Project 3966: A. E. Fernandes, O. Mateus, B. Andres, M. J. Polcyn, A. S. Schulp, A. O. Gonçalves, L. L. Jacobs. 2022. Pterosaurs from the Late Cretaceous of Angola. Diversity. 14 (9):741.
Abstract
Here, we describe the first pterosaur remains from Angola, an assemblage of fourteen bones from the Lower Maastrichtian marine deposits of Bentiaba, Namibe Province. One new species is introduced, Epapatelo otyikokolo, gen. et sp. nov., which comprises an articulated partial left humerus and ulna as well as an articulated left ulna and radius (from a second individual). Phylogenetic analysis confirms a non-nyctosaurid pteranodontian attribution for this new taxon and supports a new apomorphy-based clade, Aponyctosauria, which is here defined. Late Cretaceous pteranodontians are rare in Sub-Saharan Africa and throughout the Southern Hemisphere. Preliminary histological analysis also reveals a likely sub-adult age for one of the specimens. This fossil assemblage provides a first glimpse of Angolan pterosaur paleobiodiversity providing further insight into the Gondwanan ecosystems of the Upper Cretaceous.Read the article »
Article DOI: 10.3390/d14090741
Project DOI: 10.7934/P3966, http://dx.doi.org/10.7934/P3966
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 14584 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 271 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 548 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:245/26 |
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MorphoBank Project 3966
MorphoBank Project 3966
- Creation Date:
25 March 2021 - Publication Date:
03 March 2023 - Project views: 7668
- Media downloads: 2
- Matrix downloads: 5
Authors' Institutions
- University of Sheffield
- Universiteit Utrecht
- Museu da Lourinha
- Bayerische Staatssammlung fuer Palaeontologie und Geologie
- Naturalis Biodiversity Center
- Universidade Nova de Lisboa
- Southern Methodist University
- Universidade Agostinho Neto
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Alexandra Fernandes Project Administrator | 139 | 16 | 16 | 271 | 0 | 0 | 14584 (12177, 0, 2407) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29172' (132 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Euparkeria capensis Last Modified in 03/06/23 | 47 | 189 | 189 | 0 | 35 | 0 | 0 | 1 |
[2] Ornithosuchus longidens Last Modified in 03/06/23 | 73 | 165 | 164 | 0 | 34 | 0 | 0 | 1 |
[3] Herrerasaurus ischigualastensis Last Modified in 03/06/23 | 64 | 173 | 173 | 0 | 34 | 0 | 0 | 1 |
[4] Scleromochlus taylori Last Modified in 03/06/23 | 114 | 122 | 122 | 0 | 35 | 0 | 0 | 1 |
