Project 3587: C. Olivier, B. Battail, S. Bourquin, C. Rossignol, J. Steyer, N. Jalil. 2019. New dicynodonts (Therapsida, Anomodontia) from near the Permo-Triassic boundary of Laos: implications for dicynodont survivorship across the Permo-Triassic mass extinction and the paleobiogeography of Southeast Asian blocks. Journal of Vertebrate Paleontology. 39 (2):e1584745.
Abstract
The dicynodonts are an emblematic group of herbivorous therapsids that survived the Permo-Triassic (P-Tr) crisis. Laotian dicynodonts from stratigraphically constrained beds, recently dated using the U-Pb zircon method, yield new insights into terrestrial faunas of Southeast Asia during the latest Permian and earliest Triassic. Summarily described, they were originally attributed to the genus Dicynodon. We provide a new phylogenetic analysis for Laotian dicynodonts, based on three well-preserved skulls, indicating that they belong to two new taxa: Counillonia superoculis, gen. et sp. nov., and Repelinosaurus robustus, gen. et sp. nov. Our phylogenetic analysis of Dicynodontia indicates that (1) Counillonia is closely related to some ‘Dicynodon'-grade taxa and (2) Repelinosaurus is a kannemeyeriiform. The phylogenetic affinities of these new Laotian dicynodonts allow discussion of the survivorship of multiple lineages (Kannemeyeriiformes and ‘Dicynodon'grade dicynodontoids) across the P-Tr crisis. The Laotian dicynodonts also shed new light on the paleobiogeography of Southeast Asia from the late Paleozoic to the early Mesozoic, particularly the timing of collisions between the Indochina, the South China, and the North China blocks. The presence of dicynodonts in Laos, most likely in the Early Triassic, thus implies that the connection between the Indochina Block and the South China Block occurred no later than the latest Permian or earliest Triassic (i.e., when the dicynodonts provide direct evidence for a connection).Read the article »
Article DOI: 10.1080/02724634.2019.1584745
Project DOI: 10.7934/P3587, http://dx.doi.org/10.7934/P3587
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 14436 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 194 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 424 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:1/193 |
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MorphoBank Project 3587
MorphoBank Project 3587
- Creation Date:
15 November 2019 - Publication Date:
15 January 2020 - Project views: 17450
- Matrix downloads: 2
Authors' Institutions
- Muséum national d'Histoire naturelle, Paris (Natural History Museum, Paris)
- Centre National de la Recherche Scientifique (CNRS)
- University of Rennes
- Universidade de Sao Paulo (USP, University of Sao Paulo)
