Project 4532: N. Bekkouche, L. Gąsiorowski. 2022. Careful amendment of morphological data sets improves phylogenetic frameworks: re-evaluating placement of the fossil Amiskwia sagittiformis. Journal of Systematic Palaeontology. 20 (1):1-14.
Abstract
The Cambrian fossil Amiskwia sagittiformis has puzzled palaeontologists for more than a century, but recent re-investigation of its morphology suggested a close relationship with the bilaterian clade Gnathifera, comprising Rotifera, Gnathostomulida and Micrognathozoa. Since Amiskwia has already been considered closely related to Chaetognatha, this new interpretation of its morphology supports recent molecular and developmental studies finding a close phylogenetic relationship between Chaetognatha and Gnathifera. The recent re-description of the jaw apparatus of Amiskwia with the first phylogenetic analysis by Vinther & Parry (2019) found a rather surprising topology with Amiskwia and Chaetognatha being sister groups and nested inside Gnathifera. Furthermore, a subsequent paper from Caron & Cheung (2019) re-described the jaws of Amiskwia as very similar to the configuration found in Gnathostomulida, but did not include any phylogenetic analysis. Here we test the topology of Vinther & Parry with various parsimony and Bayesian analyses, taking into account the new description of Caron & Cheung with careful re-amendment of the matrix of Vinther & Parry. According to our results and the recent findings on the molecular phylogeny of their extant members, we suggest a new systematization of these taxa. We recovered Amiskwia as a stem-group chaetognath within a clade Cucullophora nov., rejecting the Caron & Cheung hypothesis of its close affinity to Gnathostomulida, and found a more consensual topology with monophyletic Gnathifera sister group to Cucullophora, all together forming the clade Chaetognathifera. Furthermore, Rotifera + Micrognathozoa form a clade named Gynognathifera nov. We discuss the characters supporting each clade and the reasons that account for the topology found by Vinther & Parry. Finally, we hope that this carefully amended matrix focused on Gnathifera, Amiskwia and Chaetognatha, together with a formal classification and robust phylogeny, will be of use for future studies on the palaeontology and morphology of these clades.Read the article »
Article DOI: 10.1080/14772019.2022.2109217
Project DOI: 10.7934/P4532, http://dx.doi.org/10.7934/P4532