[5] Eudimorphodon ranzii Last Modified in 03/06/23 | 78 | 166 | 166 | 0 | 27 | 0 | 0 | 1 |
[6] Eudimorphodon rosenfeldi Last Modified in 03/06/23 | 62 | 167 | 167 | 0 | 42 | 0 | 0 | 1 |
[7] Eudimorphodon cromptonellus Last Modified in 03/06/23 | 166 | 91 | 90 | 0 | 15 | 0 | 0 | 1 |
[8] Peteinosaurus zambellii Last Modified in 03/06/23 | 212 | 52 | 52 | 0 | 7 | 0 | 0 | 1 |
[9] Caviramus schesaplanensis Last Modified in 03/06/23 | 229 | 40 | 39 | 0 | 3 | 0 | 0 | 1 |
[10] Raeticodactylus filisurensis Last Modified in 03/06/23 | 102 | 141 | 141 | 0 | 28 | 0 | 0 | 1 |
[11] Austriadactylus cristatus Last Modified in 03/06/23 | 101 | 141 | 140 | 0 | 30 | 0 | 0 | 1 |
[12] Preondactylus buffarinii Last Modified in 03/06/23 | 100 | 131 | 131 | 0 | 40 | 0 | 0 | 1 |
[13] Dimorphodon macronyx Last Modified in 03/06/23 | 55 | 173 | 173 | 0 | 43 | 0 | 0 | 1 |
[14] Parapsicephalus purdoni Last Modified in 03/06/23 | 192 | 74 | 73 | 0 | 6 | 0 | 0 | 1 |
[15] Campylognathoides liasicus Last Modified in 03/06/23 | 37 | 188 | 188 | 0 | 46 | 0 | 0 | 1 |
[16] Campylognathoides zitteli Last Modified in 03/06/23 | 71 | 160 | 160 | 0 | 40 | 0 | 0 | 1 |
[17] Sericipterus wucaiwanensis Last Modified in 03/06/23 | 134 | 123 | 123 | 0 | 14 | 0 | 0 | 1 |
[18] Angustinaripterus longicephalus Last Modified in 03/06/23 | 155 | 104 | 104 | 0 | 12 | 0 | 0 | 1 |
[19] Harpactognathus gentryii Last Modified in 03/06/23 | 218 | 52 | 51 | 0 | 2 | 0 | 0 | 1 |
[20] Cacibupteryx caribensis Last Modified in 03/06/23 | 188 | 77 | 76 | 0 | 7 | 0 | 0 | 1 |
[21] Rhamphorhynchus muensteri Last Modified in 03/06/23 | 30 | 198 | 195 | 0 | 46 | 0 | 0 | 1 |
[22] Qinglongopterus guoi Last Modified in 03/06/23 | 113 | 120 | 120 | 0 | 38 | 0 | 0 | 1 |
[23] Nesodactylus hesperius Last Modified in 03/06/23 | 202 | 53 | 53 | 0 | 16 | 0 | 0 | 1 |
[24] Dorygnathus banthensis Last Modified in 03/06/23 | 29 | 197 | 196 | 0 | 46 | 0 | 0 | 1 |
[25] Scaphognathus crassirostris Last Modified in 03/06/23 | 48 | 179 | 178 | 0 | 45 | 0 | 0 | 1 |
[26] Sordes pilosus Last Modified in 03/06/23 | 36 | 191 | 190 | 0 | 45 | 0 | 0 | 1 |
[27] Darwinopterus modularis Last Modified in 03/06/23 | 64 | 166 | 166 | 0 | 41 | 0 | 0 | 1 |
[28] Wukongopterus lii Last Modified in 03/06/23 | 131 | 111 | 111 | 0 | 29 | 0 | 0 | 1 |
[29] Pterorhynchus wellnhoferi Last Modified in 03/06/23 | 96 | 141 | 141 | 0 | 34 | 0 | 0 | 1 |
[30] Changchengopterus pani Last Modified in 03/06/23 | 203 | 48 | 48 | 0 | 20 | 0 | 0 | 1 |
[31] Batrachognathus volans Last Modified in 03/06/23 | 114 | 132 | 132 | 0 | 25 | 0 | 0 | 1 |