- Sorbonne Université
- Université Cadi Ayyad
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Ana Andruchow-Colombo Project Administrator | 106 | 0 | 0 | 194 | 0 | 0 | 14436 (12941, 0, 1495) | 0 | 0 | 0 | ||||
Chloe Olivier Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M26359' (106 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Biarmosuchus Last Modified in 01/15/20 | 33 | 150 | 150 | 0 | 11 | 0 | 0 | 2 |
[2] † Hipposaurus boonstrai Last Modified in 01/15/20 | 33 | 148 | 148 | 0 | 13 | 0 | 0 | 2 |
[3] † Archaeosyodon praeventor Last Modified in 01/15/20 | 85 | 106 | 106 | 0 | 3 | 0 | 0 | 2 |
[4] † Titanophoneus potens Last Modified in 01/15/20 | 22 | 159 | 158 | 0 | 14 | 0 | 0 | 2 |
[5] † Gorgonops torvus Last Modified in 01/15/20 | 25 | 161 | 161 | 0 | 8 | 0 | 0 | 2 |
[6] † Lycosuchus vanderrieti Last Modified in 01/15/20 | 27 | 153 | 153 | 0 | 14 | 0 | 0 | 2 |
[7] † Glanosuchus macrops Last Modified in 01/15/20 | 18 | 161 | 161 | 0 | 15 | 0 | 0 | 2 |
[8] † Biseridens qilianicus Last Modified in 01/15/20 | 75 | 115 | 115 | 0 | 4 | 0 | 0 | 2 |
[9] † Anomocephalus africanus Last Modified in 01/15/20 | 166 | 28 | 28 | 0 | 0 | 0 | 0 | 2 |
[10] † Tiarajudens eccentricus Last Modified in 01/15/20 | 139 | 53 | 53 | 0 | 2 | 0 | 0 | 2 |
[11] † Otsheria netzvetajevi Last Modified in 01/15/20 | 97 | 86 | 86 | 0 | 11 | 0 | 0 | 2 |
[12] † Ulemica Last Modified in 01/15/20 | 55 | 126 | 125 | 0 | 14 | 0 | 0 | 2 |
[13] † Suminia getmanovi Last Modified in 01/15/20 | 22 | 152 | 152 | 0 | 20 | 0 | 0 | 2 |
[14] † Patranomodon nyaphulii Last Modified in 01/15/20 | 51 | 129 | 129 | 0 | 14 | 0 | 0 | 2 |
[15] † Galeops whaitsi Last Modified in 01/15/20 | 63 | 117 | 117 | 0 | 14 | 0 | 0 | 2 |
[16] † Galepus jouberti Last Modified in 01/15/20 | 151 | 40 | 40 | 0 | 3 | 0 | 0 | 2 |
[17] † Galechirus scholtzi Last Modified in 01/15/20 | 132 | 57 | 57 | 0 | 5 | 0 | 0 | 2 |
[18] † Eodicynodon oelofseni Last Modified in 01/15/20 | 94 | 93 | 93 | 0 | 7 | 0 | 0 | 2 |
[19] † Eodicynodon oosthuizeni Last Modified in 01/15/20 | 6 | 166 | 166 | 0 | 22 | 0 | 0 | 2 |
[20] † Colobodectes cluveri Last Modified in 01/15/20 | 79 | 104 | 104 | 0 | 11 | 0 | 0 | 2 |
[21] † Lanthanostegus mohoii Last Modified in 01/15/20 | 133 | 59 | 59 | 0 | 2 | 0 | 0 | 2 |
[22] † Eosimops newtoni Last Modified in 01/15/20 | 18 | 158 | 156 | 0 | 20 | 0 | 0 | 2 |
[23] † Prosictodon dubei Last Modified in 01/15/20 | 53 | 126 | 126 | 0 | 15 | 0 | 0 | 2 |
[24] † Diictodon feliceps Last Modified in 01/15/20 | 8 | 167 | 164 | 0 | 22 | 0 | 0 | 2 |
[25] † Robertia broomiana Last Modified in 01/15/20 | 14 | 160 | 160 | 0 | 20 | 0 | 0 | 2 |
[26] † Pristerodon mackayi Last Modified in 01/15/20 | 7 | 165 | 165 | 0 | 22 | 0 | 0 | 2 |
