Project 3637: S. Hoffmann, R. M. Beck, J. R. Wible, G. W. Rougier, D. W. Krause. 2020. Phylogenetic placement of Adalatherium hui (Mammalia, Gondwanatheria) from the Late Cretaceous of Madagascar: implications for allotherian relationships. Journal of Vertebrate Paleontology. 40 (sup1):213-234.
Abstract
The phylogenetic position of Gondwanatheria within Mammaliaformes has historically been controversial. The well-preserved skeleton of Adalatherium hui from the Late Cretaceous of Madagascar offers a unique opportunity to address this issue, based on morphological data from the whole skeleton. Gondwanatheria were, until recently, known only from fragmentary dental and mandibular material, as well as a single cranium. The holotype of A. hui provides the first postcranial skeleton for gondwanatherians and substantially increases the amount of character data available to score. We sampled 530 characters and 84 cynodonts (including 34 taxa historically affiliated with Allotheria) to test the phylogenetic relationships of Gondwanatheria and Allotheria using parsimony, undated Bayesian, and tip-dated Bayesian methods. We tested three lower dental formulae for Adalatherium, because its postcanines are distinctly different from those of other mammaliaforms and cannot readily be homologized with any known dental pattern. In all analyses, Adalatherium is recovered within Gondwanatheria, most frequently outside of Sudamericidae or Ferugliotheriidae, which is congruent with establishment of the family Adalatheriidae. The different dental coding schemes do not greatly impact the position of Adalatherium, although there are differences in character optimization. In all analyses, Gondwanatheria are placed within Allotheria, either as sister to Multituberculata, nested within Multituberculata, or as sister to Cifelliodon (and Euharamiyida), or in a polytomy with other allotherians. The composition of Allotheria varies in our analyses. The haramiyidans Haramiyavia and Thomasia are placed outside of Allotheria in the parsimony and tip-dated Bayesian analyses, but in a polytomy with other allotherians in the undated Bayesian analyses.Read the article »
Article DOI: 10.1080/02724634.2020.1801706
Project DOI: 10.7934/P3637, http://dx.doi.org/10.7934/P3637
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 27858 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 530 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 1256 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:530/0 |
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MorphoBank Project 3637
MorphoBank Project 3637
- Creation Date:
12 February 2020 - Publication Date:
18 December 2020 - Project views: 13366
- Matrix downloads: 15
This research
supported by
Authors' Institutions
