Project 4438: K. D. Angielczyk, J. Liu, W. Yang. 2021. A redescription of Kunpania scopulusa, a bidentalian dicynodont (Therapsida, Anomodontia) from the Guadalupian of northwestern China. Journal of Vertebrate Paleontology. 41 (1):null.
Abstract
Permian dicynodonts were discovered in strata exposed on the flanks of the Bogda Mountains (Xinjiang, China) in 1928. Nearly all known specimens were collected in the Guodikeng Formation (= upper Wutonggou low-order cycle); the single exception is the holotype of Kunpania scopulusa, which originated in the underlying Quanzijie Formation. The Quanzijie Formation is generally considered Capitanian in age, which would make Kunpania one of the oldest known bidentalian dicynodonts, but the type of K. scopulusa has received little attention. Here we redescribe K. scopulusa and investigate its phylogenetic relationships. Kunpania scopulusa is a valid species diagnosed by the absence of anterior palatal ridges; a robust crista oesophagea; a prominent lateral dentary shelf; a posteriorly curving projection on the angular; and a large, triangular attachment area for M. latissimus dorsi on the humerus. Our phylogenetic analysis recovered K. scopulusa as a stemward member of Dicynodontoidea and the sister taxon of the South African species Sintocephalus alticeps. Depending on the precise age of the uppermost Quanzijie Formation, three scenarios describe the potential phylogenetic implications of K. scopulusa: (1) bidentalian dicynodonts diversified before the end-Guadalupian extinction, with little of their early history preserved; (2) the main diversifications of cryptodont and dicynodontoid bidentalians occurred in the aftermath of the end-Guadalupian extinction, with K. scopulusa representing the latter; (3) K. scopulusa is a contemporary of other mid–late Wuchiapingian dicynodontoids and the initial radiation of Dicynodontoidea is not preserved in the known fossil record.Read the article »
Article DOI: 10.1080/02724634.2021.1922428
Project DOI: 10.7934/P4438, http://dx.doi.org/10.7934/P4438
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 13807 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 174 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 418 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:174/0 |
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MorphoBank Project 4438
MorphoBank Project 4438
- Creation Date:
09 October 2022 - Publication Date:
24 August 2023 - Project views: 3431
- Matrix downloads: 1
This research
supported by
Authors' Institutions
- Institute of Vertebrate Paleontology and Paleoanthropology (IVPP), CAS
- Field Museum of Natural History
- Missouri University of Science and Technology
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Bidane Martinez Huerta Project Administrator | 114 | 1 | 1 | 174 | 0 | 0 | 13807 (13807, 0, 0) | 0 | 0 | 0 | ||||