[27] † Brachyprosopus broomi Last Modified in 01/15/20 | 23 | 152 | 152 | 0 | 19 | 0 | 0 | 2 |
[28] † Endothiodon bathystoma Last Modified in 01/15/20 | 15 | 160 | 157 | 0 | 22 | 0 | 0 | 2 |
[29] † Endothiodon tolani Last Modified in 01/15/20 | 61 | 124 | 123 | 0 | 10 | 0 | 0 | 2 |
[30] † Niassodon mfumukasi Last Modified in 01/15/20 | 54 | 130 | 130 | 0 | 10 | 0 | 0 | 2 |
[31] † Digalodon rubidgei Last Modified in 01/15/20 | 63 | 120 | 120 | 0 | 11 | 0 | 0 | 2 |
[32] † Emydops Last Modified in 01/15/20 | 8 | 165 | 164 | 0 | 22 | 0 | 0 | 2 |
[33] † Dicynodontoides Last Modified in 01/15/20 | 10 | 163 | 163 | 0 | 21 | 0 | 0 | 2 |
[34] † Kombuisia frerensis Last Modified in 01/15/20 | 65 | 119 | 119 | 0 | 10 | 0 | 0 | 2 |
[35] † Myosaurus gracilis Last Modified in 01/15/20 | 30 | 146 | 146 | 0 | 18 | 0 | 0 | 2 |
[36] † Sauroscaptor tharavati Last Modified in 01/15/20 | 91 | 94 | 94 | 0 | 9 | 0 | 0 | 2 |
[37] † Cistecephalus microrhinus Last Modified in 01/15/20 | 13 | 162 | 160 | 0 | 21 | 0 | 0 | 2 |
[38] † Cistecephaloides boonstrai Last Modified in 01/15/20 | 60 | 121 | 121 | 0 | 13 | 0 | 0 | 2 |
[39] † Kawingasaurus fossilis Last Modified in 01/15/20 | 42 | 136 | 136 | 0 | 16 | 0 | 0 | 2 |
[40] † Rhachiocephalus magnus Last Modified in 01/15/20 | 22 | 154 | 153 | 0 | 19 | 0 | 0 | 2 |
[41] † Kitchinganomodon crassus Last Modified in 01/15/20 | 43 | 134 | 134 | 0 | 17 | 0 | 0 | 2 |
[42] † Oudenodon bainii Last Modified in 01/15/20 | 6 | 166 | 165 | 0 | 23 | 0 | 0 | 2 |
[43] † Tropidostoma dubium Last Modified in 01/15/20 | 17 | 155 | 155 | 0 | 22 | 0 | 0 | 2 |
[44] † Australobarbarus Last Modified in 01/15/20 | 41 | 131 | 131 | 0 | 22 | 0 | 0 | 2 |
[45] † Odontocyclops whaitsi Last Modified in 01/15/20 | 21 | 150 | 150 | 0 | 23 | 0 | 0 | 2 |
[46] † Idelesaurus tataricus Last Modified in 01/15/20 | 46 | 132 | 132 | 0 | 16 | 0 | 0 | 2 |
[47] † Aulacephalodon bainii Last Modified in 01/15/20 | 6 | 170 | 165 | 0 | 23 | 0 | 0 | 2 |
[48] † Pelanomodon moschops Last Modified in 01/15/20 | 69 | 111 | 110 | 0 | 15 | 0 | 0 | 2 |
[49] † Geikia locusticeps Last Modified in 01/15/20 | 40 | 138 | 138 | 0 | 16 | 0 | 0 | 2 |
[50] † Geikia elginensis Last Modified in 01/15/20 | 88 | 96 | 96 | 0 | 10 | 0 | 0 | 2 |
[51] † Elph borealis Last Modified in 01/15/20 | 69 | 113 | 113 | 0 | 12 | 0 | 0 | 2 |
[52] † Interpresosaurus blomi Last Modified in 01/15/20 | 150 | 44 | 44 | 0 | 0 | 0 | 0 | 2 |
[53] † Katumbia parringtoni Last Modified in 01/15/20 | 60 | 120 | 120 | 0 | 14 | 0 | 0 | 2 |
[54] † Delectosaurus arefjevi Last Modified in 01/15/20 | 74 | 108 | 107 | 0 | 13 | 0 | 0 | 2 |
[55] † Dicynodon lacerticeps Last Modified in 01/15/20 | 42 | 137 | 136 | 0 | 16 | 0 | 0 | 2 |
