hesperopalpus sp. Last Modified in 06/12/18 | 1 | 105 | 105 | 0 | 7 | 0 | 0 | 2 |
[76] Misantlius gebieni Last Modified in 06/12/18 | 0 | 107 | 107 | 0 | 6 | 0 | 0 | 2 |
[77] Hesperosoma pederseni Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[78] Tasgius pedator Last Modified in 06/12/18 | 0 | 107 | 107 | 0 | 6 | 0 | 0 | 2 |
[79] Rhynchocheilus sp. Last Modified in 06/12/18 | 0 | 105 | 105 | 0 | 8 | 0 | 0 | 2 |
[80] Ontholestes murinus Last Modified in 06/12/18 | 0 | 107 | 107 | 0 | 6 | 0 | 0 | 2 |
[81] Tolmerinus sp. (laos) Last Modified in 06/12/18 | 0 | 107 | 107 | 0 | 6 | 0 | 0 | 2 |
[82] Creophilus maxillosus Last Modified in 06/12/18 | 0 | 105 | 105 | 0 | 8 | 0 | 0 | 2 |
[83] Thinopinus pictus Last Modified in 06/12/18 | 1 | 101 | 101 | 0 | 11 | 0 | 0 | 2 |
[84] Platydracus cinnamopterus Last Modified in 06/12/18 | 0 | 107 | 107 | 0 | 6 | 0 | 0 | 2 |
[85] Anisolinus sp. Last Modified in 06/12/18 | 5 | 102 | 102 | 0 | 6 | 0 | 0 | 2 |
[86] Philothalpus bilobus Last Modified in 06/12/18 | 1 | 106 | 106 | 0 | 6 | 0 | 0 | 2 |
[87] Philothalpus falini Last Modified in 06/12/18 | 0 | 108 | 108 | 0 | 5 | 0 | 0 | 2 |
[88] Tympanophorus sp. (laos) Last Modified in 06/12/18 | 0 | 107 | 107 | 0 | 6 | 0 | 0 | 2 |
[89] Eccoptolonthus sp. Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[90] Dinothenarus saphyrinus Last Modified in 06/12/18 | 0 | 107 | 107 | 0 | 6 | 0 | 0 | 2 |
[91] Gastrisus sp. Last Modified in 06/12/18 | 5 | 102 | 102 | 0 | 6 | 0 | 0 | 2 |
[92] Xanthopygus chapareanus Last Modified in 06/12/18 | 2 | 104 | 104 | 0 | 7 | 0 | 0 | 2 |
[93] Phanolinus sp. Last Modified in 06/12/18 | 2 | 104 | 104 | 0 | 7 | 0 | 0 | 2 |
[94] Isanopus sallaei Last Modified in 06/12/18 | 0 | 104 | 104 | 0 | 9 | 0 | 0 | 2 |
[95] Glenus flohri Last Modified in 06/12/18 | 5 | 102 | 102 | 0 | 6 | 0 | 0 | 2 |
[96] Hemiquedius sp. Last Modified in 06/12/18 | 2 | 104 | 104 | 0 | 7 | 0 | 0 | 2 |
[97] Anaquedius vernix Last Modified in 06/12/18 | 2 | 104 | 104 | 0 | 7 | 0 | 0 | 2 |
[98] Quedionuchus plagiatus Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[99] Quedius beesoni Last Modified in 06/12/18 | 2 | 104 | 104 | 0 | 7 | 0 | 0 | 2 |
[100] Quedius simplicifrons Last Modified in 06/12/18 | 2 | 104 | 104 | 0 | 7 | 0 | 0 | 2 |
[101] Plociopterus sp. Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[102] Indoquedius bicornutus Last Modified in 06/12/18 | 2 | 102 | 102 | 0 | 9 | 0 | 0 | 2 |
[103] Indoquedius sp. Last Modified in 06/12/18 | 2 | 102 | 102 | 0 | 9 | 0 | 0 | 2 |
[104] Cyrtoquedius labiatus Last Modified in 06/12/18 | 2 | 102 | 102 | 0 | 9 | 0 | 0 | 2 |
[105] Antimerus punctipennis Last Modified in 06/12/18 | 0 | 101 | 101 | 0 | 12 | 0 | 0 | 2 |
[106] Hesperus roepki Last Modified in 06/12/18 | 0 | 105 | 105 | 0 | 8 | 0 | 0 | 2 |
[107] Hesperus laevigatus Last Modified in 06/12/18 | 0 | 105 | 105 | 0 | 8 | 0 | 0 | 2 |
[108] Hesperus amabilis Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[109] Hesperu haemorrhoidalis Last Modified in 06/12/18 | 3 | 103 | 103 | 0 | 7 | 0 | 0 | 2 |
[110] Hesperus semirufus Last Modified in 06/12/18 | 3 | 103 | 103 | 0 | 7 | 0 | 0 | 2 |
