Project 4192: G. Marramà, G. Carnevale, K. M. Claeson, G. J. Naylor, J. Kriwet. 2020. Revision of the Eocene ‘Platyrhina’ species from the Bolca Lagerstätte (Italy) reveals the first panray (Batomorphii: Zanobatidae) in the fossil record. Journal of Systematic Palaeontology. 18 (18):1519-1542.
Abstract
The fossil-Lagerstätte of Bolca (Italy) is well known for the diversity and exquisite preservation of its bony and cartilaginous fishes documenting tropical shallow-water marine environments associated with coral reefs in the western Tethys during the early Eocene. In this study, the taxonomic, systematic and phylogenetic position of two batoid species traditionally assigned to the living thornback ray genus Platyrhina is re-evaluated. †Platyrhina bolcensis Heckel, 1851 is recognized as a separate species of the Platyrhinidae because of its plate-like antorbital cartilage with an irregular outline and a small horn on the nasal capsules. Also, the rostral cartilage does not reach the anterior border of the disc. Support for the placement of this species within the new genus †Eoplatyrhina gen. nov. is based on a combination of morphological and meristic features (e.g. nasal capsules at right angles to the rostrum; large space between the hyomandibulae and mandibular arch; approximately 132 vertebral centra; 15–16 rib pairs; 81–87 pectoral radials; 18–21 pelvic radials; short, straight and stout claspers; 40–50 caudal-fin radials; thorns absent). A second species, †Platyrhina egertoni (De Zigno, 1876), is more closely related to the living panray Zanobatus than Platyrhina and is assigned here to †Plesiozanobatus gen. nov. because of a combination of characters that support its placement within the family Zanobatidae (tail stout and short, distinctly demarcated from disc; two dorsal fins and complete caudal fin; small dermal denticles and scattered thorns covering disc and tail; rostral cartilage absent; nasal capsules without horn-like processes; mesopterygium absent). The systematic position of a third taxon, †Platyrhina gigantea (Blainville, 1818), is currently impossible to establish due to the poor preservation of the only known specimen, and therefore we propose to consider it a nomen dubium. Palaeoecological and biogeographic features of the Eocene platyrhinids and zanobatids from Bolca are also discussed.Read the article »
Article DOI: 10.1080/14772019.2020.1783380
Project DOI: 10.7934/P4192, http://dx.doi.org/10.7934/P4192
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 7326 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 208 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 240 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:208/0 |