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 14928 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 570 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 295 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:570/0 |
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MorphoBank Project 4532
MorphoBank Project 4532
- Creation Date:
20 January 2023 - Publication Date:
20 January 2023 - Project views: 3762
- Matrix downloads: 6
Authors' Institutions
- Sorbonne Université
- Max Planck Institute for Multidisciplinary Sciences
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 33 | 1 | 1 | 570 | 0 | 0 | 14928 (12598, 0, 2330) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29114' (33 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Bryozoa Last Modified in 01/20/23 | 2 | 57 | 57 | 0 | 15 | 0 | 0 | 1 |
[2] Brachiopoda Last Modified in 01/20/23 | 1 | 61 | 60 | 0 | 13 | 0 | 0 | 1 |
[3] Phoronida Last Modified in 01/20/23 | 5 | 58 | 58 | 0 | 11 | 0 | 0 | 1 |
[4] Annelida Last Modified in 01/20/23 | 0 | 64 | 64 | 0 | 10 | 0 | 0 | 1 |
[5] Sipuncula Last Modified in 01/20/23 | 9 | 53 | 53 | 0 | 12 | 0 | 0 | 1 |
[6] Mollusca Last Modified in 01/20/23 | 1 | 74 | 72 | 0 | 1 | 0 | 0 | 1 |
[7] † Wiwaxia Last Modified in 01/20/23 | 46 | 25 | 25 | 0 | 3 | 0 | 0 | 1 |
[8] † Odontogriphus Last Modified in 01/20/23 | 45 | 26 | 26 | 0 | 3 | 0 | 0 | 1 |
[9] † Canadia Last Modified in 01/20/23 | 36 | 28 | 28 | 0 | 10 | 0 | 0 | 1 |
[10] Entoprocta Last Modified in 01/20/23 | 9 | 52 | 52 | 0 | 13 | 0 | 0 | 1 |
[11] Platyhelminthes Last Modified in 01/20/23 | 0 | 58 | 58 | 0 | 16 | 0 | 0 | 1 |
[12] Gastrotricha Last Modified in 01/20/23 | 9 | 50 | 50 | 0 | 15 | 0 | 0 | 1 |
[13] Gnathostumulida Last Modified in 01/20/23 | 9 | 58 | 57 | 0 | 8 | 0 | 0 | 1 |
[14] Micrognathozoa Last Modified in 01/20/23 | 13 | 54 | 54 | 0 | 7 | 0 | 0 | 1 |
[15] † Inquicus fellatus Last Modified in 01/20/23 | 50 | 23 | 23 | 0 | 1 | 0 | 0 | 1 |
[16] † Amiskwia Last Modified in 01/20/23 | 48 | 26 | 26 | 0 | 0 | 0 | 0 | 1 |
[17] Flacisaggita Last Modified in 01/20/23 | 0 | 68 | 68 | 0 | 6 | 0 | 0 | 1 |
[18] Spadella Last Modified in 01/20/23 | 0 | 68 | 68 | 0 | 6 | 0 | 0 | 1 |
[19] Eukrohnia Last Modified in 01/20/23 | 6 | 62 | 62 | 0 | 6 | 0 | 0 | 1 |
[20] † Protosagitta Last Modified in 01/20/23 | 57 | 16 | 16 | 0 | 1 | 0 | 0 | 1 |
[21] † Ankalodous Last Modified in 01/20/23 | 63 | 10 | 10 | 0 | 1 | 0 | 0 | 1 |
[22] † Capinatator Last Modified in 01/20/23 | 56 | 17 | 17 | 0 | 1 | 0 | 0 | 1 |
