Project 4422: H. O. Berks, M. Lunde Nielsen, J. FlanneryâSutherland, A. Thorshøj Nielsen, T. S. Park, J. Vinther. 2023. A possibly deep branching artiopodan arthropod from the lower Cambrian Sirius Passet Lagerstätte (North Greenland). Papers in Palaeontology. 9 (3):e1495.
Abstract
Artiopoda was a diverse group of Palaeozoic euarthropods that proliferated in the early Palaeozoic, epitomized by the ubiquitous trilobites. Their possible phylogenetic position outside mandibulates and chelicerates offers the potential for understanding the evolution of euarthropods in more detail. However, this opportunity remains unexploited given that identification of deep‐splitting artiopodans remains to be fully explored. Here, we describe a new non‐trilobite artiopodan from the lower Cambrian Sirius Passet Lagerstätte, North Greenland. Thulaspis tholops gen. et sp. nov. is a large species with a broad, domed head shield, followed by a trunk consisting of 15 thoracic tergites and a small pygidium, giving the body an ovoid appearance when viewed dorsally. Thulaspis is distinctive with its rounded genae and anterior thoracic pleural tips, as well as short pleural spines posteriorly. A heart‐shaped hypostome with an anterior lobe is present. Appendages, partly obscured by the tergal skeleton, have many moderate length gnathobasic spines, and large flap‐like exopods with a fringe of small setae. Cladistic analyses recover Thulaspis as the sister taxon to Squamacula, a genus found in the Chengjiang and Emu Bay Shale biotas, in either a polytomy with a number of artiopodan taxa or as a sister group to all other artiopodans, indicating an important role in understanding the roots of artiopodan anatomy and evolution.Read the article »
Article DOI: 10.1002/spp2.1495
Project DOI: 10.7934/P4422, http://dx.doi.org/10.7934/P4422
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 5091 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 93 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 214 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:93/0 |
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MorphoBank Project 4422
MorphoBank Project 4422
- Creation Date:
27 September 2022 - Publication Date:
18 May 2023 - Project views: 4160
- Media downloads: 1
- Matrix downloads: 2
Authors' Institutions
- Copenhagen University
- University of Bristol
- British Geological Survey
- Korea Polar Research Institute
- University of Science & Technology, Daejeon
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Harry Berks Project Administrator | 67 | 1 | 1 | 93 | 0 | 0 | 5091 (3866, 0, 1225) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29111' (67 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Fortiforceps foliosa Last Modified in 05/18/23 | 1 | 72 | 72 | 0 | 20 | 0 | 0 | 1 |
[2] † Acanthomeridion anacanthus Last Modified in 05/18/23 | 25 | 48 | 48 | 0 | 20 | 0 | 0 | 1 |
[3] † Acanthomeridion serratum Last Modified in 05/18/23 | 25 | 48 | 48 | 0 | 20 | 0 | 0 | 1 |
[4] † Aglaspella granulifera Last Modified in 05/18/23 | 23 | 60 | 60 | 0 | 10 | 0 | 0 | 1 |
