Project 4397: D. Geraads, S. Zouhri. 2021. A new late Miocene elasmotheriine rhinoceros from Morocco. Acta Palaeontologica Polonica. 66 (4):753-765.
Abstract
We describe here the first definite representative of the subfamily Elasmotheriinae in North Africa. It comes from theupper Miocene site of Skoura near Ouarzazate, on the southern slope of the Central High Atlas in Morocco. It consistsof a virtually complete skull with articulated mandible and a few fragmentary postcranial remains, making it by far thebest known elasmotheriine from the African late Miocene. We assign it to a new taxon, Eoazara xerrii gen. et sp. nov.The skull is characterized by long nasal bones indicating a strong horn and long, anteriorly expanded, edentulous premaxillae.Compared to other Rhinocerotidae, the face is moderately elongated; the lower incisors are of medium size; andthe premolar row is short. The upper molars have a strongly pinched protocone, a long antecrochet, and an unexpandedcentral valley. Eoazara xerrii gen. et sp. nov. is at a lower evolutionary grade than the Chinese species of Ningxiatherium and Parelasmotherium, but probably comparable to the very incomplete remains from the East African late Mioceneforms. We regard Eoazara as a member of a chiefly Eurasian clade, rather than as a survivor of a hypothetical Africanelasmotheriine branch. Parsimony analysis confirms the monophyly of the Elasmotheriinae, but that of the remainingRhinocerotidae is questionable.Read the article »
Article DOI: 10.4202/app.00904.2021
Project DOI: 10.7934/P4397, http://dx.doi.org/10.7934/P4397
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 11333 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 223 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 537 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:15/208 |
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MorphoBank Project 4397
MorphoBank Project 4397
- Creation Date:
09 September 2022 - Publication Date:
16 September 2022 - Project views: 9894
- Media downloads: 1
- Matrix downloads: 5
Authors' Institutions
- Muséum national d'Histoire naturelle, Paris (Natural History Museum, Paris)
- Centre National de la Recherche Scientifique (CNRS)
- Sorbonne Université
- Hassan II University of Casablanca
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 75 | 12 | 10 | 223 | 0 | 0 | 11333 (10568, 0, 765) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M28982' (75 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Tapirus terrestris Last Modified in 09/16/22 | 15 | 186 | 186 | 0 | 22 | 0 | 0 | 1 |
[2] † Aceratherium incisivum Last Modified in 09/16/22 | 44 | 172 | 172 | 0 | 7 | 0 | 0 | 1 |
[3] † Acerorhinus fuguensis Last Modified in 09/16/22 | 43 | 170 | 170 | 0 | 10 | 0 | 0 | 1 |
[4] † Acerorhinus hezhengensis Last Modified in 09/16/22 | 35 | 176 | 176 | 0 | 12 | 0 | 0 | 1 |
[5] † Acerorhinus palaeosinensis Last Modified in 09/16/22 | 69 | 146 | 146 | 0 | 8 | 0 | 0 | 1 |
[6] † Acerorhinus tsaidamensis Last Modified in 09/16/22 | 34 | 176 | 176 | 0 | 13 | 0 | 0 | 1 |
[7] † Acerorhinus yuanmouensis Last Modified in 09/16/22 | 80 | 134 | 134 | 0 | 9 | 0 | 0 | 1 |
[8] † Acerorhinus zernowi Last Modified in 09/16/22 | 59 | 155 | 155 | 0 | 9 | 0 | 0 | 1 |
[9] † Alicornops complanatum Last Modified in 09/16/22 | 102 | 114 | 114 | 0 | 7 | 0 | 0 | 1 |
[10] † Alicornops simorrense Last Modified in 09/16/22 | 35 | 179 | 179 | 0 | 9 | 0 | 0 | 1 |
