Project 4434: M. Medrano, A. B. Kury, A. C. Mendes. 2022. Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae). Zoological Journal of the Linnean Society. 195 (2):585-672.
Abstract
The cosmetids are conspicuous harvestmen, remarkably diverse in size, shape and colour. However, the effectiveness of all these morphological traits for diagnosing groups is scarcely explored in the literature. Since the early 20th century, the family Cosmetidae has been divided into two subfamilies, Cosmetinae and Discosomaticinae, and there has been no further effort to delimit additional or alternative subfamilial groups.In this analysis, we aim to test the issue of the basal dichotomy of Cosmetinae and Discosomaticinae. Thus, we propose a more comprehensive phylogenetic analysis using both parsimony and Bayesian approach, comprising 103 terminals of Cosmetidae, plus seven outgroup terminals scored for 130 morphological characters. Discosomaticinae is revisited and all its species were included in our matrix. To offer a real challenge to the monophyly of Discosomaticinae, members of 36 genera of Cosmetinae have also been scored. Our results support neither Cosmetinae nor Discosomaticinae sensu Pickard-Cambridge/Roewer as monophyletic groups. We found that Cosmetidae may be organized into a few major clades, which are here diagnosed. Accordingly, taxonomic changes in the arrangement of the family are introduced: description of one new tribe, five new subfamilies and revalidation of a sixth, description of three new genera, and proposition of synonymies, transfers and revalidations at generic level.Read the article »
Article DOI: 10.1093/zoolinnean/zlab043
Project DOI: 10.7934/P4434, http://dx.doi.org/10.7934/P4434
This project contains | Matrices |
---|---|
Download Project SDD File | Total scored cells: 12727 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 130 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 334 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:123/7 |
Currently Viewing:
MorphoBank Project 4434
MorphoBank Project 4434
- Creation Date:
06 October 2022 - Publication Date:
06 October 2022 - Project views: 5742
- Matrix downloads: 7
Authors' Institutions
- Universidade Federal do Rio De Janeiro (UFRJ)
- Universidade do Estado do Rio de Janeiro
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Miguel Medrano Project Administrator | 111 | 1 | 1 | 130 | 0 | 0 | 12727 (10976, 0, 1751) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29010' (110 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Pseudopucrolia mutica Last Modified in 10/06/22 | 4 | 99 | 99 | 0 | 27 | 0 | 0 | 1 |
[2] Quindina bella Last Modified in 10/06/22 | 4 | 102 | 102 | 0 | 24 | 0 | 0 | 1 |
[3] Syncranaus cribrum Last Modified in 10/06/22 | 6 | 102 | 102 | 0 | 22 | 0 | 0 | 1 |
[4] Chacoikeontus clavifemur Last Modified in 10/06/22 | 5 | 98 | 98 | 0 | 27 | 0 | 0 | 1 |
[5] Incasarcus dianae Last Modified in 10/06/22 | 7 | 100 | 99 | 0 | 24 | 0 | 0 | 1 |
[6] Prostygnus vestitus Last Modified in 10/06/22 | 5 | 105 | 105 | 0 | 20 | 0 | 0 | 1 |
[7] Yania flavolimbata Last Modified in 10/06/22 | 4 | 99 | 99 | 0 | 27 | 0 | 0 | 1 |
[8] Cynorta lithoclasica Last Modified in 10/06/22 | 13 | 105 | 105 | 0 | 12 | 0 | 0 | 1 |
