Project 4186: S. Y. Smith, R. A. Stockey. 2002. Permineralized Pine Cones from the Cretaceous of Vancouver Island, British Columbia. International Journal of Plant Sciences. 163 (1):185-196.
Abstract
Two abraded, cylindrical cone specimens found in calcareous concretions from the Cretaceous Spray Formation (Late Campanian) of Vancouver Island, British Columbia, were sectioned using the cellulose acetate peel technique and characterized anatomically. Their sclerenchymatous pith is surrounded by a ring of separate secondary xylem bundles that lack resin canals. The outer cortical zone is sclerenchymatous and covered in a dense ramentum of trichomes that is also present on ovuliferous scale and bract bases. Vascular traces to the ovuliferous scales and bracts arise independently. The bract, with a terete trace and two lateral resin canals, lacks a distinct abaxial lobe. Ovuliferous scales are sclerotic with resin canals adaxial, abaxial, and between the vascular bundles. The ovuliferous scales have an interseminal ridge that is prominent and thick near the micropylar end of the seed, thins out near the seed chalaza, and attaches to the seed wing tissue. There are two winged seeds per scale, and the edges of the ovuliferous scale turn upward and partly enclose the seeds near the micropylar end. Seeds have a ridged sclerotesta; nucellus, megagametophyte, and embryos with eight cotyledons are preserved. Cone structure most closely resembles fossil Pinaceae of the genus Pityostrobus. These cones have a unique combination of characters that distinguish them from the previously described taxa and are described as Pityostrobus beardii sp. nov. A phylogenetic analysis using morphological data from the ovulate cones of extant and fossil taxa of Pinaceae with Cryptomeria japonica (L. f.) D. Don and Sciadopitys verticillata (Thunb.) Siebold et Zucc. as outgroups was undertaken to assess the phylogenetic position of P. beardii within Pinaceae. Pityostrobus beardii appears to be most closely related to Pityostrobus hokodzensis from the Cretaceous of Russia. These cones provide further evidence that the Pinaceae, like the angiosperms, were undergoing a rapid Cretaceous radiation.Read the article »
Article DOI: 10.1086/324553
Project DOI: 10.7934/P4186, http://dx.doi.org/10.7934/P4186
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 1572 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 33 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 96 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:33/0 |
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MorphoBank Project 4186
MorphoBank Project 4186
- Creation Date:
30 January 2022 - Publication Date:
03 August 2022 - Project views: 4471
- Matrix downloads: 2
Authors' Institutions
- University of Alberta
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 48 | 1 | 1 | 33 | 0 | 0 | 1572 (1572, 0, 0) | 0 | 0 | 0 | ||||
Selena Smith Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M28695' (48 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Abies Last Modified in 01/30/22 | 0 | 39 | 33 | 0 | 0 | 0 | 0 | 2 |
[2] Cathaya Last Modified in 01/30/22 | 3 | 30 | 30 | 0 | 0 | 0 | 0 | 2 |
[3] Cedrus Last Modified in 01/30/22 | 0 | 36 | 33 | 0 | 0 | 0 | 0 | 2 |
[4] Larix Last Modified in 01/30/22 | 0 | 41 | 33 | 0 | 0 | 0 | 0 | 2 |
[5] Keteleeria Last Modified in 01/30/22 | 0 | 35 | 33 | 0 | 0 | 0 | 0 | 2 |
[6] Picea Last Modified in 01/30/22 | 0 | 42 | 33 | 0 | 0 | 0 | 0 | 2 |
[7] Pinus Last Modified in 01/30/22 | 0 | 40 | 33 | 0 | 0 | 0 | 0 | 2 |
[8] Pseudolarix Last Modified in 01/30/22 | 2 | 31 | 31 | 0 | 0 | 0 | 0 | 2 |
[9] Pseudotsuga Last Modified in 01/30/22 | 0 | 36 | 33 | 0 | 0 | 0 | 0 | 2 |
[10] Tsuga Last Modified in 01/30/22 | 0 | 38 | 33 | 0 | 0 | 0 | 0 | 2 |
[11] † Pseudoaraucaria arnoldii Last Modified in 01/30/22 | 0 | 34 | 33 | 0 | 0 | 0 | 0 | 2 |