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MorphoBank Project 4192
MorphoBank Project 4192
- Creation Date:
08 February 2022 - Publication Date:
08 March 2022 - Project views: 6875
- Matrix downloads: 2
Authors' Institutions
- University of Florida
- Università degli Studi di Torino (University of Turin)
- University of Vienna
- Philadelphia College of Osteopathic Medicine
- Florida Museum of Natural History
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 42 | 1 | 1 | 208 | 0 | 0 | 7326 (7012, 0, 314) | 0 | 0 | 0 | ||||
Giuseppe Marrama Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M28706' (42 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Chimaeridae Last Modified in 02/08/22 | 11 | 87 | 87 | 0 | 6 | 0 | 0 | 2 |
[2] Hexanchidae Last Modified in 02/08/22 | 7 | 91 | 90 | 0 | 7 | 0 | 0 | 2 |
[3] Heterodontus Last Modified in 02/08/22 | 6 | 84 | 83 | 0 | 14 | 0 | 0 | 2 |
[4] Squalus Last Modified in 02/08/22 | 21 | 71 | 71 | 0 | 12 | 0 | 0 | 2 |
[5] Raja Last Modified in 02/08/22 | 0 | 103 | 103 | 0 | 1 | 0 | 0 | 2 |
[6] Bathyraja Last Modified in 02/08/22 | 0 | 102 | 102 | 0 | 2 | 0 | 0 | 2 |
[7] Torpedo Last Modified in 02/08/22 | 6 | 92 | 92 | 0 | 6 | 0 | 0 | 2 |
[8] Hypnos Last Modified in 02/08/22 | 9 | 89 | 89 | 0 | 6 | 0 | 0 | 2 |
[9] Narcine Last Modified in 02/08/22 | 5 | 95 | 94 | 0 | 4 | 0 | 0 | 2 |
[10] Narke Last Modified in 02/08/22 | 6 | 95 | 93 | 0 | 4 | 0 | 0 | 2 |
[11] Temera Last Modified in 02/08/22 | 7 | 91 | 90 | 0 | 6 | 0 | 0 | 2 |
[12] Platyrhina Last Modified in 02/08/22 | 0 | 102 | 98 | 0 | 3 | 0 | 0 | 2 |
[13] † Platyrhina bolcensis Last Modified in 02/08/22 | 54 | 48 | 48 | 0 | 2 | 0 | 0 | 2 |
[14] Platyrhinoidis Last Modified in 02/08/22 | 0 | 101 | 100 | 0 | 3 | 0 | 0 | 2 |
[15] † Tingitanius Last Modified in 02/08/22 | 51 | 50 | 50 | 0 | 3 | 0 | 0 | 2 |
[16] † Tethybatis Last Modified in 02/08/22 | 60 | 42 | 42 | 0 | 2 | 0 | 0 | 2 |
[17] Kimmerobatis Last Modified in 02/08/22 | 42 | 61 | 61 | 0 | 1 | 0 | 0 | 2 |
[18] Asterodermus Last Modified in 02/08/22 | 40 | 62 | 62 | 0 | 2 | 0 | 0 | 2 |
[19] Spathobatis Last Modified in 02/08/22 | 34 | 68 | 68 | 0 | 2 | 0 | 0 | 2 |
[20] Belemnobatis Last Modified in 02/08/22 | 35 | 67 | 67 | 0 | 2 | 0 | 0 | 2 |
[21] Pristis Last Modified in 02/08/22 | 3 | 98 | 97 | 0 | 3 | 0 | 0 | 2 |
[22] Rhynchobatus Last Modified in 02/08/22 | 4 | 98 | 98 | 0 | 2 | 0 | 0 | 2 |
[23] Glaucostegus Last Modified in 02/08/22 | 14 | 87 | 86 | 0 | 3 | 0 | 0 | 2 |
[24] Rhina Last Modified in 02/08/22 | 6 | 95 | 95 | 0 | 3 | 0 | 0 | 2 |
[25] Rhinobatos Last Modified in 02/08/22 | 0 | 102 | 102 | 0 | 2 | 0 | 0 | 2 |
[26] Trygonorrhina Last Modified in 02/08/22 | 1 | 101 | 101 | 0 | 2 | 0 | 0 | 2 |