[9] Cynorta quibijana Last Modified in 10/06/22 | 16 | 103 | 103 | 0 | 11 | 0 | 0 | 1 |
[10] Cynortoides cubana cubana Last Modified in 10/06/22 | 15 | 104 | 104 | 0 | 11 | 0 | 0 | 1 |
[11] Cynortoides v-album Last Modified in 10/06/22 | 8 | 112 | 111 | 0 | 11 | 0 | 0 | 1 |
[12] Vonones octotuberculatus Last Modified in 10/06/22 | 40 | 78 | 78 | 0 | 12 | 0 | 0 | 1 |
[13] Vonones planus Last Modified in 10/06/22 | 5 | 113 | 113 | 0 | 12 | 0 | 0 | 1 |
[14] Holovonones compressus Last Modified in 10/06/22 | 13 | 101 | 101 | 0 | 16 | 0 | 0 | 1 |
[15] Vonones ornatus Last Modified in 10/06/22 | 4 | 112 | 111 | 0 | 15 | 0 | 0 | 1 |
[16] Vonones sayi Last Modified in 10/06/22 | 6 | 111 | 111 | 0 | 13 | 0 | 0 | 1 |
[17] Libitioides albolineata Last Modified in 10/06/22 | 5 | 112 | 112 | 0 | 13 | 0 | 0 | 1 |
[18] Arucillus armasi Last Modified in 10/06/22 | 10 | 102 | 101 | 0 | 19 | 0 | 0 | 1 |
[19] Trinimontius darlingtoni Last Modified in 10/06/22 | 4 | 110 | 110 | 0 | 16 | 0 | 0 | 1 |
[20] Erginulus clavotibialis Last Modified in 10/06/22 | 5 | 102 | 102 | 0 | 23 | 0 | 0 | 1 |
[21] Meterginus basalis Last Modified in 10/06/22 | 6 | 111 | 111 | 0 | 13 | 0 | 0 | 1 |
[22] Poecilaemula signata Last Modified in 10/06/22 | 8 | 107 | 107 | 0 | 15 | 0 | 0 | 1 |
[23] Paragryne quadrimaculata Last Modified in 10/06/22 | 35 | 82 | 82 | 0 | 13 | 0 | 0 | 1 |
[24] Metagryne ferruginea Last Modified in 10/06/22 | 35 | 82 | 82 | 0 | 13 | 0 | 0 | 1 |
[25] Metagryne albireticulata Last Modified in 10/06/22 | 39 | 81 | 81 | 0 | 10 | 0 | 0 | 1 |
[26] Peruvicynorta oxapampa Last Modified in 10/06/22 | 38 | 80 | 80 | 0 | 12 | 0 | 0 | 1 |
[27] Acantholibitia pustulosa Last Modified in 10/06/22 | 38 | 81 | 81 | 0 | 11 | 0 | 0 | 1 |
[28] Gryne pluriarcuata Last Modified in 10/06/22 | 3 | 115 | 115 | 0 | 12 | 0 | 0 | 1 |
[29] Fortalezius excellens Last Modified in 10/06/22 | 36 | 83 | 83 | 0 | 11 | 0 | 0 | 1 |
[30] Bodunius biocellatus Last Modified in 10/06/22 | 3 | 115 | 115 | 0 | 12 | 0 | 0 | 1 |
[31] Metalibitia paraguayensis Last Modified in 10/06/22 | 2 | 94 | 94 | 0 | 34 | 0 | 0 | 1 |
[32] Ferkeria vestita Last Modified in 10/06/22 | 6 | 106 | 106 | 0 | 18 | 0 | 0 | 1 |
[33] Platygyndes titicaca Last Modified in 10/06/22 | 12 | 84 | 84 | 0 | 34 | 0 | 0 | 1 |
[34] Paecilaemella festae Last Modified in 10/06/22 | 39 | 71 | 70 | 0 | 21 | 0 | 0 | 1 |
[35] Paecilaemella multimaculata Last Modified in 10/06/22 | 45 | 65 | 65 | 0 | 20 | 0 | 0 | 1 |
[36] Protus ornatus Last Modified in 10/06/22 | 37 | 73 | 73 | 0 | 20 | 0 | 0 | 1 |
[37] Flirtea picta Last Modified in 10/06/22 | 4 | 110 | 110 | 0 | 16 | 0 | 0 | 1 |
[38] Flirtea valida Last Modified in 10/06/22 | 8 | 106 | 106 | 0 | 16 | 0 | 0 | 1 |
[39] Acritas bilineatus Last Modified in 10/06/22 | 3 | 103 | 103 | 0 | 24 | 0 | 0 | 1 |
[40] Vononoides unimaculatus Last Modified in 10/06/22 | 7 | 106 | 106 | 0 | 17 | 0 | 0 | 1 |
[41] Chusgonobius weyrauchi Last Modified in 10/06/22 | 42 | 76 | 76 | 0 | 12 | 0 | 0 | 1 |
[42] Vononella maculata Last Modified in 10/06/22 | 47 | 69 | 69 | 0 | 14 | 0 | 0 | 1 |
[43] Eucynortella cryptogramma Last Modified in 10/06/22 | 30 | 86 | 86 | 0 | 14 | 0 | 0 | 1 |
[44] Cynortopyga pauper Last Modified in 10/06/22 | 23 | 92 | 92 | 0 | 15 | 0 | 0 | 1 |