- University of Louisville
- University of Salford
- Carnegie Museum of Natural History
- Denver Museum of Nature and Science
- New York Institute of Technology
- Stony Brook University
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Simone Hoffmann Project Administrator | 86 | 1 | 1 | 530 | 0 | 0 | 27858 (20099, 0, 7759) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M26451' (86 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Thrinaxodon Last Modified in 12/18/20 | 38 | 322 | 315 | 0 | 177 | 0 | 0 | 1 |
[2] † Probainognathus Last Modified in 12/18/20 | 51 | 310 | 309 | 0 | 170 | 0 | 0 | 1 |
[3] † Cynognathus Last Modified in 12/18/20 | 103 | 253 | 253 | 0 | 174 | 0 | 0 | 1 |
[4] † Diademodon Last Modified in 12/18/20 | 73 | 285 | 283 | 0 | 174 | 0 | 0 | 1 |
[5] † Exaeretodon Last Modified in 12/18/20 | 89 | 288 | 281 | 0 | 160 | 0 | 0 | 1 |
[6] † Oligokyphus Last Modified in 12/18/20 | 69 | 319 | 318 | 0 | 143 | 0 | 0 | 1 |
[7] † Kayentatherium Last Modified in 12/18/20 | 111 | 279 | 278 | 0 | 141 | 0 | 0 | 1 |
[8] † Yunnanodon Last Modified in 12/18/20 | 258 | 148 | 148 | 0 | 124 | 0 | 0 | 1 |
[9] † Bocatherium Last Modified in 12/18/20 | 292 | 119 | 119 | 0 | 119 | 0 | 0 | 1 |
[10] † Bienotheroides Last Modified in 12/18/20 | 125 | 264 | 264 | 0 | 141 | 0 | 0 | 1 |
[11] † Bienotherium Last Modified in 12/18/20 | 185 | 211 | 211 | 0 | 134 | 0 | 0 | 1 |
[12] † Pachygenelus Last Modified in 12/18/20 | 91 | 297 | 297 | 0 | 142 | 0 | 0 | 1 |
[13] † Brasilodon Last Modified in 12/18/20 | 97 | 301 | 292 | 0 | 141 | 0 | 0 | 1 |
[14] † Riograndia Last Modified in 12/18/20 | 154 | 243 | 238 | 0 | 138 | 0 | 0 | 1 |
[15] † Sinoconodon Last Modified in 12/18/20 | 108 | 351 | 348 | 0 | 74 | 0 | 0 | 1 |
[16] † Morganucodon Last Modified in 12/18/20 | 49 | 416 | 412 | 0 | 69 | 0 | 0 | 1 |
[17] † Megazostrodon Last Modified in 12/18/20 | 150 | 317 | 317 | 0 | 63 | 0 | 0 | 1 |
[18] † Hadrocodium Last Modified in 12/18/20 | 233 | 237 | 237 | 0 | 60 | 0 | 0 | 1 |
[19] † Haldanodon Last Modified in 12/18/20 | 113 | 354 | 352 | 0 | 65 | 0 | 0 | 1 |
[20] † Pseudotribos Last Modified in 12/18/20 | 374 | 118 | 118 | 0 | 38 | 0 | 0 | 1 |
[21] Ornithorhynchus Last Modified in 02/12/20 | 44 | 393 | 390 | 0 | 96 | 0 | 0 | 1 |
[22] † Steropodon Last Modified in 12/18/20 | 458 | 48 | 48 | 0 | 24 | 0 | 0 | 1 |
[23] † Teinolophos Last Modified in 12/18/20 | 416 | 90 | 89 | 0 | 25 | 0 | 0 | 1 |
[24] † Obdurodon Last Modified in 12/18/20 | 202 | 273 | 272 | 0 | 56 | 0 | 0 | 1 |
[25] † Megaconus Last Modified in 12/18/20 | 293 | 143 | 143 | 0 | 94 | 0 | 0 | 1 |
[26] † Haramiyavia Last Modified in 12/18/20 | 335 | 104 | 103 | 0 | 92 | 0 | 0 | 1 |
[27] † Thomasia Last Modified in 12/18/20 | 392 | 52 | 52 | 0 | 86 | 0 | 0 | 1 |