[23] Adineta Last Modified in 01/20/23 | 3 | 65 | 65 | 0 | 6 | 0 | 0 | 1 |
[24] † Branchionus Last Modified in 01/20/23 | 3 | 65 | 65 | 0 | 6 | 0 | 0 | 1 |
[25] Nemertea Last Modified in 01/20/23 | 1 | 61 | 61 | 0 | 12 | 0 | 0 | 1 |
[26] Onychophora Last Modified in 01/20/23 | 0 | 63 | 63 | 0 | 11 | 0 | 0 | 1 |
[27] Arthropoda Last Modified in 01/20/23 | 0 | 62 | 62 | 0 | 12 | 0 | 0 | 1 |
[28] Tardigrada Last Modified in 01/20/23 | 8 | 52 | 52 | 0 | 14 | 0 | 0 | 1 |
[29] Nematoda Last Modified in 01/20/23 | 7 | 51 | 51 | 0 | 16 | 0 | 0 | 1 |
[30] Nematomorpha Last Modified in 01/20/23 | 7 | 51 | 51 | 0 | 16 | 0 | 0 | 1 |
[31] Priapulida Last Modified in 01/20/23 | 0 | 62 | 62 | 0 | 12 | 0 | 0 | 1 |
[32] Kinorhyncha Last Modified in 01/20/23 | 10 | 52 | 52 | 0 | 12 | 0 | 0 | 1 |
[33] Loricifera Last Modified in 01/20/23 | 9 | 53 | 53 | 0 | 12 | 0 | 0 | 1 |
Taxonomic Overview for Matrix 'M29115' (33 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Bryozoa Last Modified in 01/20/23 | 2 | 56 | 56 | 0 | 14 | 0 | 0 | 1 |
[2] Brachiopoda Last Modified in 01/20/23 | 1 | 60 | 59 | 0 | 12 | 0 | 0 | 1 |
[3] Phoronida Last Modified in 01/20/23 | 5 | 57 | 57 | 0 | 10 | 0 | 0 | 1 |
[4] Annelida Last Modified in 01/20/23 | 0 | 63 | 63 | 0 | 9 | 0 | 0 | 1 |
[5] Sipuncula Last Modified in 01/20/23 | 9 | 52 | 52 | 0 | 11 | 0 | 0 | 1 |
[6] Mollusca Last Modified in 01/20/23 | 1 | 73 | 71 | 0 | 0 | 0 | 0 | 1 |
[7] † Wiwaxia Last Modified in 01/20/23 | 46 | 24 | 24 | 0 | 2 | 0 | 0 | 1 |
[8] † Odontogriphus Last Modified in 01/20/23 | 45 | 25 | 25 | 0 | 2 | 0 | 0 | 1 |
[9] † Canadia Last Modified in 01/20/23 | 36 | 27 | 27 | 0 | 9 | 0 | 0 | 1 |
[10] Entoprocta Last Modified in 01/20/23 | 9 | 51 | 51 | 0 | 12 | 0 | 0 | 1 |
[11] Platyhelminthes Last Modified in 01/20/23 | 0 | 57 | 57 | 0 | 15 | 0 | 0 | 1 |
[12] Gastrotricha Last Modified in 01/20/23 | 9 | 49 | 49 | 0 | 14 | 0 | 0 | 1 |
[13] Gnathostumulida Last Modified in 01/20/23 | 9 | 57 | 56 | 0 | 7 | 0 | 0 | 1 |
[14] Micrognathozoa Last Modified in 01/20/23 | 13 | 53 | 53 | 0 | 6 | 0 | 0 | 1 |
[15] † Inquicus fellatus Last Modified in 01/20/23 | 50 | 22 | 22 | 0 | 0 | 0 | 0 | 1 |
[16] † Amiskwia Last Modified in 01/20/23 | 48 | 24 | 24 | 0 | 0 | 0 | 0 | 1 |
[17] Flacisaggita Last Modified in 01/20/23 | 0 | 66 | 66 | 0 | 6 | 0 | 0 | 1 |
[18] Spadella Last Modified in 01/20/23 | 0 | 66 | 66 | 0 | 6 | 0 | 0 | 1 |
[19] Eukrohnia Last Modified in 01/20/23 | 6 | 60 | 60 | 0 | 6 | 0 | 0 | 1 |