[27] Zapteryx Last Modified in 02/08/22 | 0 | 102 | 102 | 0 | 2 | 0 | 0 | 2 |
[28] Aptychotrema Last Modified in 02/08/22 | 11 | 91 | 91 | 0 | 2 | 0 | 0 | 2 |
[29] Zanobatus Last Modified in 02/08/22 | 0 | 99 | 97 | 0 | 5 | 0 | 0 | 2 |
[30] † Platyrhina egertoni Last Modified in 02/08/22 | 66 | 35 | 35 | 0 | 3 | 0 | 0 | 2 |
[31] Urolophus Last Modified in 02/08/22 | 2 | 97 | 96 | 0 | 5 | 0 | 0 | 2 |
[32] Urobatis Last Modified in 02/08/22 | 5 | 95 | 94 | 0 | 4 | 0 | 0 | 2 |
[33] Urotygon Last Modified in 02/08/22 | 2 | 97 | 96 | 0 | 5 | 0 | 0 | 2 |
[34] Gymnura Last Modified in 02/08/22 | 3 | 96 | 95 | 0 | 5 | 0 | 0 | 2 |
[35] Myliobatis Last Modified in 02/08/22 | 2 | 99 | 99 | 0 | 3 | 0 | 0 | 2 |
[36] Aetobatus Last Modified in 02/08/22 | 3 | 98 | 98 | 0 | 3 | 0 | 0 | 2 |
[37] Rhinoptera Last Modified in 02/08/22 | 2 | 99 | 98 | 0 | 3 | 0 | 0 | 2 |
[38] Mobula Last Modified in 02/08/22 | 7 | 92 | 92 | 0 | 5 | 0 | 0 | 2 |
[39] † Ptychotrygon sp Last Modified in 02/08/22 | 49 | 52 | 52 | 0 | 3 | 0 | 0 | 2 |
[40] † Sclerorhynchus atavus Last Modified in 02/08/22 | 41 | 60 | 60 | 0 | 3 | 0 | 0 | 2 |
[41] † Libanopristis hiram Last Modified in 02/08/22 | 38 | 63 | 63 | 0 | 3 | 0 | 0 | 2 |
[42] † Asflapristis chistadentis Last Modified in 02/08/22 | 55 | 47 | 47 | 0 | 2 | 0 | 0 | 2 |
Taxonomic Overview for Matrix 'M28707' (42 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Chimaeridae Last Modified in 02/08/22 | 11 | 87 | 87 | 0 | 6 | 0 | 0 | 2 |
[2] Hexanchidae Last Modified in 02/08/22 | 7 | 91 | 90 | 0 | 7 | 0 | 0 | 2 |
[3] Heterodontus Last Modified in 02/08/22 | 6 | 86 | 85 | 0 | 12 | 0 | 0 | 2 |
[4] Squalus Last Modified in 02/08/22 | 21 | 73 | 73 | 0 | 10 | 0 | 0 | 2 |
[5] Raja Last Modified in 02/08/22 | 0 | 103 | 103 | 0 | 1 | 0 | 0 | 2 |
[6] Bathyraja Last Modified in 02/08/22 | 0 | 102 | 102 | 0 | 2 | 0 | 0 | 2 |
[7] Torpedo Last Modified in 02/08/22 | 6 | 92 | 92 | 0 | 6 | 0 | 0 | 2 |
[8] Hypnos Last Modified in 02/08/22 | 9 | 89 | 89 | 0 | 6 | 0 | 0 | 2 |
[9] Narcine Last Modified in 02/08/22 | 5 | 95 | 94 | 0 | 4 | 0 | 0 | 2 |
[10] Narke Last Modified in 02/08/22 | 6 | 95 | 93 | 0 | 4 | 0 | 0 | 2 |
[11] Temera Last Modified in 02/08/22 | 7 | 91 | 90 | 0 | 6 | 0 | 0 | 2 |
[12] Platyrhina Last Modified in 02/08/22 | 0 | 102 | 98 | 0 | 3 | 0 | 0 | 2 |
[13] † Platyrhina bolcensis Last Modified in 02/08/22 | 54 | 48 | 48 | 0 | 2 | 0 | 0 | 2 |
[14] Platyrhinoidis Last Modified in 02/08/22 | 0 | 101 | 100 | 0 | 3 | 0 | 0 | 2 |
[15] † Tingitanius Last Modified in 02/08/22 | 51 | 50 | 50 | 0 | 3 | 0 | 0 | 2 |
[16] † Tethybatis Last Modified in 02/08/22 | 60 | 42 | 42 | 0 | 2 | 0 | 0 | 2 |
[17] Kimmerobatis Last Modified in 02/08/22 | 42 | 61 | 61 | 0 | 1 | 0 | 0 | 2 |