[32] Jeholopterus ningchengensis Last Modified in 03/06/23 | 93 | 138 | 138 | 0 | 40 | 0 | 0 | 1 |
[33] Anurognathus ammoni Last Modified in 03/06/23 | 87 | 144 | 144 | 0 | 40 | 0 | 0 | 1 |
[34] Dendrorhynchoides curvidentatus Last Modified in 03/06/23 | 141 | 104 | 104 | 0 | 26 | 0 | 0 | 1 |
[35] Kryptodrakon progenitor Last Modified in 03/06/23 | 253 | 16 | 16 | 0 | 2 | 0 | 0 | 1 |
[36] Gnathosaurus subulatus Last Modified in 03/06/23 | 159 | 102 | 102 | 0 | 10 | 0 | 0 | 1 |
[37] Gnathosaurus macrurus Last Modified in 03/06/23 | 226 | 40 | 40 | 0 | 5 | 0 | 0 | 1 |
[38] Plataleorhynchus streptophorodon Last Modified in 03/06/23 | 239 | 31 | 31 | 0 | 1 | 0 | 0 | 1 |
[39] Huanhepterus quingyangensis Last Modified in 03/06/23 | 167 | 85 | 85 | 0 | 19 | 0 | 0 | 1 |
[40] Moganopterus zhuiana Last Modified in 03/06/23 | 163 | 95 | 95 | 0 | 13 | 0 | 0 | 1 |
[41] Elanodactylus prolatus Last Modified in 03/06/23 | 186 | 65 | 65 | 0 | 20 | 0 | 0 | 1 |
[42] Kepodactylus insperatus Last Modified in 03/06/23 | 241 | 28 | 28 | 0 | 2 | 0 | 0 | 1 |
[43] Ctenochasma elegans Last Modified in 03/06/23 | 56 | 174 | 173 | 0 | 42 | 0 | 0 | 1 |
[44] Pterodaustro guinazui Last Modified in 03/06/23 | 66 | 166 | 166 | 0 | 39 | 0 | 0 | 1 |
[45] Beipiaopterus chenianus Last Modified in 03/06/23 | 208 | 37 | 37 | 0 | 26 | 0 | 0 | 1 |
[46] Gegepterus changae Last Modified in 03/06/23 | 147 | 107 | 106 | 0 | 18 | 0 | 0 | 1 |
[47] Feilongus youngi Last Modified in 03/06/23 | 149 | 109 | 109 | 0 | 13 | 0 | 0 | 1 |
[48] Cycnorhamphus suevicus Last Modified in 03/06/23 | 61 | 175 | 174 | 0 | 36 | 0 | 0 | 1 |
[49] Ardeadactylus longicollum Last Modified in 03/06/23 | 58 | 174 | 174 | 0 | 39 | 0 | 0 | 1 |
[50] Pterodactylus antiquus Last Modified in 03/06/23 | 50 | 179 | 179 | 0 | 42 | 0 | 0 | 1 |
[51] Normannognathus wellnhoferi Last Modified in 03/06/23 | 217 | 53 | 53 | 0 | 1 | 0 | 0 | 1 |
[52] Germanodactylus cristatus Last Modified in 03/06/23 | 65 | 165 | 165 | 0 | 41 | 0 | 0 | 1 |
[53] Germanodactylus rhamphastinus Last Modified in 03/06/23 | 83 | 153 | 153 | 0 | 35 | 0 | 0 | 1 |
[54] Haopterus gracilis Last Modified in 03/06/23 | 133 | 112 | 112 | 0 | 26 | 0 | 0 | 1 |
[55] Aetodactylus halli Last Modified in 03/06/23 | 216 | 51 | 51 | 0 | 4 | 0 | 0 | 1 |
[56] Anhanguera santanae Last Modified in 03/06/23 | 67 | 179 | 179 | 0 | 25 | 0 | 0 | 1 |
[57] Anhanguera piscator Last Modified in 03/06/23 | 52 | 186 | 186 | 0 | 33 | 0 | 0 | 1 |
[58] Anhanguera blittersdorffi Last Modified in 03/06/23 | 144 | 113 | 113 | 0 | 14 | 0 | 0 | 1 |