[20] † Protosagitta Last Modified in 01/20/23 | 55 | 16 | 16 | 0 | 1 | 0 | 0 | 1 |
[21] † Ankalodous Last Modified in 01/20/23 | 61 | 10 | 10 | 0 | 1 | 0 | 0 | 1 |
[22] † Capinatator Last Modified in 01/20/23 | 54 | 17 | 17 | 0 | 1 | 0 | 0 | 1 |
[23] Adineta Last Modified in 01/20/23 | 3 | 64 | 64 | 0 | 5 | 0 | 0 | 1 |
[24] † Branchionus Last Modified in 01/20/23 | 3 | 64 | 64 | 0 | 5 | 0 | 0 | 1 |
[25] Nemertea Last Modified in 01/20/23 | 1 | 60 | 60 | 0 | 11 | 0 | 0 | 1 |
[26] Onychophora Last Modified in 01/20/23 | 0 | 62 | 62 | 0 | 10 | 0 | 0 | 1 |
[27] Arthropoda Last Modified in 01/20/23 | 0 | 61 | 61 | 0 | 11 | 0 | 0 | 1 |
[28] Tardigrada Last Modified in 01/20/23 | 8 | 51 | 51 | 0 | 13 | 0 | 0 | 1 |
[29] Nematoda Last Modified in 01/20/23 | 7 | 50 | 50 | 0 | 15 | 0 | 0 | 1 |
[30] Nematomorpha Last Modified in 01/20/23 | 7 | 50 | 50 | 0 | 15 | 0 | 0 | 1 |
[31] Priapulida Last Modified in 01/20/23 | 0 | 61 | 61 | 0 | 11 | 0 | 0 | 1 |
[32] Kinorhyncha Last Modified in 01/20/23 | 10 | 51 | 51 | 0 | 11 | 0 | 0 | 1 |
[33] Loricifera Last Modified in 01/20/23 | 9 | 52 | 52 | 0 | 11 | 0 | 0 | 1 |
Taxonomic Overview for Matrix 'M29116' (32 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Bryozoa Last Modified in 01/20/23 | 2 | 57 | 57 | 0 | 15 | 0 | 0 | 1 |
[2] Brachiopoda Last Modified in 01/20/23 | 1 | 61 | 60 | 0 | 13 | 0 | 0 | 1 |
[3] Phoronida Last Modified in 01/20/23 | 5 | 58 | 58 | 0 | 11 | 0 | 0 | 1 |
[4] Annelida Last Modified in 01/20/23 | 0 | 64 | 64 | 0 | 10 | 0 | 0 | 1 |
[5] Sipuncula Last Modified in 01/20/23 | 9 | 53 | 53 | 0 | 12 | 0 | 0 | 1 |
[6] Mollusca Last Modified in 01/20/23 | 1 | 74 | 72 | 0 | 1 | 0 | 0 | 1 |
[7] † Wiwaxia Last Modified in 01/20/23 | 46 | 25 | 25 | 0 | 3 | 0 | 0 | 1 |
[8] † Odontogriphus Last Modified in 01/20/23 | 45 | 26 | 26 | 0 | 3 | 0 | 0 | 1 |
[9] † Canadia Last Modified in 01/20/23 | 36 | 28 | 28 | 0 | 10 | 0 | 0 | 1 |
[10] Entoprocta Last Modified in 01/20/23 | 9 | 52 | 52 | 0 | 13 | 0 | 0 | 1 |
[11] Platyhelminthes Last Modified in 01/20/23 | 0 | 58 | 58 | 0 | 16 | 0 | 0 | 1 |
[12] Gastrotricha Last Modified in 01/20/23 | 9 | 50 | 50 | 0 | 15 | 0 | 0 | 1 |
[13] Gnathostumulida Last Modified in 01/20/23 | 9 | 58 | 57 | 0 | 8 | 0 | 0 | 1 |
[14] Micrognathozoa Last Modified in 01/20/23 | 13 | 54 | 54 | 0 | 7 | 0 | 0 | 1 |
[15] † Inquicus fellatus Last Modified in 01/20/23 | 50 | 23 | 23 | 0 | 1 | 0 | 0 | 1 |
[16] Flacisaggita Last Modified in 01/20/23 | 0 | 68 | 68 | 0 | 6 | 0 | 0 | 1 |