[45] Vononana peruviana Last Modified in 10/06/22 | 43 | 72 | 72 | 0 | 15 | 0 | 0 | 1 |
[46] Guaporeius patellaris Last Modified in 10/06/22 | 14 | 102 | 102 | 0 | 14 | 0 | 0 | 1 |
[47] Guaporeius sanctissimus Last Modified in 10/06/22 | 26 | 90 | 90 | 0 | 14 | 0 | 0 | 1 |
[48] Taito adrik Last Modified in 10/06/22 | 5 | 102 | 102 | 0 | 23 | 0 | 0 | 1 |
[49] Taito juruensis Last Modified in 10/06/22 | 5 | 109 | 109 | 0 | 16 | 0 | 0 | 1 |
[50] Taito spaceinvaders Last Modified in 10/06/22 | 4 | 110 | 110 | 0 | 16 | 0 | 0 | 1 |
[51] Taito honda Last Modified in 10/06/22 | 5 | 109 | 109 | 0 | 16 | 0 | 0 | 1 |
[52] Rhauculus insignitus Last Modified in 10/06/22 | 5 | 111 | 111 | 0 | 14 | 0 | 0 | 1 |
[53] Rhaucoides ornatus Last Modified in 10/06/22 | 1 | 117 | 117 | 0 | 12 | 0 | 0 | 1 |
[54] Rhaucoides riveti Last Modified in 10/06/22 | 3 | 115 | 115 | 0 | 12 | 0 | 0 | 1 |
[55] Eucynorta virescens Last Modified in 10/06/22 | 2 | 116 | 116 | 0 | 12 | 0 | 0 | 1 |
[56] Rhaucus vulneratus Last Modified in 10/06/22 | 1 | 115 | 115 | 0 | 14 | 0 | 0 | 1 |
[57] Rhaucus robustus Last Modified in 10/06/22 | 1 | 115 | 115 | 0 | 14 | 0 | 0 | 1 |
[58] Eulibitia scalaris Last Modified in 10/06/22 | 1 | 113 | 112 | 0 | 17 | 0 | 0 | 1 |
[59] Eulibitia h-inscriptum Last Modified in 10/06/22 | 1 | 112 | 112 | 0 | 17 | 0 | 0 | 1 |
[60] Ambatoiella vigilans Last Modified in 10/06/22 | 3 | 105 | 105 | 0 | 22 | 0 | 0 | 1 |
[61] Oligovonones brunneus Last Modified in 10/06/22 | 11 | 95 | 95 | 0 | 24 | 0 | 0 | 1 |
[62] Libitia cordata Last Modified in 10/06/22 | 1 | 104 | 104 | 0 | 25 | 0 | 0 | 1 |
[63] Libitia bipunctata Last Modified in 10/06/22 | 1 | 104 | 104 | 0 | 25 | 0 | 0 | 1 |
[64] Neocynorta marmorata Last Modified in 10/06/22 | 1 | 115 | 114 | 0 | 15 | 0 | 0 | 1 |
[65] Neocynorta venezuelensis Last Modified in 10/06/22 | 4 | 111 | 110 | 0 | 16 | 0 | 0 | 1 |
[66] Cynorta conspersa Last Modified in 10/06/22 | 3 | 112 | 111 | 0 | 16 | 0 | 0 | 1 |
[67] Gryne pustulata Last Modified in 10/06/22 | 41 | 77 | 77 | 0 | 12 | 0 | 0 | 1 |
[68] Cosmetus varius Last Modified in 10/06/22 | 33 | 82 | 82 | 0 | 15 | 0 | 0 | 1 |
[69] Cosmetus delicatus Last Modified in 10/06/22 | 7 | 110 | 110 | 0 | 13 | 0 | 0 | 1 |
[70] Cosmetus arietinus Last Modified in 10/06/22 | 9 | 104 | 104 | 0 | 17 | 0 | 0 | 1 |
[71] Metavononoides barbacenensis Last Modified in 10/06/22 | 3 | 116 | 116 | 0 | 11 | 0 | 0 | 1 |
[72] Metavononoides guttulosus Last Modified in 10/06/22 | 3 | 116 | 116 | 0 | 11 | 0 | 0 | 1 |
[73] Metavononoides orientalis Last Modified in 10/06/22 | 34 | 85 | 85 | 0 | 11 | 0 | 0 | 1 |
[74] Paecilaema albisectum Last Modified in 10/06/22 | 2 | 116 | 116 | 0 | 12 | 0 | 0 | 1 |
[75] Paecilaema u-flavum Last Modified in 10/06/22 | 1 | 117 | 117 | 0 | 12 | 0 | 0 | 1 |
[76] Paecilaema melanacanthum Last Modified in 10/06/22 | 6 | 112 | 112 | 0 | 12 | 0 | 0 | 1 |
[77] Paecilaema lyra Last Modified in 10/06/22 | 38 | 82 | 82 | 0 | 10 | 0 | 0 | 1 |
[78] Paecilaema muticum Last Modified in 10/06/22 | 36 | 78 | 78 | 0 | 16 | 0 | 0 | 1 |
[79] Paecilaema ornatissimum Last Modified in 10/06/22 | 32 | 88 | 88 | 0 | 10 | 0 | 0 | 1 |
[80] Paecilaema preciosum Last Modified in 10/06/22 | 33 | 87 | 87 | 0 | 10 | 0 | 0 | 1 |