Brooke Long-Fox Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29024' (114 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Biarmosuchus tener Last Modified in 10/15/22 | 22 | 152 | 152 | 0 | 0 | 0 | 0 | 2 |
[2] † Hipposaurus boonstrai Last Modified in 08/24/23 | 24 | 150 | 150 | 0 | 0 | 0 | 0 | 2 |
[3] † Archaeosyodon praeventor Last Modified in 10/15/22 | 67 | 107 | 107 | 0 | 0 | 0 | 0 | 2 |
[4] † Titanophoneus potens Last Modified in 08/24/23 | 15 | 160 | 159 | 0 | 0 | 0 | 0 | 2 |
[5] † Gorgonops torvus Last Modified in 08/24/23 | 12 | 162 | 162 | 0 | 0 | 0 | 0 | 2 |
[6] † Lycosuchus vanderrieti Last Modified in 08/24/23 | 20 | 154 | 154 | 0 | 0 | 0 | 0 | 2 |
[7] † Glanosuchus macrops Last Modified in 08/24/23 | 12 | 162 | 162 | 0 | 0 | 0 | 0 | 2 |
[8] † Biseridens qilianicus Last Modified in 10/15/22 | 59 | 115 | 115 | 0 | 0 | 0 | 0 | 2 |
[9] † Anomocephalus africanus Last Modified in 10/15/22 | 146 | 28 | 28 | 0 | 0 | 0 | 0 | 2 |
[10] † Tiarajudens eccentricus Last Modified in 08/24/23 | 120 | 54 | 54 | 0 | 0 | 0 | 0 | 2 |
[11] † Otsheria netzvetajevi Last Modified in 08/24/23 | 88 | 86 | 86 | 0 | 0 | 0 | 0 | 2 |
[12] † Ulemica Last Modified in 08/24/23 | 48 | 127 | 126 | 0 | 0 | 0 | 0 | 2 |
[13] † Suminia getmanovi Last Modified in 08/24/23 | 20 | 154 | 154 | 0 | 0 | 0 | 0 | 2 |
[14] † Patranomodon nyaphulii Last Modified in 08/24/23 | 44 | 130 | 130 | 0 | 0 | 0 | 0 | 2 |
[15] † Galeops whaitsi Last Modified in 10/15/22 | 56 | 118 | 118 | 0 | 0 | 0 | 0 | 2 |
[16] † Galepus jouberti Last Modified in 10/15/22 | 134 | 40 | 40 | 0 | 0 | 0 | 0 | 2 |
[17] † Galechirus scholtzi Last Modified in 10/15/22 | 117 | 57 | 57 | 0 | 0 | 0 | 0 | 2 |
[18] † Eodicynodon oelofseni Last Modified in 10/15/22 | 80 | 94 | 94 | 0 | 0 | 0 | 0 | 2 |
[19] † Eodicynodon oosthuizeni Last Modified in 10/15/22 | 7 | 167 | 167 | 0 | 0 | 0 | 0 | 2 |
[20] † Colobodectes cluveri Last Modified in 10/15/22 | 71 | 103 | 103 | 0 | 0 | 0 | 0 | 2 |
[21] † Lanthanostegus mohoii Last Modified in 08/24/23 | 115 | 59 | 59 | 0 | 0 | 0 | 0 | 2 |
[22] † Eosimops newtoni Last Modified in 10/15/22 | 17 | 159 | 157 | 0 | 0 | 0 | 0 | 2 |
[23] † Diictodon feliceps Last Modified in 10/15/22 | 9 | 168 | 165 | 0 | 0 | 0 | 0 | 2 |
[24] † Robertia broomiana Last Modified in 08/24/23 | 13 | 161 | 161 | 0 | 0 | 0 | 0 | 2 |
[25] † Prosictodon dubei Last Modified in 08/24/23 | 49 | 125 | 125 | 0 | 0 | 0 | 0 | 2 |
[26] † Endothiodon bathystoma Last Modified in 10/15/22 | 15 | 162 | 159 | 0 | 0 | 0 | 0 | 2 |
[27] † Endothiodon tolani Last Modified in 10/15/22 | 48 | 127 | 126 | 0 | 0 | 0 | 0 | 2 |
[28] † Niassodon mfumukasi Last Modified in 08/24/23 | 44 | 130 | 130 | 0 | 0 | 0 | 0 | 2 |