[12] † Pseudoaraucaria benstedii Last Modified in 01/30/22 | 7 | 26 | 26 | 0 | 0 | 0 | 0 | 2 |
[13] † Pseudoaraucaria gibbosa Last Modified in 01/30/22 | 1 | 32 | 32 | 0 | 0 | 0 | 0 | 2 |
[14] † Pseudoaraucaria heeri Last Modified in 01/30/22 | 0 | 33 | 33 | 0 | 0 | 0 | 0 | 2 |
[15] † Pseudoaraucaria loppinetti Last Modified in 01/30/22 | 1 | 32 | 32 | 0 | 0 | 0 | 0 | 2 |
[16] † Pseudoaraucaria major Last Modified in 01/30/22 | 0 | 33 | 33 | 0 | 0 | 0 | 0 | 2 |
[17] † Obirastrobus kokubunii Last Modified in 01/30/22 | 0 | 34 | 33 | 0 | 0 | 0 | 0 | 2 |
[18] † Obirastrobus nihongii Last Modified in 01/30/22 | 0 | 34 | 33 | 0 | 0 | 0 | 0 | 2 |
[19] † Pityostrobus andraei Last Modified in 01/30/22 | 6 | 27 | 27 | 0 | 0 | 0 | 0 | 2 |
[20] † Pityostrobus argonnensis Last Modified in 01/30/22 | 2 | 31 | 31 | 0 | 0 | 0 | 0 | 2 |
[21] † Pityostrobus beardii Last Modified in 01/30/22 | 0 | 33 | 33 | 0 | 0 | 0 | 0 | 2 |
[22] † Pityostrobus bommeri Last Modified in 01/30/22 | 1 | 32 | 32 | 0 | 0 | 0 | 0 | 2 |
[23] † Pityostrobus californiensis Last Modified in 01/30/22 | 0 | 33 | 33 | 0 | 0 | 0 | 0 | 2 |
[24] † Pityostrobus cliffwoodensis Last Modified in 01/30/22 | 4 | 29 | 29 | 0 | 0 | 0 | 0 | 2 |
[25] † Pityostrobus corneti Last Modified in 01/30/22 | 1 | 32 | 32 | 0 | 0 | 0 | 0 | 2 |
[26] † Pityostrobus hallii Last Modified in 01/30/22 | 1 | 32 | 32 | 0 | 0 | 0 | 0 | 2 |
[27] † Pityostrobus hautrageanus Last Modified in 01/30/22 | 3 | 30 | 30 | 0 | 0 | 0 | 0 | 2 |
[28] † Pityostrobus hokodzensis Last Modified in 01/30/22 | 0 | 33 | 33 | 0 | 0 | 0 | 0 | 2 |
[29] † Pityostrobus hueberi Last Modified in 01/30/22 | 0 | 33 | 33 | 0 | 0 | 0 | 0 | 2 |
[30] † Pityostrobus jacksonii Last Modified in 01/30/22 | 5 | 28 | 28 | 0 | 0 | 0 | 0 | 2 |
[31] † Pityostrobus kayeii Last Modified in 01/30/22 | 0 | 33 | 33 | 0 | 0 | 0 | 0 | 2 |
[32] † Pityostrobus leckenbyi Last Modified in 01/30/22 | 3 | 31 | 30 | 0 | 0 | 0 | 0 | 2 |
[33] † Pityostrobus lynnii Last Modified in 01/30/22 | 3 | 31 | 30 | 0 | 0 | 0 | 0 | 2 |
[34] † Pityostrobus macrocephalus Last Modified in 01/30/22 | 4 | 29 | 29 | 0 | 0 | 0 | 0 | 2 |
[35] † Pityostrobus makahensis Last Modified in 01/30/22 | 0 | 34 | 33 | 0 | 0 | 0 | 0 | 2 |
[36] † Pityostrobus mcmurryensis Last Modified in 01/30/22 | 1 | 32 | 32 | 0 | 0 | 0 | 0 | 2 |
[37] † Pityostrobus matsubarae Last Modified in 01/30/22 | 0 | 33 | 33 | 0 | 0 | 0 | 0 | 2 |
[38] † Pityostrobus milleri Last Modified in 01/30/22 | 3 | 30 | 30 | 0 | 0 | 0 | 0 | 2 |
[39] † Pityostrobus oblongus Last Modified in 01/30/22 | 0 | 33 | 33 | 0 | 0 | 0 | 0 | 2 |
[40] † Pityostrobus palmeri Last Modified in 01/30/22 | 0 | 34 | 33 | 0 | 0 | 0 | 0 | 2 |
[41] † Pityostrobus pubescens Last Modified in 01/30/22 | 0 | 33 | 33 | 0 | 0 | 0 | 0 | 2 |
[42] † Pityostrobus ramentosa Last Modified in 01/30/22 | 3 | 30 | 30 | 0 | 0 | 0 | 0 | 2 |
[43] † Pityostrobus shastaensis Last Modified in 01/30/22 | 2 | 31 | 31 | 0 | 0 | 0 | 0 | 2 |
[44] † Pityostrobus villerotensis Last Modified in 01/30/22 | 1 | 32 | 32 | 0 | 0 | 0 | 0 | 2 |
[45] † Pityostrobus virginiana Last Modified in 01/30/22 | 2 | 31 | 31 | 0 | 0 | 0 | 0 | 2 |
[46] Cryptomeria Last Modified in 01/30/22 | 3 | 30 | 30 | 0 | 0 | 0 | 0 | 2 |
[47] † Pararaucaria Last Modified in 01/30/22 | 1 | 33 | 32 | 0 | 0 | 0 | 0 | 2 |
[48] Sciadopitys Last Modified in 01/30/22 | 0 | 33 | 33 | 0 | 0 | 0 | 0 | 2 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 4471 | |
Project overview | 354 | |
Documents list | 233 | |
Media views | 1001 | Media search (847 views); M838524 (154 views); |
Matrix views | 270 | Matrix landing page (249 views); Matrix from Smith & Stockey 2002 (21 views); |
Taxon list | 1539 | |
Specimen list | 539 | |
Bibliography | 196 | |
Views for media list | 339 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 3 | |
Project downloads | 1 | |
Matrix downloads | 2 | Matrix from Smith & Stockey 2002 (2 downloads); |