[11] † Aprotodon fatehjangense Last Modified in 09/16/22 | 89 | 130 | 130 | 0 | 4 | 0 | 0 | 1 |
[12] † Begertherium grimmi Last Modified in 09/16/22 | 85 | 128 | 128 | 0 | 10 | 0 | 0 | 1 |
[13] † Brachypotherium brachypus Last Modified in 09/16/22 | 33 | 177 | 177 | 0 | 13 | 0 | 0 | 1 |
[14] † Brachypotherium minor Last Modified in 09/16/22 | 92 | 119 | 119 | 0 | 12 | 0 | 0 | 1 |
[15] † Caementodon caucasicum Last Modified in 09/16/22 | 94 | 119 | 119 | 0 | 10 | 0 | 0 | 1 |
[16] Ceratotherium simum Last Modified in 09/16/22 | 18 | 187 | 187 | 0 | 18 | 0 | 0 | 1 |
[17] † Chilotheridium pattersoni Last Modified in 09/16/22 | 96 | 119 | 119 | 0 | 8 | 0 | 0 | 1 |
[18] † Chilotherium anderssoni Last Modified in 09/16/22 | 16 | 193 | 193 | 0 | 14 | 0 | 0 | 1 |
[19] † Chilotherium wimani Last Modified in 09/16/22 | 26 | 182 | 182 | 0 | 15 | 0 | 0 | 1 |
[20] † Coelodonta antiquitatis Last Modified in 09/16/22 | 28 | 181 | 181 | 0 | 14 | 0 | 0 | 1 |
[21] † Diaceratherium aginense Last Modified in 09/16/22 | 72 | 142 | 142 | 0 | 9 | 0 | 0 | 1 |
[22] † Diceratherium armatum Last Modified in 09/16/22 | 16 | 195 | 195 | 0 | 12 | 0 | 0 | 1 |
[23] Dicerorhinus sumatrensis Last Modified in 09/16/22 | 30 | 179 | 179 | 0 | 14 | 0 | 0 | 1 |
[24] Diceros bicornis Last Modified in 09/16/22 | 13 | 189 | 189 | 0 | 21 | 0 | 0 | 1 |
[25] † Elasmotherium sibiricum Last Modified in 09/16/22 | 43 | 149 | 149 | 0 | 31 | 0 | 0 | 1 |
[26] † Eoazara xerrii Last Modified in 09/16/22 | 86 | 122 | 122 | 0 | 15 | 0 | 0 | 1 |
[27] † Gaindatherium browni Last Modified in 09/16/22 | 57 | 153 | 153 | 0 | 13 | 0 | 0 | 1 |
[28] † Hispanotherium beonense Last Modified in 09/16/22 | 38 | 177 | 177 | 0 | 8 | 0 | 0 | 1 |
[29] † Hispanotherium corcolense Last Modified in 09/16/22 | 93 | 125 | 125 | 0 | 5 | 0 | 0 | 1 |
[30] † Hispanotherium matritense Last Modified in 09/16/22 | 75 | 142 | 142 | 0 | 6 | 0 | 0 | 1 |
[31] † Hispanotherium tungurense Last Modified in 09/16/22 | 28 | 184 | 184 | 0 | 11 | 0 | 0 | 1 |
[32] † Hoploaceratherium tetradactylum Last Modified in 09/16/22 | 37 | 177 | 177 | 0 | 9 | 0 | 0 | 1 |
[33] † Hyrachyus eximius Last Modified in 09/16/22 | 32 | 168 | 168 | 0 | 23 | 0 | 0 | 1 |
[34] † Iranotherium morgani Last Modified in 09/16/22 | 42 | 167 | 167 | 0 | 14 | 0 | 0 | 1 |
[35] † Lartetotherium sansaniense Last Modified in 09/16/22 | 44 | 166 | 166 | 0 | 13 | 0 | 0 | 1 |
[36] † Menoceras arikarense Last Modified in 09/16/22 | 18 | 194 | 194 | 0 | 11 | 0 | 0 | 1 |
[37] † Mesaceratherium gaimersheimense Last Modified in 09/16/22 | 69 | 144 | 144 | 0 | 10 | 0 | 0 | 1 |
[38] † Mesaceratherium paulhiacense Last Modified in 09/16/22 | 94 | 122 | 122 | 0 | 7 | 0 | 0 | 1 |
[39] † Mesaceratherium welcommi Last Modified in 09/16/22 | 73 | 138 | 138 | 0 | 12 | 0 | 0 | 1 |
[40] † Molassitherium albigense Last Modified in 09/16/22 | 39 | 174 | 174 | 0 | 10 | 0 | 0 | 1 |
[41] † Ningxiatherium euryrhinus Last Modified in 09/16/22 | 99 | 117 | 117 | 0 | 7 | 0 | 0 | 1 |
[42] † Ningxiatherium longirhinus Last Modified in 09/16/22 | 103 | 113 | 113 | 0 | 7 | 0 | 0 | 1 |
[43] † Parelasmotherium lingxiaense Last Modified in 09/16/22 | 90 | 124 | 124 | 0 | 9 | 0 | 0 | 1 |
[44] † Plesiaceratherium gracile Last Modified in 09/16/22 | 51 | 159 | 159 | 0 | 13 | 0 | 0 | 1 |
[45] † Plesiaceratherium mirallesi Last Modified in 09/16/22 | 49 | 165 | 165 | 0 | 9 | 0 | 0 | 1 |
[46] † Pleuroceros blanfordi Last Modified in 09/16/22 | 48 | 168 | 168 | 0 | 7 | 0 | 0 | 1 |