[56] † Dicynodon huenei Last Modified in 01/15/20 | 15 | 157 | 156 | 0 | 23 | 0 | 0 | 2 |
[57] † Daptocephalus leoniceps Last Modified in 01/15/20 | 32 | 145 | 145 | 0 | 17 | 0 | 0 | 2 |
[58] † Daqingshanodon limbus Last Modified in 01/15/20 | 58 | 121 | 121 | 0 | 15 | 0 | 0 | 2 |
[59] † Dinanomodon gilli Last Modified in 01/15/20 | 56 | 123 | 123 | 0 | 15 | 0 | 0 | 2 |
[60] † Peramodon amalitzkii Last Modified in 01/15/20 | 87 | 95 | 95 | 0 | 12 | 0 | 0 | 2 |
[61] † Vivaxosaurus trautscholdi Last Modified in 01/15/20 | 35 | 142 | 140 | 0 | 19 | 0 | 0 | 2 |
[62] † Jimusaria sinkiangensis Last Modified in 01/15/20 | 61 | 120 | 120 | 0 | 13 | 0 | 0 | 2 |
[63] † Sintocephalus alticeps Last Modified in 01/15/20 | 101 | 80 | 80 | 0 | 13 | 0 | 0 | 2 |
[64] † Basilodon woodwardi Last Modified in 01/15/20 | 70 | 111 | 111 | 0 | 13 | 0 | 0 | 2 |
[65] † Turfanodon bogdaensis Last Modified in 01/15/20 | 103 | 86 | 86 | 0 | 5 | 0 | 0 | 2 |
[66] † Keyseria benjamini Last Modified in 01/15/20 | 83 | 100 | 100 | 0 | 11 | 0 | 0 | 2 |
[67] † Gordonia traquairi Last Modified in 01/15/20 | 81 | 101 | 101 | 0 | 12 | 0 | 0 | 2 |
[68] † Syops vanhoepeni Last Modified in 01/15/20 | 105 | 87 | 87 | 0 | 2 | 0 | 0 | 2 |
[69] † Euptychognathus bathyrhynchus Last Modified in 01/15/20 | 66 | 113 | 113 | 0 | 15 | 0 | 0 | 2 |
[70] † Lystrosaurus hedini Last Modified in 01/15/20 | 46 | 132 | 132 | 0 | 16 | 0 | 0 | 2 |
[71] † Lystrosaurus maccaigi Last Modified in 01/15/20 | 37 | 138 | 138 | 0 | 19 | 0 | 0 | 2 |
[72] † Lystrosaurus curvatus Last Modified in 01/15/20 | 33 | 138 | 138 | 0 | 23 | 0 | 0 | 2 |
[73] † Lystrosaurus declivis Last Modified in 01/15/20 | 12 | 159 | 159 | 0 | 23 | 0 | 0 | 2 |
[74] † Lystrosaurus murrayi Last Modified in 01/15/20 | 12 | 159 | 159 | 0 | 23 | 0 | 0 | 2 |
[75] † Angonisaurus cruickshanki Last Modified in 01/15/20 | 49 | 126 | 126 | 0 | 19 | 0 | 0 | 2 |
[76] † Tetragonias njalilus Last Modified in 01/15/20 | 10 | 163 | 162 | 0 | 22 | 0 | 0 | 2 |
[77] † Shansiodon Last Modified in 01/15/20 | 34 | 138 | 138 | 0 | 22 | 0 | 0 | 2 |
[78] † Vinceria andina Last Modified in 01/15/20 | 86 | 96 | 96 | 0 | 12 | 0 | 0 | 2 |
[79] † Rhinodicynodon gracile Last Modified in 01/15/20 | 45 | 130 | 130 | 0 | 19 | 0 | 0 | 2 |
[80] † Dinodontosaurus Last Modified in 01/15/20 | 15 | 156 | 156 | 0 | 23 | 0 | 0 | 2 |
[81] † Kannemeyeria simocephalus Last Modified in 01/15/20 | 11 | 161 | 161 | 0 | 22 | 0 | 0 | 2 |
[82] † Kannemeyeria lophorhinus Last Modified in 01/15/20 | 53 | 129 | 129 | 0 | 12 | 0 | 0 | 2 |
[83] † Dolichuranus primaevus Last Modified in 01/15/20 | 35 | 141 | 141 | 0 | 18 | 0 | 0 | 2 |
[84] † Wadiasaurus indicus Last Modified in 01/15/20 | 68 | 108 | 107 | 0 | 19 | 0 | 0 | 2 |