[5] † Aglaspis spinifer Last Modified in 05/18/23 | 19 | 64 | 64 | 0 | 10 | 0 | 0 | 1 |
[6] † Alalcomenaeus cambricus Last Modified in 05/18/23 | 1 | 71 | 71 | 0 | 21 | 0 | 0 | 1 |
[7] † Australaglaspis stonyensis Last Modified in 05/18/23 | 30 | 53 | 53 | 0 | 10 | 0 | 0 | 1 |
[8] † Australimicola spriggi Last Modified in 05/18/23 | 19 | 51 | 51 | 0 | 23 | 0 | 0 | 1 |
[9] † Beckwithia typa Last Modified in 05/18/23 | 31 | 52 | 52 | 0 | 10 | 0 | 0 | 1 |
[10] † Brachyaglaspis singularis Last Modified in 05/18/23 | 35 | 43 | 43 | 0 | 15 | 0 | 0 | 1 |
[11] † Buenaspis Last Modified in 05/18/23 | 28 | 47 | 47 | 0 | 18 | 0 | 0 | 1 |
[12] † Burgessia bella Last Modified in 05/18/23 | 1 | 62 | 62 | 0 | 30 | 0 | 0 | 1 |
[13] † Cheloniellon calmani Last Modified in 05/18/23 | 4 | 71 | 71 | 0 | 18 | 0 | 0 | 1 |
[14] † Chlupacaris dubia Last Modified in 05/18/23 | 40 | 43 | 43 | 0 | 10 | 0 | 0 | 1 |
[15] † Chraspedops Last Modified in 05/18/23 | 32 | 50 | 50 | 0 | 11 | 0 | 0 | 1 |
[16] † Cindarella eucalla Last Modified in 05/18/23 | 0 | 73 | 73 | 0 | 20 | 0 | 0 | 1 |
[17] † Cyclopites vulgaris Last Modified in 05/18/23 | 26 | 57 | 57 | 0 | 10 | 0 | 0 | 1 |
[18] † Duslia insignis Last Modified in 05/18/23 | 29 | 51 | 51 | 0 | 13 | 0 | 0 | 1 |
[19] † Emeraldella brockii Last Modified in 05/18/23 | 1 | 71 | 71 | 0 | 21 | 0 | 0 | 1 |
[20] † Eoredlichia intermedia Last Modified in 05/18/23 | 1 | 74 | 74 | 0 | 18 | 0 | 0 | 1 |
[21] † Eozetetes gemelli Last Modified in 05/18/23 | 31 | 51 | 51 | 0 | 11 | 0 | 0 | 1 |
[22] † Flobertia kochi Last Modified in 05/18/23 | 46 | 43 | 43 | 0 | 4 | 0 | 0 | 1 |
[23] † Glypharthrus magnoculus Last Modified in 05/18/23 | 36 | 47 | 47 | 0 | 10 | 0 | 0 | 1 |
[24] † Glypharthrus simplex Last Modified in 05/18/23 | 24 | 58 | 58 | 0 | 11 | 0 | 0 | 1 |
[25] † Glypharthrus thomasi Last Modified in 05/18/23 | 33 | 50 | 50 | 0 | 10 | 0 | 0 | 1 |
[26] † Glypharthrus trispinicaudatus Last Modified in 05/18/23 | 35 | 49 | 49 | 0 | 9 | 0 | 0 | 1 |
[27] † Gogglops Last Modified in 05/18/23 | 36 | 48 | 48 | 0 | 9 | 0 | 0 | 1 |
[28] † Haifengella corona Last Modified in 05/18/23 | 25 | 50 | 50 | 0 | 18 | 0 | 0 | 1 |
[29] † Haikoucaris Last Modified in 05/18/23 | 0 | 72 | 72 | 0 | 21 | 0 | 0 | 1 |
[30] † Helmetia expansa Last Modified in 05/18/23 | 21 | 54 | 54 | 0 | 18 | 0 | 0 | 1 |
[31] † Kodymirus vagans Last Modified in 05/18/23 | 12 | 63 | 63 | 0 | 18 | 0 | 0 | 1 |
[32] † Kuamaia Last Modified in 05/18/23 | 4 | 68 | 68 | 0 | 21 | 0 | 0 | 1 |
[33] † Kwanyinaspis maotiashanensis Last Modified in 05/18/23 | 14 | 66 | 66 | 0 | 13 | 0 | 0 | 1 |
[34] † Leanchoilia superlata Last Modified in 05/18/23 | 2 | 69 | 69 | 0 | 22 | 0 | 0 | 1 |
[35] † Liwia convexa Last Modified in 05/18/23 | 19 | 53 | 53 | 0 | 21 | 0 | 0 | 1 |
[36] † Luohuilinella deletres Last Modified in 05/18/23 | 6 | 66 | 66 | 0 | 21 | 0 | 0 | 1 |
[37] † Luohuilinella rarus Last Modified in 05/18/23 | 42 | 40 | 40 | 0 | 11 | 0 | 0 | 1 |
[38] † Marrella splendens Last Modified in 05/18/23 | 2 | 57 | 57 | 0 | 34 | 0 | 0 | 1 |