[28] † Arboroharamiya j Last Modified in 12/18/20 | 284 | 138 | 138 | 0 | 108 | 0 | 0 | 1 |
[29] † Arboroharamiya a Last Modified in 12/18/20 | 212 | 207 | 207 | 0 | 111 | 0 | 0 | 1 |
[30] † Xianshou l Last Modified in 12/18/20 | 248 | 170 | 170 | 0 | 112 | 0 | 0 | 1 |
[31] † Xianshou s Last Modified in 12/18/20 | 278 | 147 | 147 | 0 | 105 | 0 | 0 | 1 |
[32] † Shenshou Last Modified in 12/18/20 | 172 | 240 | 240 | 0 | 118 | 0 | 0 | 1 |
[33] † Qishou Last Modified in 12/18/20 | 191 | 227 | 227 | 0 | 112 | 0 | 0 | 1 |
[34] † Maiopatagium Last Modified in 12/18/20 | 324 | 146 | 146 | 0 | 60 | 0 | 0 | 1 |
[35] † Vilevolodon Last Modified in 12/18/20 | 234 | 181 | 181 | 0 | 115 | 0 | 0 | 1 |
[36] † Cifelliodon Last Modified in 12/18/20 | 377 | 137 | 137 | 0 | 16 | 0 | 0 | 1 |
[37] † Adalatherium 1004 Last Modified in 12/18/20 | 139 | 258 | 256 | 0 | 135 | 0 | 0 | 1 |
[38] † Adalatherium 1013 Last Modified in 12/18/20 | 139 | 264 | 263 | 0 | 128 | 0 | 0 | 1 |
[39] † Adalatherium 1022 Last Modified in 12/18/20 | 139 | 267 | 266 | 0 | 125 | 0 | 0 | 1 |
[40] † Vintana Last Modified in 12/18/20 | 252 | 213 | 213 | 0 | 65 | 0 | 0 | 1 |
[41] † Ferugliotherium Last Modified in 12/18/20 | 389 | 48 | 48 | 0 | 93 | 0 | 0 | 1 |
[42] † Trapalcotherium Last Modified in 12/18/20 | 430 | 19 | 19 | 0 | 81 | 0 | 0 | 1 |
[43] † Bharattherium Last Modified in 12/18/20 | 429 | 16 | 16 | 0 | 85 | 0 | 0 | 1 |
[44] † Gondwanatherium Last Modified in 12/18/20 | 391 | 32 | 32 | 0 | 107 | 0 | 0 | 1 |
[45] † Lavanify Last Modified in 12/18/20 | 429 | 16 | 16 | 0 | 85 | 0 | 0 | 1 |
[46] † Sudamerica Last Modified in 12/18/20 | 369 | 48 | 47 | 0 | 114 | 0 | 0 | 1 |
[47] † Galulatherium Last Modified in 12/18/20 | 368 | 42 | 42 | 0 | 120 | 0 | 0 | 1 |
[48] † Greniodon Last Modified in 12/18/20 | 427 | 19 | 19 | 0 | 84 | 0 | 0 | 1 |
[49] † Jeholbaatar Last Modified in 12/18/20 | 188 | 244 | 244 | 0 | 98 | 0 | 0 | 1 |
[50] † Rugosodon Last Modified in 12/18/20 | 248 | 190 | 190 | 0 | 92 | 0 | 0 | 1 |
[51] † Guimarota paulchoffatiidae Last Modified in 12/18/20 | 215 | 233 | 217 | 0 | 98 | 0 | 0 | 1 |
[52] † Sinobaatar Last Modified in 12/18/20 | 211 | 222 | 220 | 0 | 99 | 0 | 0 | 1 |
[53] † Plagiaulacidae Last Modified in 12/18/20 | 322 | 116 | 113 | 0 | 95 | 0 | 0 | 1 |
[54] † Ptilodus Last Modified in 12/18/20 | 105 | 320 | 319 | 0 | 106 | 0 | 0 | 1 |
[55] † Taeniolabis Last Modified in 12/18/20 | 241 | 176 | 176 | 0 | 113 | 0 | 0 | 1 |
[56] † Lambdopsalis Last Modified in 12/18/20 | 136 | 278 | 276 | 0 | 118 | 0 | 0 | 1 |
[57] † Catopsbaatar Last Modified in 12/18/20 | 148 | 280 | 277 | 0 | 105 | 0 | 0 | 1 |
[58] † Chulsanbaatar Last Modified in 12/18/20 | 93 | 332 | 330 | 0 | 107 | 0 | 0 | 1 |
[59] † Nemegtbaatar Last Modified in 12/18/20 | 105 | 319 | 318 | 0 | 107 | 0 | 0 | 1 |