[85] † Rabidosaurus cristatus Last Modified in 01/15/20 | 118 | 73 | 73 | 0 | 3 | 0 | 0 | 2 |
[86] † Rhadiodromus Last Modified in 01/15/20 | 87 | 95 | 94 | 0 | 13 | 0 | 0 | 2 |
[87] † Rechnisaurus cristarhynchus Last Modified in 01/15/20 | 112 | 75 | 75 | 0 | 7 | 0 | 0 | 2 |
[88] † Uralokannemeyeria vjuschkovi Last Modified in 01/15/20 | 104 | 79 | 79 | 0 | 11 | 0 | 0 | 2 |
[89] † Shaanbeikannemeyeria Last Modified in 01/15/20 | 58 | 121 | 121 | 0 | 15 | 0 | 0 | 2 |
[90] † Xiyukannemeyeria brevirostris Last Modified in 01/15/20 | 55 | 125 | 125 | 0 | 14 | 0 | 0 | 2 |
[91] † Parakannemeyeria Last Modified in 01/15/20 | 22 | 150 | 149 | 0 | 23 | 0 | 0 | 2 |
[92] † Sinokannemeyeria Last Modified in 01/15/20 | 27 | 145 | 145 | 0 | 22 | 0 | 0 | 2 |
[93] † Stahleckeria potens Last Modified in 01/15/20 | 12 | 161 | 161 | 0 | 21 | 0 | 0 | 2 |
[94] † Eubrachiosaurus browni Last Modified in 01/15/20 | 183 | 9 | 9 | 0 | 2 | 0 | 0 | 2 |
[95] † Sangusaurus parringtonii Last Modified in 01/15/20 | 46 | 134 | 134 | 0 | 14 | 0 | 0 | 2 |
[96] † Ischigualastia jenseni Last Modified in 01/15/20 | 24 | 150 | 150 | 0 | 20 | 0 | 0 | 2 |
[97] † Jachaleria Last Modified in 01/15/20 | 45 | 130 | 130 | 0 | 19 | 0 | 0 | 2 |
[98] † Zambiasaurus submersus Last Modified in 01/15/20 | 132 | 60 | 60 | 0 | 2 | 0 | 0 | 2 |
[99] † Placerias hesternus Last Modified in 01/15/20 | 72 | 120 | 120 | 0 | 2 | 0 | 0 | 2 |
[100] † Moghreberia nmachouensis Last Modified in 01/15/20 | 97 | 97 | 97 | 0 | 0 | 0 | 0 | 2 |
[101] † Rastodon procurvidens Last Modified in 01/15/20 | 80 | 101 | 101 | 0 | 13 | 0 | 0 | 2 |
[102] † Bulbasaurus phylloxyron Last Modified in 01/15/20 | 59 | 125 | 121 | 0 | 14 | 0 | 0 | 2 |
[103] † Compsodon helmoedi Last Modified in 01/15/20 | 70 | 111 | 111 | 0 | 13 | 0 | 0 | 2 |
[104] † Counillonia superoculis Last Modified in 01/15/20 | 107 | 78 | 78 | 0 | 9 | 0 | 0 | 2 |
[105] † Repelinosaurus robustus (1993-2) Last Modified in 01/15/20 | 113 | 75 | 75 | 0 | 6 | 0 | 0 | 2 |
[106] † Repelinosaurus robustus (1995-9) Last Modified in 01/15/20 | 106 | 82 | 82 | 0 | 6 | 0 | 0 | 2 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 17450 | |
Project overview | 1172 | |
Matrix views | 807 | Laos (118 views); Matrix landing page (689 views); |
Documents list | 687 | |
Media views | 2511 | Media search (2122 views); M683406 (389 views); |
Bibliography | 538 | |
Taxon list | 9629 | |
Views for media list | 768 | |
Specimen list | 1338 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 127 | |
Project downloads | 124 | |
Document downloads | 1 | Character list and original tnt matrix (without character names) (1 download); |
Matrix downloads | 2 | Laos (2 downloads); |