[17] Spadella Last Modified in 01/20/23 | 0 | 68 | 68 | 0 | 6 | 0 | 0 | 1 |
[18] Eukrohnia Last Modified in 01/20/23 | 6 | 62 | 62 | 0 | 6 | 0 | 0 | 1 |
[19] † Protosagitta Last Modified in 01/20/23 | 57 | 16 | 16 | 0 | 1 | 0 | 0 | 1 |
[20] † Ankalodous Last Modified in 01/20/23 | 63 | 10 | 10 | 0 | 1 | 0 | 0 | 1 |
[21] † Capinatator Last Modified in 01/20/23 | 56 | 17 | 17 | 0 | 1 | 0 | 0 | 1 |
[22] Adineta Last Modified in 01/20/23 | 3 | 65 | 65 | 0 | 6 | 0 | 0 | 1 |
[23] † Branchionus Last Modified in 01/20/23 | 3 | 65 | 65 | 0 | 6 | 0 | 0 | 1 |
[24] Nemertea Last Modified in 01/20/23 | 1 | 61 | 61 | 0 | 12 | 0 | 0 | 1 |
[25] Onychophora Last Modified in 01/20/23 | 0 | 63 | 63 | 0 | 11 | 0 | 0 | 1 |
[26] Arthropoda Last Modified in 01/20/23 | 0 | 62 | 62 | 0 | 12 | 0 | 0 | 1 |
[27] Tardigrada Last Modified in 01/20/23 | 8 | 52 | 52 | 0 | 14 | 0 | 0 | 1 |
[28] Nematoda Last Modified in 01/20/23 | 7 | 51 | 51 | 0 | 16 | 0 | 0 | 1 |
[29] Nematomorpha Last Modified in 01/20/23 | 7 | 51 | 51 | 0 | 16 | 0 | 0 | 1 |
[30] Priapulida Last Modified in 01/20/23 | 0 | 62 | 62 | 0 | 12 | 0 | 0 | 1 |
[31] Kinorhyncha Last Modified in 01/20/23 | 10 | 52 | 52 | 0 | 12 | 0 | 0 | 1 |
[32] Loricifera Last Modified in 01/20/23 | 9 | 53 | 53 | 0 | 12 | 0 | 0 | 1 |
Taxonomic Overview for Matrix 'M29117' (33 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Bryozoa Last Modified in 01/20/23 | 2 | 110 | 110 | 0 | 26 | 0 | 0 | 1 |
[2] Brachiopoda Last Modified in 01/20/23 | 1 | 117 | 115 | 0 | 22 | 0 | 0 | 1 |
[3] Phoronida Last Modified in 01/20/23 | 5 | 110 | 110 | 0 | 23 | 0 | 0 | 1 |
[4] Annelida Last Modified in 01/20/23 | 0 | 120 | 120 | 0 | 18 | 0 | 0 | 1 |
[5] Sipuncula Last Modified in 01/20/23 | 9 | 106 | 106 | 0 | 23 | 0 | 0 | 1 |
[6] Mollusca Last Modified in 01/20/23 | 1 | 132 | 129 | 0 | 8 | 0 | 0 | 1 |
[7] † Wiwaxia Last Modified in 01/20/23 | 91 | 39 | 39 | 0 | 8 | 0 | 0 | 1 |
[8] † Odontogriphus Last Modified in 01/20/23 | 93 | 37 | 37 | 0 | 8 | 0 | 0 | 1 |
[9] † Canadia Last Modified in 01/20/23 | 80 | 43 | 43 | 0 | 15 | 0 | 0 | 1 |
[10] Entoprocta Last Modified in 01/20/23 | 10 | 104 | 104 | 0 | 24 | 0 | 0 | 1 |
[11] Platyhelminthes Last Modified in 01/20/23 | 1 | 109 | 109 | 0 | 28 | 0 | 0 | 1 |
[12] Gastrotricha Last Modified in 01/20/23 | 10 | 101 | 101 | 0 | 27 | 0 | 0 | 1 |
[13] Gnathostumulida Last Modified in 01/20/23 | 9 | 109 | 108 | 0 | 21 | 0 | 0 | 1 |