[18] Asterodermus Last Modified in 02/08/22 | 40 | 62 | 62 | 0 | 2 | 0 | 0 | 2 |
[19] Spathobatis Last Modified in 02/08/22 | 34 | 68 | 68 | 0 | 2 | 0 | 0 | 2 |
[20] Belemnobatis Last Modified in 02/08/22 | 35 | 67 | 67 | 0 | 2 | 0 | 0 | 2 |
[21] Pristis Last Modified in 02/08/22 | 3 | 98 | 97 | 0 | 3 | 0 | 0 | 2 |
[22] Rhynchobatus Last Modified in 02/08/22 | 4 | 98 | 98 | 0 | 2 | 0 | 0 | 2 |
[23] Glaucostegus Last Modified in 02/08/22 | 14 | 87 | 86 | 0 | 3 | 0 | 0 | 2 |
[24] Rhina Last Modified in 02/08/22 | 6 | 95 | 95 | 0 | 3 | 0 | 0 | 2 |
[25] Rhinobatos Last Modified in 02/08/22 | 0 | 102 | 102 | 0 | 2 | 0 | 0 | 2 |
[26] Trygonorrhina Last Modified in 02/08/22 | 1 | 101 | 101 | 0 | 2 | 0 | 0 | 2 |
[27] Zapteryx Last Modified in 02/08/22 | 0 | 102 | 102 | 0 | 2 | 0 | 0 | 2 |
[28] Aptychotrema Last Modified in 02/08/22 | 11 | 91 | 91 | 0 | 2 | 0 | 0 | 2 |
[29] Zanobatus Last Modified in 02/08/22 | 0 | 99 | 97 | 0 | 5 | 0 | 0 | 2 |
[30] † Platyrhina egertoni Last Modified in 02/08/22 | 66 | 35 | 35 | 0 | 3 | 0 | 0 | 2 |
[31] Urolophus Last Modified in 02/08/22 | 2 | 97 | 96 | 0 | 5 | 0 | 0 | 2 |
[32] Urobatis Last Modified in 02/08/22 | 5 | 95 | 94 | 0 | 4 | 0 | 0 | 2 |
[33] Urotygon Last Modified in 02/08/22 | 2 | 97 | 96 | 0 | 5 | 0 | 0 | 2 |
[34] Gymnura Last Modified in 02/08/22 | 3 | 96 | 95 | 0 | 5 | 0 | 0 | 2 |
[35] Myliobatis Last Modified in 02/08/22 | 2 | 99 | 99 | 0 | 3 | 0 | 0 | 2 |
[36] Aetobatus Last Modified in 02/08/22 | 3 | 98 | 98 | 0 | 3 | 0 | 0 | 2 |
[37] Rhinoptera Last Modified in 02/08/22 | 2 | 99 | 98 | 0 | 3 | 0 | 0 | 2 |
[38] Mobula Last Modified in 02/08/22 | 7 | 92 | 92 | 0 | 5 | 0 | 0 | 2 |
[39] † Ptychotrygon sp Last Modified in 02/08/22 | 49 | 52 | 52 | 0 | 3 | 0 | 0 | 2 |
[40] † Sclerorhynchus atavus Last Modified in 02/08/22 | 41 | 60 | 60 | 0 | 3 | 0 | 0 | 2 |
[41] † Libanopristis hiram Last Modified in 02/08/22 | 38 | 63 | 63 | 0 | 3 | 0 | 0 | 2 |
[42] † Asflapristis chistadentis Last Modified in 02/08/22 | 55 | 47 | 47 | 0 | 2 | 0 | 0 | 2 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 6875 | |
Matrix views | 401 | Matrix landing page (327 views); Data matrix (DS coding) from Supplemental file 3 (48 views); Data matrix (CH coding) from Supplemental File 2 (26 views); |
Project overview | 447 | |
Media views | 1265 | Media search (1070 views); M838569 (195 views); |
Documents list | 315 | |
Views for media list | 645 | |
Taxon list | 2972 | |
Bibliography | 253 | |
Specimen list | 577 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 11 | |
Project downloads | 9 | |
Matrix downloads | 2 | Data matrix (DS coding) from Supplemental file 3 (1 download); Data matrix (CH coding) from Supplemental File 2 (1 download); |