[60] † Kryptobaatar Last Modified in 12/18/20 | 41 | 388 | 382 | 0 | 107 | 0 | 0 | 1 |
[61] † Gobiconodon Last Modified in 12/18/20 | 163 | 305 | 297 | 0 | 70 | 0 | 0 | 1 |
[62] † Repenomamus Last Modified in 12/18/20 | 116 | 347 | 345 | 0 | 69 | 0 | 0 | 1 |
[63] † Yanoconodon Last Modified in 12/18/20 | 244 | 220 | 220 | 0 | 66 | 0 | 0 | 1 |
[64] † Jeholodens Last Modified in 12/18/20 | 256 | 211 | 209 | 0 | 65 | 0 | 0 | 1 |
[65] † Akidolestes Last Modified in 12/18/20 | 306 | 171 | 171 | 0 | 53 | 0 | 0 | 1 |
[66] † Zhangheotherium Last Modified in 12/18/20 | 215 | 269 | 268 | 0 | 47 | 0 | 0 | 1 |
[67] † Maotherium Last Modified in 12/18/20 | 173 | 301 | 297 | 0 | 60 | 0 | 0 | 1 |
[68] † Cronopio Last Modified in 12/18/20 | 239 | 239 | 238 | 0 | 53 | 0 | 0 | 1 |
[69] † Henkelotherium Last Modified in 12/18/20 | 223 | 261 | 261 | 0 | 46 | 0 | 0 | 1 |
[70] † Dryolestes Last Modified in 12/18/20 | 266 | 220 | 219 | 0 | 45 | 0 | 0 | 1 |
[71] † Vincelestes Last Modified in 12/18/20 | 38 | 441 | 440 | 0 | 52 | 0 | 0 | 1 |
[72] † Eomaia Last Modified in 12/18/20 | 223 | 256 | 255 | 0 | 52 | 0 | 0 | 1 |
[73] † Juramaia Last Modified in 12/18/20 | 338 | 161 | 161 | 0 | 31 | 0 | 0 | 1 |
[74] † Asioryctes Last Modified in 12/18/20 | 138 | 343 | 341 | 0 | 51 | 0 | 0 | 1 |
[75] † Ukhaatherium Last Modified in 12/18/20 | 120 | 358 | 357 | 0 | 53 | 0 | 0 | 1 |
[76] † Zalambdalestes Last Modified in 12/18/20 | 85 | 397 | 391 | 0 | 54 | 0 | 0 | 1 |
[77] Erinaceus Last Modified in 02/12/20 | 4 | 463 | 461 | 0 | 65 | 0 | 0 | 1 |
[78] † Gomphos Last Modified in 12/18/20 | 132 | 323 | 323 | 0 | 75 | 0 | 0 | 1 |
[79] † Tribosphenomys Last Modified in 12/18/20 | 299 | 175 | 171 | 0 | 60 | 0 | 0 | 1 |
[80] Dasypus Last Modified in 02/12/20 | 91 | 350 | 345 | 0 | 94 | 0 | 0 | 1 |
[81] † Asiatherium Last Modified in 12/18/20 | 212 | 258 | 258 | 0 | 60 | 0 | 0 | 1 |
[82] † Deltatheridium Last Modified in 12/18/20 | 184 | 282 | 280 | 0 | 66 | 0 | 0 | 1 |
[83] † Pucadelphys Last Modified in 12/18/20 | 56 | 402 | 400 | 0 | 74 | 0 | 0 | 1 |
[84] Didelphis Last Modified in 02/12/20 | 7 | 448 | 446 | 0 | 77 | 0 | 0 | 1 |
[85] † Sinodelphys Last Modified in 12/18/20 | 287 | 203 | 203 | 0 | 40 | 0 | 0 | 1 |
[86] † Ambolestes Last Modified in 12/18/20 | 266 | 227 | 227 | 0 | 37 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 13366 | |
Project overview | 825 | |
Matrix views | 690 | Adalatherium_SVP Memoir (137 views); Matrix landing page (553 views); |
Media views | 2118 | Media search (1791 views); M738318 (327 views); |
Specimen list | 1038 | |
Bibliography | 385 | |
Views for media list | 638 | |
Taxon list | 7668 | |
Documents list | 4 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 121 | |
Matrix downloads | 15 | Adalatherium_SVP Memoir (15 downloads); |
Project downloads | 106 |