[59] Anhanguera araripensis Last Modified in 03/06/23 | 136 | 123 | 123 | 0 | 12 | 0 | 0 | 1 |
[60] Liaoningopterus gui Last Modified in 03/06/23 | 173 | 94 | 93 | 0 | 5 | 0 | 0 | 1 |
[61] Siroccopteryx moroccensis Last Modified in 03/06/23 | 236 | 35 | 35 | 0 | 0 | 0 | 0 | 1 |
[62] Tropeognathus mesembrinus Last Modified in 03/06/23 | 139 | 117 | 117 | 0 | 15 | 0 | 0 | 1 |
[63] Coloborhynchus clavirostris Last Modified in 03/06/23 | 241 | 30 | 30 | 0 | 0 | 0 | 0 | 1 |
[64] Coloborhynchus wadleighi Last Modified in 03/06/23 | 236 | 35 | 35 | 0 | 0 | 0 | 0 | 1 |
[65] Ornithocheirus simus Last Modified in 03/06/23 | 244 | 27 | 27 | 0 | 0 | 0 | 0 | 1 |
[66] Brasileodactylus araripensis Last Modified in 03/06/23 | 233 | 38 | 38 | 0 | 0 | 0 | 0 | 1 |
[67] Ludodactylus sibbicki Last Modified in 03/06/23 | 147 | 111 | 111 | 0 | 13 | 0 | 0 | 1 |
[68] Boreopterus cuiae Last Modified in 03/06/23 | 144 | 104 | 103 | 0 | 24 | 0 | 0 | 1 |
[69] Guidraco venator Last Modified in 03/06/23 | 161 | 97 | 97 | 0 | 13 | 0 | 0 | 1 |
[70] Zhenyuanopterus longiristris Last Modified in 03/06/23 | 99 | 140 | 139 | 0 | 33 | 0 | 0 | 1 |
[71] Cearadactylus atrox Last Modified in 03/06/23 | 157 | 103 | 103 | 0 | 11 | 0 | 0 | 1 |
[72] Hongshanopterus lacustris Last Modified in 03/06/23 | 191 | 73 | 72 | 0 | 8 | 0 | 0 | 1 |
[73] Lonchodectes compressirostris Last Modified in 03/06/23 | 235 | 35 | 35 | 0 | 1 | 0 | 0 | 1 |
[74] Lonchodraco giganteus Last Modified in 03/06/23 | 199 | 70 | 70 | 0 | 2 | 0 | 0 | 1 |
[75] Cimoliopterus cuvieri Last Modified in 03/06/23 | 229 | 41 | 41 | 0 | 1 | 0 | 0 | 1 |
[76] Cimoliopterus dunni Last Modified in 03/06/23 | 231 | 39 | 39 | 0 | 1 | 0 | 0 | 1 |
[77] Nurhachius ignaciobritoi Last Modified in 03/06/23 | 86 | 152 | 151 | 0 | 34 | 0 | 0 | 1 |
[78] Liaoxipterus brachyognathus Last Modified in 03/06/23 | 235 | 32 | 32 | 0 | 4 | 0 | 0 | 1 |
[79] Istiodactylus sinensis Last Modified in 03/06/23 | 120 | 123 | 123 | 0 | 28 | 0 | 0 | 1 |
[80] Istiodactylus latidens Last Modified in 03/06/23 | 109 | 140 | 140 | 0 | 22 | 0 | 0 | 1 |
[81] Pteranodon longiceps Last Modified in 03/06/23 | 66 | 171 | 171 | 0 | 34 | 0 | 0 | 1 |
[82] Pteranodon sternbergi Last Modified in 03/06/23 | 136 | 100 | 100 | 0 | 35 | 0 | 0 | 1 |
[83] Tethydraco regalis Last Modified in 03/06/23 | 258 | 12 | 12 | 0 | 1 | 0 | 0 | 1 |
[84] Alamodactylus byrdi Last Modified in 03/06/23 | 255 | 15 | 15 | 0 | 1 | 0 | 0 | 1 |
[85] Nyctosaurus grandis Last Modified in 03/06/23 | 238 | 26 | 26 | 0 | 7 | 0 | 0 | 1 |
[86] Alcione elainus Last Modified in 03/06/23 | 227 | 37 | 37 | 0 | 7 | 0 | 0 | 1 |