[47] † Pleuroceros pleuroceros Last Modified in 09/16/22 | 61 | 156 | 156 | 0 | 6 | 0 | 0 | 1 |
[48] † Prosantorhinus douvillei Last Modified in 09/16/22 | 50 | 163 | 163 | 0 | 10 | 0 | 0 | 1 |
[49] † Protaceratherium minutum Last Modified in 09/16/22 | 55 | 158 | 158 | 0 | 10 | 0 | 0 | 1 |
[50] Rhinoceros sondaicus Last Modified in 09/16/22 | 15 | 192 | 192 | 0 | 16 | 0 | 0 | 1 |
[51] Rhinoceros unicornis Last Modified in 09/16/22 | 17 | 192 | 192 | 0 | 14 | 0 | 0 | 1 |
[52] † Ronzotherium filholi Last Modified in 09/16/22 | 19 | 190 | 190 | 0 | 14 | 0 | 0 | 1 |
[53] † Rusingaceros leakeyi Last Modified in 09/16/22 | 92 | 123 | 123 | 0 | 8 | 0 | 0 | 1 |
[54] † Shansirhinus ringstromi Last Modified in 09/16/22 | 41 | 170 | 170 | 0 | 12 | 0 | 0 | 1 |
[55] † Subhyracodon occidentalis Last Modified in 09/16/22 | 43 | 171 | 171 | 0 | 9 | 0 | 0 | 1 |
[56] † Teleoceras fossiger Last Modified in 09/16/22 | 18 | 190 | 190 | 0 | 15 | 0 | 0 | 1 |
[57] † Trigonias osborni Last Modified in 09/16/22 | 15 | 192 | 192 | 0 | 16 | 0 | 0 | 1 |
[58] † Turkanatherium acutirostratum Last Modified in 09/16/22 | 88 | 126 | 126 | 0 | 9 | 0 | 0 | 1 |
[59] † Victoriaceros kenyensis Last Modified in 09/16/22 | 52 | 157 | 157 | 0 | 14 | 0 | 0 | 1 |
[60] † Aceratherium huadeensis Last Modified in 09/16/22 | 141 | 77 | 77 | 0 | 5 | 0 | 0 | 1 |
[61] † Acerorhinus lufengensis Last Modified in 09/16/22 | 131 | 86 | 86 | 0 | 6 | 0 | 0 | 1 |
[62] † Alicornops alfambrense Last Modified in 09/16/22 | 186 | 36 | 36 | 0 | 1 | 0 | 0 | 1 |
[63] † Begertherium borissiaki Last Modified in 09/16/22 | 155 | 65 | 65 | 0 | 3 | 0 | 0 | 1 |
[64] † Beliajevina tekkayai Last Modified in 09/16/22 | 118 | 102 | 102 | 0 | 3 | 0 | 0 | 1 |
[65] † Bugtirhinus praecursor Last Modified in 09/16/22 | 135 | 85 | 85 | 0 | 3 | 0 | 0 | 1 |
[66] † Caementodon oettingenae Last Modified in 09/16/22 | 126 | 94 | 94 | 0 | 3 | 0 | 0 | 1 |
[67] † Elasmotherium caucasicum Last Modified in 09/16/22 | 111 | 90 | 90 | 0 | 22 | 0 | 0 | 1 |
[68] † Kenyatherium bishopi Last Modified in 09/16/22 | 184 | 38 | 38 | 0 | 1 | 0 | 0 | 1 |
[69] † Ougandatherium napakense Last Modified in 09/16/22 | 146 | 69 | 69 | 0 | 8 | 0 | 0 | 1 |
[70] † Parelasmotherium schansiense Last Modified in 09/16/22 | 183 | 39 | 39 | 0 | 1 | 0 | 0 | 1 |
[71] † Procoelodonta mongoliense Last Modified in 09/16/22 | 142 | 77 | 77 | 0 | 4 | 0 | 0 | 1 |
[72] † Shennongtherium hyposodontus Last Modified in 09/16/22 | 184 | 37 | 37 | 0 | 2 | 0 | 0 | 1 |
[73] † Sinotherium lagrelii Last Modified in 09/16/22 | 121 | 95 | 95 | 0 | 7 | 0 | 0 | 1 |
[74] † Subchilotherium intermedium Last Modified in 09/16/22 | 115 | 102 | 102 | 0 | 6 | 0 | 0 | 1 |
[75] † Victoriaceros hooijeri Last Modified in 09/16/22 | 116 | 100 | 100 | 0 | 7 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 9894 | |
Project overview | 469 | |
Media views | 2499 | Media search (875 views); M846869 (163 views); M846868 (156 views); M846871 (143 views); M846870 (174 views); M846863 (162 views); M846862 (158 views); M846872 (160 views); M846867 (163 views); M846873 (173 views); M846866 (172 views); |
Matrix views | 291 | Matrix landing page (271 views); matrix (20 views); |
Taxon list | 2974 | |
Views for media list | 592 | |
Bibliography | 223 | |
Specimen list | 2840 | |
Documents list | 6 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 7 | |
Matrix downloads | 5 | matrix (5 downloads); |
Project downloads | 1 | |
Media downloads | 1 | M846872 (1 download); |