[29] † Brachyprosopus broomi Last Modified in 10/15/22 | 22 | 152 | 152 | 0 | 0 | 0 | 0 | 2 |
[30] † Pristerodon mackayi Last Modified in 08/24/23 | 8 | 166 | 166 | 0 | 0 | 0 | 0 | 2 |
[31] † Emydops Last Modified in 10/16/22 | 9 | 166 | 165 | 0 | 0 | 0 | 0 | 2 |
[32] † Digalodon rubidgei Last Modified in 10/15/22 | 54 | 120 | 120 | 0 | 0 | 0 | 0 | 2 |
[33] † Dicynodontoides Last Modified in 10/16/22 | 9 | 165 | 165 | 0 | 0 | 0 | 0 | 2 |
[34] † Kombuisia frerensis Last Modified in 08/24/23 | 55 | 119 | 119 | 0 | 0 | 0 | 0 | 2 |
[35] † Myosaurus gracilis Last Modified in 08/24/23 | 28 | 146 | 146 | 0 | 0 | 0 | 0 | 2 |
[36] † Cistecephalus microrhinus Last Modified in 10/15/22 | 12 | 164 | 162 | 0 | 0 | 0 | 0 | 2 |
[37] † Cistecephaloides boonstrai Last Modified in 10/15/22 | 52 | 122 | 122 | 0 | 0 | 0 | 0 | 2 |
[38] † Kawingasaurus fossilis Last Modified in 08/24/23 | 35 | 139 | 139 | 0 | 0 | 0 | 0 | 2 |
[39] † Daqingshanodon limbus Last Modified in 10/15/22 | 52 | 122 | 122 | 0 | 0 | 0 | 0 | 2 |
[40] † Keyseria benjamini Last Modified in 08/24/23 | 75 | 99 | 99 | 0 | 0 | 0 | 0 | 2 |
[41] † Rhachiocephalus magnus Last Modified in 08/24/23 | 19 | 156 | 155 | 0 | 0 | 0 | 0 | 2 |
[42] † Kitchinganomodon crassus Last Modified in 08/24/23 | 40 | 134 | 134 | 0 | 0 | 0 | 0 | 2 |
[43] † Oudenodon bainii Last Modified in 08/24/23 | 7 | 168 | 167 | 0 | 0 | 0 | 0 | 2 |
[44] † Tropidostoma dubium Last Modified in 08/24/23 | 18 | 156 | 156 | 0 | 0 | 0 | 0 | 2 |
[45] † Australobarbarus Last Modified in 10/16/22 | 42 | 132 | 132 | 0 | 0 | 0 | 0 | 2 |
[46] † Syops vanhoepeni Last Modified in 08/24/23 | 87 | 87 | 87 | 0 | 0 | 0 | 0 | 2 |
[47] † Odontocyclops whaitsi Last Modified in 08/24/23 | 22 | 152 | 152 | 0 | 0 | 0 | 0 | 2 |
[48] † Idelesaurus tataricus Last Modified in 08/24/23 | 42 | 132 | 132 | 0 | 0 | 0 | 0 | 2 |
[49] † Aulacephalodon bainii Last Modified in 10/15/22 | 7 | 172 | 167 | 0 | 0 | 0 | 0 | 2 |
[50] † Pelanomodon moschops Last Modified in 08/24/23 | 64 | 111 | 110 | 0 | 0 | 0 | 0 | 2 |
[51] † Geikia locusticeps Last Modified in 10/15/22 | 36 | 138 | 138 | 0 | 0 | 0 | 0 | 2 |
[52] † Geikia elginensis Last Modified in 08/24/23 | 78 | 96 | 96 | 0 | 0 | 0 | 0 | 2 |
[53] † Elph borealis Last Modified in 10/15/22 | 60 | 114 | 114 | 0 | 0 | 0 | 0 | 2 |
[54] † Interpresosaurus blomi Last Modified in 08/24/23 | 130 | 44 | 44 | 0 | 0 | 0 | 0 | 2 |
[55] † Katumbia parringtoni Last Modified in 08/24/23 | 54 | 120 | 120 | 0 | 0 | 0 | 0 | 2 |
[56] † Delectosaurus arefjevi Last Modified in 10/15/22 | 67 | 108 | 107 | 0 | 0 | 0 | 0 | 2 |
[57] † Dicynodon lacerticeps Last Modified in 10/15/22 | 37 | 138 | 137 | 0 | 0 | 0 | 0 | 2 |
[58] † Daptocephalus leoniceps Last Modified in 10/15/22 | 27 | 147 | 147 | 0 | 0 | 0 | 0 | 2 |
[59] † Dinanomodon gilli Last Modified in 10/15/22 | 51 | 123 | 123 | 0 | 0 | 0 | 0 | 2 |
