Project 4826: A. Bignon, C. Cronier. 2013. The systematics and phylogeny of the Devonian subfamily Asteropyginae (Trilobita: Phacopida). Journal of Systematic Palaeontology. 12 (6):637-668.
Abstract
Traditionally, the evolutionary scenarios of Asteropyginae have been organized in two major clades involving four pygidial patterns. The first cladistic analysis performed on the subfamily maintained the organization in two clades, though the phylogenetic relationships between genera were unexpected compared to the traditional scenarios. Moreover, the previous systematic scheme based on pygidial segmentation was rendered obsolete. In this study, we performed a new phylogenetic analysis from well-known taxa and recent discoveries. Sixty species assigned to 36 genera have been analysed from a dataset of 72 characters. Three more species are used as the outgroup. The 79 most parsimonious trees have a length of 492 steps and a retention index of 0.735. Our analysis confirms the monophyly of Asteropyginae excluding the genus Protacanthina. The phylogenetic pattern is unconventional in that a largely pectinate topology is resolved, rather than two clades corresponding to formerly delimited pygidial morphs. Nevertheless, most relationships between genera suggested in the traditional evolutionary scenarios of the subfamily have been recovered, the main differences being the origin of the group. Whereas Treveropyge was usually considered as the origin of the remaining Asteropyginae, the present results instead identify Destombesina as the sister group to all remaining members of the subfamily. From the new phylogenetic pattern, the diagnoses of genera are rewritten in a standard way and to delimit clades more accurately. This work implies modified generic assignments for some species and the erection of four new genera: Minicryphaeus gen. nov., Gandlops gen. nov., Morzadecops gen. nov. and Pennarbedops gen. nov. The systematic position of Erbenochile is also discussed, the genus no longer being assigned to Asteropyginae.Read the article »
Article DOI: 10.1080/14772019.2013.804008
Project DOI: 10.7934/P4826, http://dx.doi.org/10.7934/P4826
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 4432 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 72 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 180 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:72/0 |
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MorphoBank Project 4826
MorphoBank Project 4826
- Creation Date:
10 August 2023 - Publication Date:
04 September 2023 - Project views: 3155
Authors' Institutions
- Universite de Lille
- Yale University
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Shreya Jariwala Project Administrator | 63 | 4 | 4 | 72 | 0 | 0 | 4432 (4432, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29434' (63 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Neocalmonia yazdii Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[2] † Neocalmonia quadricosta Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[3] † Neocalmonia lutensis Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[4] † Neocalmonia brinkmanni Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[5] † Radiopyge heratensis Last Modified in 08/10/23 | 1 | 71 | 71 | 0 | 0 | 0 | 0 | 1 |
[6] † Radiopyge sardarensis Last Modified in 08/10/23 | 1 | 71 | 71 | 0 | 0 | 0 | 0 | 1 |
[7] † Jaylopygiella medusa Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[8] † Bradocryphaeus nodosa Last Modified in 08/10/23 | 1 | 71 | 71 | 0 | 0 | 0 | 0 | 1 |
[9] † Bradocryphaeus maillieuxi Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[10] † Bradocryphaeus vanherlei Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[11] † Bradocryphaeus hispanica Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[12] † Bradocryphaeus porsguenensis Last Modified in 08/10/23 | 1 | 71 | 71 | 0 | 0 | 0 | 0 | 1 |
[13] † Bradocryphaeus sp. cf. mosanus Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[14] † Heliopyge helios Last Modified in 08/10/23 | 0 | 72 | 72 | 0 | 0 | 0 | 0 | 1 |
[15] † Hallandclarkeops cantarmorica Last Modified in 08/10/23 | 1 | 71 | 71 | 0 | 0 | 0 | 0 | 1 |
[16] † Alcaldops troaonensis Last Modified in 08/10/23 | 0 | 72 | 72 | 0 | 0 | 0 | 0 | 1 |
[17] † Alcaldops renaudae Last Modified in 08/10/23 | 0 | 72 | 72 | 0 | 0 | 0 | 0 | 1 |
[18] † Alcaldops camfroutensis Last Modified in 08/10/23 | 0 | 72 | 72 | 0 | 0 | 0 | 0 | 1 |
[19] † Comura tuberculata Last Modified in 08/10/23 | 0 | 72 | 72 | 0 | 0 | 0 | 0 | 1 |
[20] † Psychopyge elegans Last Modified in 08/10/23 | 1 | 71 | 71 | 0 | 0 | 0 | 0 | 1 |
[21] † Psychopyge termierorum Last Modified in 08/10/23 | 1 | 71 | 71 | 0 | 0 | 0 | 0 | 1 |
[22] † Walliserops trifurcatus Last Modified in 08/10/23 | 1 | 71 | 71 | 0 | 0 | 0 | 0 | 1 |
[23] † Saharops bensaidi Last Modified in 08/10/23 | 1 | 71 | 71 | 0 | 0 | 0 | 0 | 1 |
[24] † Quadrops flexuosa Last Modified in 08/10/23 | 1 | 71 | 71 | 0 | 0 | 0 | 0 | 1 |
[25] † Coltraneia celtica Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[26] † Coltraneia oufatenensis Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[27] † Coltraneia saourensis Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[28] † Treveropyge prorotundifrons Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[29] † Pilletina praecursor Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[30] † Tolkienia cancer Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[31] † Hollardops struvei Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[32] † Hollardops mesocristata Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[33] † Hollardops boudibensis Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[34] † Hollardops burtandmimiae Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[35] † Pennarbedops babini Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[36] † Morzadecops zguidensis Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[37] † Asteropyge perforata Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[38] † Mrakibina cattoi Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[39] † Metacanthina barrandei Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[40] † Bellacartwrigthia jennyae Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[41] † Bellacartwrigthia phyllocaudata Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[42] † Braunops obsoletus Last Modified in 08/10/23 | 1 | 71 | 71 | 0 | 0 | 0 | 0 | 1 |
[43] † Rheingoldium marocanensis Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[44] † Rhenops anserinus Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[45] † Paracryphaeus jonesi Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[46] † Gandlops ogivalis Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[47] † Kayserops kerfornei Last Modified in 08/10/23 | 1 | 71 | 71 | 0 | 0 | 0 | 0 | 1 |
[48] † Kayserops inflatus Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[49] † Stummiana osismorum Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[50] † Deloops haasi Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[51] † Breizhops bigranulosus Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[52] † Greenops (neometacanthus) stellifer Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[53] † Greenops grabaui Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[54] † Greenops boothi Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[55] † Pseudocryphaeus michelini Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[56] † Minicryphaeus minimus Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[57] † Minicryphaeus quaterspinosus Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[58] † Destombesina ougartensis Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[59] † Destombesina tafilaltensis Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[60] † Protacanthina vellilae Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[61] † Pelitlina goltzi Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[62] † Acastella tyro Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[63] † Mimocryphaeus altecostatus Last Modified in 08/10/23 | 2 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 3155 | |
Project overview | 172 | |
Documents list | 117 | |
Taxon list | 1201 | |
Views for media list | 148 | |
Media views | 707 | Media search (409 views); M890600 (74 views); M890602 (77 views); M890601 (73 views); M890599 (74 views); |
Matrix views | 117 | Matrix landing page (110 views); Bignon and Cronier 2013 (7 views); |
Specimen list | 603 | |
Bibliography | 90 |