[87] Simurghia robusta Last Modified in 03/06/23 | 265 | 5 | 5 | 0 | 1 | 0 | 0 | 1 |
[88] Cretornis hlavaci Last Modified in 03/06/23 | 253 | 18 | 18 | 0 | 0 | 0 | 0 | 1 |
[89] Nyctosaurus gracilis Last Modified in 03/06/23 | 70 | 163 | 163 | 0 | 38 | 0 | 0 | 1 |
[90] Nyctosaurus nanus Last Modified in 03/06/23 | 245 | 24 | 24 | 0 | 2 | 0 | 0 | 1 |
[91] Nyctosaurus lamegoi Last Modified in 03/06/23 | 265 | 5 | 5 | 0 | 1 | 0 | 0 | 1 |
[92] Muzquizopteryx coahuilensis Last Modified in 03/06/23 | 167 | 91 | 91 | 0 | 13 | 0 | 0 | 1 |
[93] Tupandactylus navigans Last Modified in 03/06/23 | 202 | 61 | 61 | 0 | 8 | 0 | 0 | 1 |
[94] Tupandactylus imperator Last Modified in 03/06/23 | 189 | 74 | 74 | 0 | 8 | 0 | 0 | 1 |
[95] Bakonydraco galaczi Last Modified in 03/06/23 | 237 | 31 | 31 | 0 | 3 | 0 | 0 | 1 |
[96] Tapejara wellnhoferi Last Modified in 03/06/23 | 63 | 172 | 172 | 0 | 36 | 0 | 0 | 1 |
[97] Europejara olcadesorum Last Modified in 03/06/23 | 223 | 45 | 45 | 0 | 3 | 0 | 0 | 1 |
[98] Vectidraco daisymorrisae Last Modified in 03/06/23 | 257 | 11 | 11 | 0 | 3 | 0 | 0 | 1 |
[99] Caiuajara dobruskii Last Modified in 03/06/23 | 114 | 136 | 135 | 0 | 22 | 0 | 0 | 1 |
[100] Huaxiapterus benxiensis Last Modified in 03/06/23 | 153 | 96 | 96 | 0 | 22 | 0 | 0 | 1 |
[101] Huaxiapterus corallatus Last Modified in 03/06/23 | 144 | 94 | 94 | 0 | 33 | 0 | 0 | 1 |
[102] Eopteranodon lii Last Modified in 03/06/23 | 157 | 90 | 90 | 0 | 24 | 0 | 0 | 1 |
[103] Huaxiapterus jii Last Modified in 03/06/23 | 142 | 100 | 100 | 0 | 29 | 0 | 0 | 1 |
[104] Sinopterus dongi Last Modified in 03/06/23 | 100 | 134 | 134 | 0 | 37 | 0 | 0 | 1 |
[105] Bennettazhia oregonensis Last Modified in 03/06/23 | 253 | 16 | 16 | 0 | 2 | 0 | 0 | 1 |
[106] Dsungaripterus weii Last Modified in 03/06/23 | 54 | 181 | 181 | 0 | 36 | 0 | 0 | 1 |
[107] Domeykodactylus ceciliae Last Modified in 03/06/23 | 240 | 30 | 30 | 0 | 1 | 0 | 0 | 1 |
[108] Noripterus parvus Last Modified in 03/06/23 | 72 | 168 | 168 | 0 | 31 | 0 | 0 | 1 |
[109] Noripterus complicidens Last Modified in 03/06/23 | 153 | 92 | 92 | 0 | 26 | 0 | 0 | 1 |
[110] Tupuxuara longicristatus Last Modified in 03/06/23 | 234 | 36 | 36 | 0 | 1 | 0 | 0 | 1 |
[111] Tupuxuara leonardii Last Modified in 03/06/23 | 89 | 149 | 149 | 0 | 33 | 0 | 0 | 1 |
[112] Alanqa saharica Last Modified in 03/06/23 | 244 | 26 | 26 | 0 | 1 | 0 | 0 | 1 |
[113] Aerotitan sudamericanus Last Modified in 03/06/23 | 255 | 14 | 14 | 0 | 2 | 0 | 0 | 1 |
[114] Thalassodromeus sethi Last Modified in 03/06/23 | 167 | 96 | 96 | 0 | 8 | 0 | 0 | 1 |