[14] Micrognathozoa Last Modified in 01/20/23 | 23 | 96 | 96 | 0 | 19 | 0 | 0 | 1 |
[15] † Inquicus fellatus Last Modified in 01/20/23 | 97 | 32 | 32 | 0 | 9 | 0 | 0 | 1 |
[16] † Amiskwia Last Modified in 01/20/23 | 93 | 37 | 37 | 0 | 8 | 0 | 0 | 1 |
[17] Flacisaggita Last Modified in 01/20/23 | 0 | 123 | 123 | 0 | 15 | 0 | 0 | 1 |
[18] Spadella Last Modified in 01/20/23 | 0 | 125 | 125 | 0 | 13 | 0 | 0 | 1 |
[19] Eukrohnia Last Modified in 01/20/23 | 6 | 119 | 119 | 0 | 13 | 0 | 0 | 1 |
[20] † Protosagitta Last Modified in 01/20/23 | 109 | 23 | 23 | 0 | 6 | 0 | 0 | 1 |
[21] † Ankalodous Last Modified in 01/20/23 | 122 | 14 | 14 | 0 | 2 | 0 | 0 | 1 |
[22] † Capinatator Last Modified in 01/20/23 | 107 | 24 | 24 | 0 | 7 | 0 | 0 | 1 |
[23] Adineta Last Modified in 01/20/23 | 3 | 116 | 116 | 0 | 19 | 0 | 0 | 1 |
[24] † Branchionus Last Modified in 01/20/23 | 3 | 116 | 116 | 0 | 19 | 0 | 0 | 1 |
[25] Nemertea Last Modified in 01/20/23 | 2 | 113 | 113 | 0 | 23 | 0 | 0 | 1 |
[26] Onychophora Last Modified in 01/20/23 | 0 | 117 | 117 | 0 | 21 | 0 | 0 | 1 |
[27] Arthropoda Last Modified in 01/20/23 | 0 | 117 | 116 | 0 | 22 | 0 | 0 | 1 |
[28] Tardigrada Last Modified in 01/20/23 | 8 | 106 | 106 | 0 | 24 | 0 | 0 | 1 |
[29] Nematoda Last Modified in 01/20/23 | 7 | 102 | 102 | 0 | 29 | 0 | 0 | 1 |
[30] Nematomorpha Last Modified in 01/20/23 | 7 | 102 | 102 | 0 | 29 | 0 | 0 | 1 |
[31] Priapulida Last Modified in 01/20/23 | 0 | 113 | 113 | 0 | 25 | 0 | 0 | 1 |
[32] Kinorhyncha Last Modified in 01/20/23 | 10 | 103 | 103 | 0 | 25 | 0 | 0 | 1 |
[33] Loricifera Last Modified in 01/20/23 | 9 | 104 | 104 | 0 | 25 | 0 | 0 | 1 |
Taxonomic Overview for Matrix 'M29118' (33 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Bryozoa Last Modified in 01/20/23 | 2 | 57 | 57 | 0 | 15 | 0 | 0 | 1 |
[2] Brachiopoda Last Modified in 01/20/23 | 1 | 61 | 60 | 0 | 13 | 0 | 0 | 1 |
[3] Phoronida Last Modified in 01/20/23 | 5 | 58 | 58 | 0 | 11 | 0 | 0 | 1 |
[4] Annelida Last Modified in 01/20/23 | 0 | 64 | 64 | 0 | 10 | 0 | 0 | 1 |
[5] Sipuncula Last Modified in 01/20/23 | 9 | 53 | 53 | 0 | 12 | 0 | 0 | 1 |
[6] Mollusca Last Modified in 01/20/23 | 1 | 74 | 72 | 0 | 1 | 0 | 0 | 1 |
[7] † Wiwaxia Last Modified in 01/20/23 | 46 | 25 | 25 | 0 | 3 | 0 | 0 | 1 |
[8] † Odontogriphus Last Modified in 01/20/23 | 45 | 26 | 26 | 0 | 3 | 0 | 0 | 1 |
[9] † Canadia Last Modified in 01/20/23 | 37 | 28 | 28 | 0 | 9 | 0 | 0 | 1 |
[10] Entoprocta Last Modified in 01/20/23 | 10 | 52 | 52 | 0 | 12 | 0 | 0 | 1 |