[60] † Peramodon amalitzkii Last Modified in 08/24/23 | 78 | 96 | 96 | 0 | 0 | 0 | 0 | 2 |
[61] † Vivaxosaurus trautscholdi Last Modified in 08/24/23 | 33 | 143 | 141 | 0 | 0 | 0 | 0 | 2 |
[62] † Jimusaria sinkiangensis Last Modified in 08/24/23 | 54 | 120 | 120 | 0 | 0 | 0 | 0 | 2 |
[63] † Sintocephalus alticeps Last Modified in 08/24/23 | 94 | 80 | 80 | 0 | 0 | 0 | 0 | 2 |
[64] † Basilodon woodwardi Last Modified in 10/15/22 | 63 | 111 | 111 | 0 | 0 | 0 | 0 | 2 |
[65] † Turfanodon bogdaensis Last Modified in 08/24/23 | 71 | 103 | 103 | 0 | 0 | 0 | 0 | 2 |
[66] † Gordonia traquairi Last Modified in 08/24/23 | 71 | 103 | 103 | 0 | 0 | 0 | 0 | 2 |
[67] † Euptychognathus bathyrhynchus Last Modified in 10/15/22 | 61 | 113 | 113 | 0 | 0 | 0 | 0 | 2 |
[68] † Lystrosaurus hedini Last Modified in 08/24/23 | 19 | 155 | 155 | 0 | 0 | 0 | 0 | 2 |
[69] † Lystrosaurus maccaigi Last Modified in 08/24/23 | 35 | 139 | 139 | 0 | 0 | 0 | 0 | 2 |
[70] † Lystrosaurus curvatus Last Modified in 08/24/23 | 34 | 140 | 140 | 0 | 0 | 0 | 0 | 2 |
[71] † Lystrosaurus declivis Last Modified in 08/24/23 | 14 | 160 | 160 | 0 | 0 | 0 | 0 | 2 |
[72] † Lystrosaurus murrayi Last Modified in 08/24/23 | 14 | 160 | 160 | 0 | 0 | 0 | 0 | 2 |
[73] † Shansiodon Last Modified in 08/24/23 | 33 | 141 | 141 | 0 | 0 | 0 | 0 | 2 |
[74] † Tetragonias njalilus Last Modified in 08/24/23 | 10 | 165 | 164 | 0 | 0 | 0 | 0 | 2 |
[75] † Vinceria andina Last Modified in 08/24/23 | 78 | 96 | 96 | 0 | 0 | 0 | 0 | 2 |
[76] † Rhinodicynodon gracile Last Modified in 08/24/23 | 41 | 133 | 133 | 0 | 0 | 0 | 0 | 2 |
[77] † Dinodontosaurus Last Modified in 10/16/22 | 17 | 157 | 157 | 0 | 0 | 0 | 0 | 2 |
[78] † Angonisaurus cruickshanki Last Modified in 10/15/22 | 45 | 129 | 129 | 0 | 0 | 0 | 0 | 2 |
[79] † Kannemeyeria simocephalus Last Modified in 08/24/23 | 12 | 162 | 162 | 0 | 0 | 0 | 0 | 2 |
[80] † Kannemeyeria lophorhinus Last Modified in 08/24/23 | 46 | 128 | 128 | 0 | 0 | 0 | 0 | 2 |
[81] † Dolichuranus primaevus Last Modified in 10/15/22 | 32 | 142 | 142 | 0 | 0 | 0 | 0 | 2 |
[82] † Sinokannemeyeria Last Modified in 08/24/23 | 26 | 148 | 148 | 0 | 0 | 0 | 0 | 2 |
[83] † Parakannemeyeria Last Modified in 08/24/23 | 24 | 151 | 150 | 0 | 0 | 0 | 0 | 2 |
[84] † Xiyukannemeyeria brevirostris Last Modified in 08/24/23 | 46 | 128 | 128 | 0 | 0 | 0 | 0 | 2 |
[85] † Rabidosaurus cristatus Last Modified in 08/24/23 | 102 | 72 | 72 | 0 | 0 | 0 | 0 | 2 |
[86] † Rhadiodromus Last Modified in 08/24/23 | 81 | 94 | 93 | 0 | 0 | 0 | 0 | 2 |
[87] † Wadiasaurus indicus Last Modified in 08/24/23 | 64 | 111 | 110 | 0 | 0 | 0 | 0 | 2 |
[88] † Shaanbeikannemeyeria Last Modified in 08/24/23 | 54 | 120 | 120 | 0 | 0 | 0 | 0 | 2 |
[89] † Rechnisaurus cristarhynchus Last Modified in 08/24/23 | 100 | 74 | 74 | 0 | 0 | 0 | 0 | 2 |