[81] Paecilaema renneri Last Modified in 10/06/22 | 6 | 112 | 112 | 0 | 12 | 0 | 0 | 1 |
[82] Kayania compressicoxa Last Modified in 10/06/22 | 43 | 76 | 76 | 0 | 11 | 0 | 0 | 1 |
[83] Kayania bicolumnata Last Modified in 10/06/22 | 37 | 82 | 82 | 0 | 11 | 0 | 0 | 1 |
[84] Marronia elegans Last Modified in 10/06/22 | 39 | 77 | 77 | 0 | 14 | 0 | 0 | 1 |
[85] Sibambea waorani sp. nov. Last Modified in 10/06/22 | 5 | 105 | 105 | 0 | 20 | 0 | 0 | 1 |
[86] Sibambea rotunda Last Modified in 10/06/22 | 2 | 109 | 109 | 0 | 19 | 0 | 0 | 1 |
[87] Discosomaticus cinctus Last Modified in 10/06/22 | 4 | 111 | 111 | 0 | 15 | 0 | 0 | 1 |
[88] Protus distinctus Last Modified in 10/06/22 | 10 | 105 | 105 | 0 | 15 | 0 | 0 | 1 |
[89] Protus bolivari Last Modified in 10/06/22 | 7 | 108 | 108 | 0 | 15 | 0 | 0 | 1 |
[90] Protus insolens Last Modified in 10/06/22 | 6 | 106 | 106 | 0 | 18 | 0 | 0 | 1 |
[91] Paraprotus speciosus Last Modified in 10/06/22 | 4 | 109 | 109 | 0 | 17 | 0 | 0 | 1 |
[92] Protus marllusi sp. nov. Last Modified in 10/06/22 | 8 | 105 | 105 | 0 | 17 | 0 | 0 | 1 |
[93] Paraprotus quadripunctatus Last Modified in 10/06/22 | 37 | 79 | 78 | 0 | 15 | 0 | 0 | 1 |
[94] Paraprotus atroluteus Last Modified in 10/06/22 | 37 | 77 | 77 | 0 | 16 | 0 | 0 | 1 |
[95] Paraprotus albithorax Last Modified in 10/06/22 | 37 | 79 | 78 | 0 | 15 | 0 | 0 | 1 |
[96] Gryne marginalis Last Modified in 10/06/22 | 5 | 113 | 112 | 0 | 13 | 0 | 0 | 1 |
[97] Gryne leprosa Last Modified in 10/06/22 | 5 | 111 | 111 | 0 | 14 | 0 | 0 | 1 |
[98] Gryne coccinelloides Last Modified in 10/06/22 | 39 | 79 | 79 | 0 | 12 | 0 | 0 | 1 |
[99] Gryne orensis Last Modified in 10/06/22 | 3 | 115 | 113 | 0 | 14 | 0 | 0 | 1 |
[100] Gryne dimorpha Last Modified in 10/06/22 | 6 | 111 | 110 | 0 | 14 | 0 | 0 | 1 |
[101] Roquettea andina Last Modified in 10/06/22 | 43 | 74 | 74 | 0 | 13 | 0 | 0 | 1 |
[102] Roquettea singularis Last Modified in 10/06/22 | 2 | 111 | 111 | 0 | 17 | 0 | 0 | 1 |
[103] Roquettea jalapensis Last Modified in 10/06/22 | 4 | 109 | 109 | 0 | 17 | 0 | 0 | 1 |
[104] Roquettea carajas Last Modified in 10/06/22 | 4 | 109 | 109 | 0 | 17 | 0 | 0 | 1 |
[105] Roquettea peba Last Modified in 10/06/22 | 4 | 110 | 110 | 0 | 16 | 0 | 0 | 1 |
[106] Roquettea scrotalis Last Modified in 10/06/22 | 6 | 105 | 105 | 0 | 19 | 0 | 0 | 1 |
[107] Roquettea taurina Last Modified in 10/06/22 | 4 | 110 | 110 | 0 | 16 | 0 | 0 | 1 |
[108] Roquettea bubalina Last Modified in 10/06/22 | 2 | 112 | 112 | 0 | 16 | 0 | 0 | 1 |
[109] Roquettea troguloides Last Modified in 10/06/22 | 2 | 112 | 112 | 0 | 16 | 0 | 0 | 1 |
[110] Roquettea decioi Last Modified in 10/06/22 | 33 | 83 | 83 | 0 | 14 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 5742 | |
Matrix views | 299 | Matrix landing page (274 views); Main - partition 2021 subfamilies (25 views); |
Media views | 984 | Media search (836 views); M847687 (148 views); |
Project overview | 356 | |
Specimen list | 487 | |
Taxon list | 3173 | |
Bibliography | 190 | |
Views for media list | 246 | |
Documents list | 7 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 8 | |
Matrix downloads | 7 | Main - partition 2021 subfamilies (7 downloads); |
Project downloads | 1 |