[11] Platyhelminthes Last Modified in 01/20/23 | 1 | 58 | 58 | 0 | 15 | 0 | 0 | 1 |
[12] Gastrotricha Last Modified in 01/20/23 | 10 | 50 | 50 | 0 | 14 | 0 | 0 | 1 |
[13] Gnathostumulida Last Modified in 01/20/23 | 9 | 58 | 57 | 0 | 8 | 0 | 0 | 1 |
[14] Micrognathozoa Last Modified in 01/20/23 | 13 | 54 | 54 | 0 | 7 | 0 | 0 | 1 |
[15] † Inquicus fellatus Last Modified in 01/20/23 | 50 | 23 | 23 | 0 | 1 | 0 | 0 | 1 |
[16] † Amiskwia Last Modified in 01/20/23 | 48 | 26 | 26 | 0 | 0 | 0 | 0 | 1 |
[17] Flacisaggita Last Modified in 01/20/23 | 0 | 68 | 68 | 0 | 6 | 0 | 0 | 1 |
[18] Spadella Last Modified in 01/20/23 | 0 | 68 | 68 | 0 | 6 | 0 | 0 | 1 |
[19] Eukrohnia Last Modified in 01/20/23 | 6 | 62 | 62 | 0 | 6 | 0 | 0 | 1 |
[20] † Protosagitta Last Modified in 01/20/23 | 57 | 16 | 16 | 0 | 1 | 0 | 0 | 1 |
[21] † Ankalodous Last Modified in 01/20/23 | 63 | 10 | 10 | 0 | 1 | 0 | 0 | 1 |
[22] † Capinatator Last Modified in 01/20/23 | 56 | 17 | 17 | 0 | 1 | 0 | 0 | 1 |
[23] Adineta Last Modified in 01/20/23 | 3 | 65 | 65 | 0 | 6 | 0 | 0 | 1 |
[24] † Branchionus Last Modified in 01/20/23 | 3 | 65 | 65 | 0 | 6 | 0 | 0 | 1 |
[25] Nemertea Last Modified in 01/20/23 | 1 | 61 | 61 | 0 | 12 | 0 | 0 | 1 |
[26] Onychophora Last Modified in 01/20/23 | 0 | 63 | 63 | 0 | 11 | 0 | 0 | 1 |
[27] Arthropoda Last Modified in 01/20/23 | 0 | 62 | 62 | 0 | 12 | 0 | 0 | 1 |
[28] Tardigrada Last Modified in 01/20/23 | 8 | 52 | 52 | 0 | 14 | 0 | 0 | 1 |
[29] Nematoda Last Modified in 01/20/23 | 7 | 51 | 51 | 0 | 16 | 0 | 0 | 1 |
[30] Nematomorpha Last Modified in 01/20/23 | 7 | 51 | 51 | 0 | 16 | 0 | 0 | 1 |
[31] Priapulida Last Modified in 01/20/23 | 0 | 62 | 62 | 0 | 12 | 0 | 0 | 1 |
[32] Kinorhyncha Last Modified in 01/20/23 | 10 | 52 | 52 | 0 | 12 | 0 | 0 | 1 |
[33] Loricifera Last Modified in 01/20/23 | 9 | 53 | 53 | 0 | 12 | 0 | 0 | 1 |
Taxonomic Overview for Matrix 'M29119' (33 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Bryozoa Last Modified in 01/20/23 | 2 | 110 | 110 | 0 | 26 | 0 | 0 | 1 |
[2] Brachiopoda Last Modified in 01/20/23 | 1 | 117 | 115 | 0 | 22 | 0 | 0 | 1 |
[3] Phoronida Last Modified in 01/20/23 | 5 | 110 | 110 | 0 | 23 | 0 | 0 | 1 |
[4] Annelida Last Modified in 01/20/23 | 0 | 120 | 120 | 0 | 18 | 0 | 0 | 1 |
[5] Sipuncula Last Modified in 01/20/23 | 9 | 106 | 106 | 0 | 23 | 0 | 0 | 1 |
[6] Mollusca Last Modified in 01/20/23 | 1 | 132 | 129 | 0 | 8 | 0 | 0 | 1 |
[7] † Wiwaxia Last Modified in 01/20/23 | 91 | 39 | 39 | 0 | 8 | 0 | 0 | 1 |