[90] † Uralokannemeyeria vjuschkovi Last Modified in 08/24/23 | 96 | 78 | 78 | 0 | 0 | 0 | 0 | 2 |
[91] † Zambiasaurus submersus Last Modified in 08/24/23 | 111 | 63 | 63 | 0 | 0 | 0 | 0 | 2 |
[92] † Placerias hesternus Last Modified in 08/24/23 | 51 | 123 | 123 | 0 | 0 | 0 | 0 | 2 |
[93] † Moghreberia nmachouensis Last Modified in 08/24/23 | 76 | 98 | 98 | 0 | 0 | 0 | 0 | 2 |
[94] † Stahleckeria potens Last Modified in 08/24/23 | 11 | 163 | 163 | 0 | 0 | 0 | 0 | 2 |
[95] † Sangusaurus parringtonii Last Modified in 08/24/23 | 40 | 134 | 134 | 0 | 0 | 0 | 0 | 2 |
[96] † Eubrachiosaurus browni Last Modified in 10/15/22 | 163 | 11 | 11 | 0 | 0 | 0 | 0 | 2 |
[97] † Jachaleria Last Modified in 08/24/23 | 44 | 130 | 130 | 0 | 0 | 0 | 0 | 2 |
[98] † Ischigualastia jenseni Last Modified in 08/24/23 | 22 | 152 | 152 | 0 | 0 | 0 | 0 | 2 |
[99] † Rastodon procurvidens Last Modified in 08/24/23 | 72 | 102 | 102 | 0 | 0 | 0 | 0 | 2 |
[100] † Sauroscaptor tharavati Last Modified in 08/24/23 | 80 | 94 | 94 | 0 | 0 | 0 | 0 | 2 |
[101] † Bulbasaurus phylloxyron Last Modified in 10/15/22 | 52 | 126 | 122 | 0 | 0 | 0 | 0 | 2 |
[102] † Compsodon helmoedi Last Modified in 10/15/22 | 64 | 110 | 110 | 0 | 0 | 0 | 0 | 2 |
[103] † Pentasaurus goggai Last Modified in 08/24/23 | 150 | 24 | 24 | 0 | 0 | 0 | 0 | 2 |
[104] † Thliptosaurus imperforatus Last Modified in 08/24/23 | 106 | 68 | 68 | 0 | 0 | 0 | 0 | 2 |
[105] † Ufudocyclops mukanelai Last Modified in 08/24/23 | 63 | 111 | 111 | 0 | 0 | 0 | 0 | 2 |
[106] † Dicynodon angielczyki Last Modified in 10/15/22 | 39 | 135 | 135 | 0 | 0 | 0 | 0 | 2 |
[107] † Daptocephalus huenei Last Modified in 10/15/22 | 65 | 109 | 109 | 0 | 0 | 0 | 0 | 2 |
[108] † Kunpania scopulusa Last Modified in 08/24/23 | 93 | 81 | 81 | 0 | 0 | 0 | 0 | 2 |
[109] † Taoheodon baizhijuni Last Modified in 08/24/23 | 67 | 107 | 107 | 0 | 0 | 0 | 0 | 2 |
[110] LPB 1993 3" Last Modified in 10/15/22 | 96 | 78 | 78 | 0 | 0 | 0 | 0 | 2 |
[111] LPB 1993 2" Last Modified in 10/15/22 | 99 | 75 | 75 | 0 | 0 | 0 | 0 | 2 |
[112] LPB 1995 9" Last Modified in 10/15/22 | 91 | 83 | 83 | 0 | 0 | 0 | 0 | 2 |
[113] † Kembawacela kitchingi Last Modified in 08/24/23 | 38 | 136 | 136 | 0 | 0 | 0 | 0 | 2 |
[114] † Abajudon kaayai Last Modified in 10/15/22 | 51 | 123 | 123 | 0 | 0 | 0 | 0 | 2 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 3431 | |
Media views | 574 | Media search (494 views); M847929 (80 views); |
Project overview | 221 | |
Matrix views | 136 | Matrix landing page (120 views); Angielczyk et al. (2021) (16 views); |
Documents list | 123 | |
Specimen list | 629 | |
Taxon list | 1504 | |
Bibliography | 96 | |
Views for media list | 148 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 1 | |
Matrix downloads | 1 | Angielczyk et al. (2021) (1 download); |