[111] Actinus sp. Last Modified in 06/12/18 | 3 | 103 | 103 | 0 | 7 | 0 | 0 | 2 |
[112] Leucitus sp. Last Modified in 06/12/18 | 5 | 100 | 100 | 0 | 8 | 0 | 0 | 2 |
[113] Erichsonius nanus Last Modified in 06/12/18 | 0 | 105 | 105 | 0 | 8 | 0 | 0 | 2 |
[114] Erichsonius sp. (laos) Last Modified in 06/12/18 | 0 | 105 | 105 | 0 | 8 | 0 | 0 | 2 |
[115] Pammegus ruficollis Last Modified in 06/12/18 | 1 | 106 | 106 | 0 | 6 | 0 | 0 | 2 |
[116] Bolitogyrus bullatus Last Modified in 06/12/18 | 2 | 102 | 102 | 0 | 9 | 0 | 0 | 2 |
[117] Elmas hanleyi Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[118] Nordus fungicola Last Modified in 06/12/18 | 0 | 105 | 105 | 0 | 8 | 0 | 0 | 2 |
[119] Algon hollowayae Last Modified in 06/12/18 | 6 | 101 | 101 | 0 | 6 | 0 | 0 | 2 |
[120] Algon oculatus group Last Modified in 06/12/18 | 6 | 101 | 101 | 0 | 6 | 0 | 0 | 2 |
[121] Atanygnathus acuminatus Last Modified in 06/12/18 | 113 | 0 | 0 | 0 | 0 | 0 | 0 | 2 |
[122] Atanygnathus sp. (laos) Last Modified in 06/12/18 | 4 | 100 | 100 | 0 | 9 | 0 | 0 | 2 |
[123] Holisus planus group Last Modified in 06/12/18 | 18 | 84 | 84 | 0 | 11 | 0 | 0 | 2 |
[124] Holisus prolixus Last Modified in 06/12/18 | 18 | 84 | 84 | 0 | 11 | 0 | 0 | 2 |
[125] Holisus sp. (costa rica) Last Modified in 06/12/18 | 113 | 0 | 0 | 0 | 0 | 0 | 0 | 2 |
[126] Rientis sp. undescribed Last Modified in 06/12/18 | 0 | 107 | 107 | 0 | 6 | 0 | 0 | 2 |
[127] Arrowinus minutus Last Modified in 06/12/18 | 0 | 99 | 99 | 0 | 14 | 0 | 0 | 2 |
[128] Barypalpus ruficornis Last Modified in 06/12/18 | 2 | 104 | 104 | 0 | 7 | 0 | 0 | 2 |
[129] Genus nr. quedius iheringhi Last Modified in 06/12/18 | 10 | 95 | 95 | 0 | 8 | 0 | 0 | 2 |
[130] Afroquedius sexpunctatus Last Modified in 06/12/18 | 0 | 106 | 106 | 0 | 7 | 0 | 0 | 2 |
[131] Heterothops sp. (costa rica) Last Modified in 06/12/18 | 1 | 101 | 101 | 0 | 11 | 0 | 0 | 2 |
[132] Amblyopinus emarginatus Last Modified in 06/12/18 | 2 | 102 | 102 | 0 | 9 | 0 | 0 | 2 |
[133] Quedius antipodum Last Modified in 06/12/18 | 0 | 105 | 105 | 0 | 8 | 0 | 0 | 2 |
[134] Atanygnatus sp. (south africa) Last Modified in 06/12/18 | 4 | 101 | 101 | 0 | 8 | 0 | 0 | 2 |
[135] Holisus sp. (ecuador) Last Modified in 06/12/18 | 4 | 98 | 98 | 0 | 11 | 0 | 0 | 2 |
[136] Quedius iheringhi Last Modified in 06/12/18 | 10 | 94 | 94 | 0 | 9 | 0 | 0 | 2 |
[137] Cheilocolpus pyrostomus Last Modified in 06/12/18 | 0 | 104 | 104 | 0 | 9 | 0 | 0 | 2 |
[138] Belonuchus haemorrhoidalis Last Modified in 06/12/18 | 2 | 107 | 107 | 0 | 4 | 0 | 0 | 2 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 19803 | |
Project overview | 1760 | |
Media views | 3262 | Media search (2750 views); M584444 (512 views); |
Matrix views | 1088 | Matrix landing page (936 views); Morphological matrix from supplemental data (152 views); |
Taxon list | 10196 | |
Views for media list | 992 | |
Bibliography | 732 | |
Specimen list | 1764 | |
Documents list | 9 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 175 | |
Project downloads | 170 | |
Matrix downloads | 5 | Morphological matrix from supplemental data (5 downloads); |