[8] † Odontogriphus Last Modified in 01/20/23 | 93 | 37 | 37 | 0 | 8 | 0 | 0 | 1 |
[9] † Canadia Last Modified in 01/20/23 | 80 | 43 | 43 | 0 | 15 | 0 | 0 | 1 |
[10] Entoprocta Last Modified in 01/20/23 | 10 | 104 | 104 | 0 | 24 | 0 | 0 | 1 |
[11] Platyhelminthes Last Modified in 01/20/23 | 1 | 109 | 109 | 0 | 28 | 0 | 0 | 1 |
[12] Gastrotricha Last Modified in 01/20/23 | 10 | 101 | 101 | 0 | 27 | 0 | 0 | 1 |
[13] Gnathostumulida Last Modified in 01/20/23 | 9 | 109 | 108 | 0 | 21 | 0 | 0 | 1 |
[14] Micrognathozoa Last Modified in 01/20/23 | 23 | 96 | 96 | 0 | 19 | 0 | 0 | 1 |
[15] † Inquicus fellatus Last Modified in 01/20/23 | 97 | 32 | 32 | 0 | 9 | 0 | 0 | 1 |
[16] † Amiskwia Last Modified in 01/20/23 | 93 | 37 | 37 | 0 | 8 | 0 | 0 | 1 |
[17] Flacisaggita Last Modified in 01/20/23 | 0 | 123 | 123 | 0 | 15 | 0 | 0 | 1 |
[18] Spadella Last Modified in 01/20/23 | 0 | 125 | 125 | 0 | 13 | 0 | 0 | 1 |
[19] Eukrohnia Last Modified in 01/20/23 | 6 | 119 | 119 | 0 | 13 | 0 | 0 | 1 |
[20] † Protosagitta Last Modified in 01/20/23 | 109 | 23 | 23 | 0 | 6 | 0 | 0 | 1 |
[21] † Ankalodous Last Modified in 01/20/23 | 122 | 14 | 14 | 0 | 2 | 0 | 0 | 1 |
[22] † Capinatator Last Modified in 01/20/23 | 107 | 24 | 24 | 0 | 7 | 0 | 0 | 1 |
[23] Adineta Last Modified in 01/20/23 | 3 | 116 | 116 | 0 | 19 | 0 | 0 | 1 |
[24] † Branchionus Last Modified in 01/20/23 | 3 | 116 | 116 | 0 | 19 | 0 | 0 | 1 |
[25] Nemertea Last Modified in 01/20/23 | 2 | 113 | 113 | 0 | 23 | 0 | 0 | 1 |
[26] Onychophora Last Modified in 01/20/23 | 0 | 117 | 117 | 0 | 21 | 0 | 0 | 1 |
[27] Arthropoda Last Modified in 01/20/23 | 0 | 117 | 116 | 0 | 22 | 0 | 0 | 1 |
[28] Tardigrada Last Modified in 01/20/23 | 8 | 106 | 106 | 0 | 24 | 0 | 0 | 1 |
[29] Nematoda Last Modified in 01/20/23 | 7 | 102 | 102 | 0 | 29 | 0 | 0 | 1 |
[30] Nematomorpha Last Modified in 01/20/23 | 7 | 102 | 102 | 0 | 29 | 0 | 0 | 1 |
[31] Priapulida Last Modified in 01/20/23 | 0 | 113 | 113 | 0 | 25 | 0 | 0 | 1 |
[32] Kinorhyncha Last Modified in 01/20/23 | 10 | 103 | 103 | 0 | 25 | 0 | 0 | 1 |
[33] Loricifera Last Modified in 01/20/23 | 9 | 104 | 104 | 0 | 25 | 0 | 0 | 1 |
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Specimen list | 370 | |
Bibliography | 174 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 7 | |
Project downloads | 1 | |
Matrix downloads | 6 | sm7151 (1 download); sm7153 (1 download); sm7154 (1 download); sm7155 (1 download); sm7157 (1 download); sm7158 (1 download); |