[115] Chaoyangopterus zhangi Last Modified in 03/06/23 | 163 | 84 | 84 | 0 | 24 | 0 | 0 | 1 |
[116] Jidapterus edentus Last Modified in 03/06/23 | 151 | 88 | 88 | 0 | 32 | 0 | 0 | 1 |
[117] Eoazhdarcho liaoxiensis Last Modified in 03/06/23 | 199 | 53 | 53 | 0 | 19 | 0 | 0 | 1 |
[118] Shenzhoupterus chaoyangensis Last Modified in 03/06/23 | 169 | 84 | 84 | 0 | 18 | 0 | 0 | 1 |
[119] Radiodactylus langstoni Last Modified in 03/06/23 | 258 | 13 | 13 | 0 | 0 | 0 | 0 | 1 |
[120] Microtuban altivolans Last Modified in 03/06/23 | 232 | 28 | 28 | 0 | 11 | 0 | 0 | 1 |
[121] Aralazhdarcho bestobensis Last Modified in 03/06/23 | 254 | 17 | 17 | 0 | 0 | 0 | 0 | 1 |
[122] Volgadraco bogolubovi Last Modified in 03/06/23 | 242 | 26 | 26 | 0 | 3 | 0 | 0 | 1 |
[123] Eurazhdarcho langendorfensis Last Modified in 03/06/23 | 259 | 10 | 10 | 0 | 2 | 0 | 0 | 1 |
[124] Montanazhdarcho minor Last Modified in 03/06/23 | 211 | 53 | 53 | 0 | 7 | 0 | 0 | 1 |
[125] Phosphatodraco mauritanicus Last Modified in 03/06/23 | 261 | 9 | 9 | 0 | 1 | 0 | 0 | 1 |
[126] Zhejiangopterus linhaiensis Last Modified in 03/06/23 | 126 | 114 | 114 | 0 | 31 | 0 | 0 | 1 |
[127] Azhdarcho lancicollis Last Modified in 03/06/23 | 184 | 84 | 84 | 0 | 3 | 0 | 0 | 1 |
[128] OCP dek GE 716 Last Modified in 03/06/23 | 263 | 7 | 7 | 0 | 1 | 0 | 0 | 1 |
[129] Hatzegopteryx thambema Last Modified in 03/06/23 | 251 | 20 | 20 | 0 | 0 | 0 | 0 | 1 |
[130] Arambourgiania philadelphiae Last Modified in 03/06/23 | 262 | 8 | 8 | 0 | 1 | 0 | 0 | 1 |
[131] Quetzalcoatlus spp Last Modified in 03/06/23 | 106 | 146 | 145 | 0 | 20 | 0 | 0 | 1 |
[132] Epapatelo otyikokolo Last Modified in 03/06/23 | 261 | 9 | 9 | 0 | 1 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 7668 | |
Project overview | 333 | |
Matrix views | 216 | Matrix landing page (200 views); (3 views); Pterosaurs from the Late Cretaceous of Angola (13 views); |
Media views | 2442 | Media search (651 views); M838532 (122 views); M838540 (116 views); M838538 (117 views); M740037 (107 views); M838537 (115 views); M838539 (113 views); M838541 (108 views); M838543 (113 views); M838544 (113 views); M838545 (112 views); M838546 (114 views); M838547 (109 views); M838548 (108 views); M838551 (107 views); M838552 (110 views); M838555 (107 views); |
Specimen list | 1676 | |
Taxon list | 2450 | |
Bibliography | 152 | |
Views for media list | 393 | |
Documents list | 6 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 8 | |
Media downloads | 2 | M838532 (2 downloads); |
Matrix downloads | 5 | Pterosaurs from the Late Cretaceous of Angola (5 downloads); |
Project downloads | 1 |