[39] † Martinssonia elongata Last Modified in 05/18/23 | 1 | 58 | 58 | 0 | 34 | 0 | 0 | 1 |
[40] † Mimetaster hexagonalis Last Modified in 05/18/23 | 1 | 64 | 64 | 0 | 28 | 0 | 0 | 1 |
[41] † Misszhouia longicaudata Last Modified in 05/18/23 | 2 | 67 | 67 | 0 | 24 | 0 | 0 | 1 |
[42] † Naraoia compacta Last Modified in 05/18/23 | 3 | 67 | 67 | 0 | 23 | 0 | 0 | 1 |
[43] † Naraoia spinosa Last Modified in 05/18/23 | 1 | 68 | 68 | 0 | 24 | 0 | 0 | 1 |
[44] Nebalia bipes Last Modified in 05/18/23 | 0 | 64 | 64 | 0 | 29 | 0 | 0 | 1 |
[45] † Neostrabops Last Modified in 05/18/23 | 43 | 43 | 43 | 0 | 7 | 0 | 0 | 1 |
[46] † Olenoides serratus Last Modified in 05/18/23 | 1 | 73 | 73 | 0 | 19 | 0 | 0 | 1 |
[47] † Phytophilaspis pergamena Last Modified in 05/18/23 | 25 | 52 | 52 | 0 | 16 | 0 | 0 | 1 |
[48] † Pygmaclypeatus daziensis Last Modified in 05/18/23 | 1 | 69 | 69 | 0 | 23 | 0 | 0 | 1 |
[49] † Quasimodaspis brentsae Last Modified in 05/18/23 | 33 | 50 | 50 | 0 | 10 | 0 | 0 | 1 |
[50] † Rebachiella kinnekullensis Last Modified in 05/18/23 | 1 | 63 | 63 | 0 | 29 | 0 | 0 | 1 |
[51] † Retifacies abnormalis Last Modified in 05/18/23 | 1 | 67 | 67 | 0 | 25 | 0 | 0 | 1 |
[52] † Saperion Last Modified in 05/18/23 | 6 | 61 | 61 | 0 | 26 | 0 | 0 | 1 |
[53] † Sidneyia inexpectans Last Modified in 05/18/23 | 1 | 75 | 75 | 0 | 17 | 0 | 0 | 1 |
[54] † Sinoburius lunaris Last Modified in 05/18/23 | 2 | 72 | 72 | 0 | 19 | 0 | 0 | 1 |
[55] † Skioldia Last Modified in 05/18/23 | 11 | 56 | 56 | 0 | 26 | 0 | 0 | 1 |
[56] † Soomaspis Last Modified in 05/18/23 | 28 | 47 | 47 | 0 | 18 | 0 | 0 | 1 |
[57] † Squamacula clypeata Last Modified in 05/18/23 | 1 | 60 | 60 | 0 | 32 | 0 | 0 | 1 |
[58] † Tariccoia Last Modified in 05/18/23 | 27 | 48 | 48 | 0 | 18 | 0 | 0 | 1 |
[59] † Tegopelte gigas Last Modified in 05/18/23 | 18 | 50 | 50 | 0 | 25 | 0 | 0 | 1 |
[60] † Tremaglaspis unite Last Modified in 05/18/23 | 26 | 49 | 49 | 0 | 18 | 0 | 0 | 1 |
[61] † Tremaglaspis vanroyi Last Modified in 05/18/23 | 34 | 43 | 43 | 0 | 16 | 0 | 0 | 1 |
[62] † Triopus draboviensis morocco Last Modified in 05/18/23 | 31 | 49 | 49 | 0 | 13 | 0 | 0 | 1 |
[63] † Uarthrus instabilis Last Modified in 05/18/23 | 31 | 53 | 53 | 0 | 9 | 0 | 0 | 1 |
[64] † Xandarella spectacullum Last Modified in 05/18/23 | 1 | 73 | 73 | 0 | 19 | 0 | 0 | 1 |
[65] † Zhiwenia coronata Last Modified in 05/18/23 | 15 | 59 | 59 | 0 | 19 | 0 | 0 | 1 |
[66] † Squamacula buckorum Last Modified in 05/18/23 | 27 | 42 | 42 | 0 | 24 | 0 | 0 | 1 |
[67] † Thulaspis tholops Last Modified in 05/18/23 | 9 | 59 | 59 | 0 | 25 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 4160 | |
Project overview | 307 | |
Media views | 709 | Media search (528 views); M862361 (181 views); |
Taxon list | 2047 | |
Matrix views | 245 | Matrix landing page (227 views); Artiopoda matrix (18 views); |
Specimen list | 355 | |
Documents list | 163 | |
Views for media list | 204 | |
Bibliography | 130 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 11 | |
Project downloads | 8 | |
Media downloads | 1 | M862361 (1 download); |
Matrix downloads | 2 | Artiopoda matrix (2 downloads); |