Project 5137: E. Martin-Silverstone, D. M. Unwin, A. R. Cuff, E. E. Brown, L. Allington-Jones, P. M. Barrett. 2024. A new pterosaur from the Middle Jurassic of Skye, Scotland and the early diversification of flying reptiles. Journal of Vertebrate Paleontology. null:null.
Abstract
The Middle Jurassic was a critical time in pterosaur evolution, witnessing the appearance of major morphological innovations that underpinned successive radiations by rhamphorhynchids, basally branching monofenestratans, and pterodactyloids. Frustratingly, this interval is particularly sparsely sampled, with a record consisting almost exclusively of isolated fragmentary remains. Here, we describe new material from the Bathonian-aged Kilmaluag Formation of Skye, Scotland, which helps close this gap. Ceoptera evansae (gen. et sp. nov.) is based on a three-dimensionally preserved partial skeleton, which represents one of the only associated Middle Jurassic pterosaurs. Ceoptera is among the first pterosaurs to be fully digitally prepared, and µCT scanning reveals multiple elements of the skeleton that remain fully embedded within the matrix and otherwise inaccessible. It is diagnosed by unique features of the pectoral and pelvic girdle. The inclusion of this new Middle Jurassic pterosaur in a novel phylogenetic analysis of pterosaur interrelationships provides additional support for the existence of the controversial clade Darwinoptera, adding to our knowledge of pterosaur diversity and evolution.Read the article »
Article DOI: 10.1080/02724634.2023.2298741
Project DOI: 10.7934/P5137, http://dx.doi.org/10.7934/P5137
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 6415 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 136 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 338 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:111/25 |
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MorphoBank Project 5137
MorphoBank Project 5137
- Creation Date:
18 March 2024 - Publication Date:
18 March 2024 - Project views: 996
Authors' Institutions
- Natural History Museum, London
- University of Birmingham
- University of Leicester
- University of Liverpool
- University of Bristol
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 69 | 1 | 1 | 136 | 0 | 0 | 6415 (6415, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29717' (69 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Euparkeria Last Modified in 03/18/24 | 10 | 126 | 126 | 0 | 0 | 0 | 0 | 1 |
[2] † Herrerasaurus Last Modified in 03/18/24 | 10 | 126 | 126 | 0 | 0 | 0 | 0 | 1 |
[3] † Preondactylus Last Modified in 03/18/24 | 31 | 105 | 105 | 0 | 0 | 0 | 0 | 1 |
[4] † Dimorphodon Last Modified in 03/18/24 | 13 | 124 | 123 | 0 | 0 | 0 | 0 | 1 |
[5] † Peteinosaurus Last Modified in 03/18/24 | 57 | 79 | 79 | 0 | 0 | 0 | 0 | 1 |
[6] † Anurognathus Last Modified in 03/18/24 | 23 | 113 | 113 | 0 | 0 | 0 | 0 | 1 |
[7] † Batrachognathus Last Modified in 03/18/24 | 41 | 95 | 95 | 0 | 0 | 0 | 0 | 1 |
[8] † Dendrorhynchoides Last Modified in 03/18/24 | 52 | 85 | 84 | 0 | 0 | 0 | 0 | 1 |
[9] † Jeholopterus Last Modified in 03/18/24 | 45 | 91 | 91 | 0 | 0 | 0 | 0 | 1 |
[10] † Austriadactylus Last Modified in 03/18/24 | 58 | 78 | 78 | 0 | 0 | 0 | 0 | 1 |
[11] † Campylognathoides Last Modified in 03/18/24 | 4 | 133 | 132 | 0 | 0 | 0 | 0 | 1 |
[12] † Eudimorphodon Last Modified in 03/18/24 | 11 | 131 | 125 | 0 | 0 | 0 | 0 | 1 |
[13] † Raeticodactylus Last Modified in 03/18/24 | 42 | 94 | 94 | 0 | 0 | 0 | 0 | 1 |
[14] † Scaphognathus Last Modified in 03/18/24 | 8 | 128 | 128 | 0 | 0 | 0 | 0 | 1 |
[15] † Sordes Last Modified in 03/18/24 | 14 | 123 | 122 | 0 | 0 | 0 | 0 | 1 |
[16] † Pterorhynchus Last Modified in 03/18/24 | 50 | 86 | 86 | 0 | 0 | 0 | 0 | 1 |
[17] † Cacibupteryx Last Modified in 03/18/24 | 90 | 46 | 46 | 0 | 0 | 0 | 0 | 1 |
[18] † Dorygnathus Last Modified in 03/18/24 | 5 | 131 | 131 | 0 | 0 | 0 | 0 | 1 |
[19] † Nesodactylus Last Modified in 03/18/24 | 94 | 42 | 42 | 0 | 0 | 0 | 0 | 1 |
[20] † Dearc Last Modified in 03/18/24 | 60 | 76 | 76 | 0 | 0 | 0 | 0 | 1 |
[21] † Rhamphorhynchus Last Modified in 03/18/24 | 1 | 136 | 135 | 0 | 0 | 0 | 0 | 1 |
[22] † Ceoptera Last Modified in 03/18/24 | 113 | 23 | 23 | 0 | 0 | 0 | 0 | 1 |
[23] † Allkaruen Last Modified in 03/18/24 | 81 | 55 | 55 | 0 | 0 | 0 | 0 | 1 |
[24] † Cuspicephalus Last Modified in 03/18/24 | 93 | 43 | 43 | 0 | 0 | 0 | 0 | 1 |
[25] † Kryptodrakon Last Modified in 03/18/24 | 123 | 13 | 13 | 0 | 0 | 0 | 0 | 1 |
[26] † Darwinopterus Last Modified in 03/18/24 | 6 | 136 | 130 | 0 | 0 | 0 | 0 | 1 |
[27] † Wukongopterus Last Modified in 03/18/24 | 57 | 79 | 79 | 0 | 0 | 0 | 0 | 1 |
[28] † Kunpengopterus Last Modified in 03/18/24 | 9 | 128 | 127 | 0 | 0 | 0 | 0 | 1 |
[29] † Changchengopterus Last Modified in 03/18/24 | 81 | 55 | 55 | 0 | 0 | 0 | 0 | 1 |
[30] † Douzhanopterus Last Modified in 03/18/24 | 94 | 42 | 42 | 0 | 0 | 0 | 0 | 1 |
[31] † Rhamphodactylus Last Modified in 03/18/24 | 7 | 130 | 129 | 0 | 0 | 0 | 0 | 1 |
[32] † Liaodactylus Last Modified in 03/18/24 | 73 | 63 | 63 | 0 | 0 | 0 | 0 | 1 |
[33] † Pterodactylus Last Modified in 03/18/24 | 5 | 134 | 131 | 0 | 0 | 0 | 0 | 1 |
[34] † Cycnorhamphus Last Modified in 03/18/24 | 15 | 121 | 121 | 0 | 0 | 0 | 0 | 1 |
[35] † Eosipterus Last Modified in 03/18/24 | 87 | 49 | 49 | 0 | 0 | 0 | 0 | 1 |
[36] † Aurorazhdarcho Last Modified in 03/18/24 | 10 | 127 | 126 | 0 | 0 | 0 | 0 | 1 |
[37] † Gnathosaurus Last Modified in 03/18/24 | 66 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[38] † Gegepterus Last Modified in 03/18/24 | 50 | 86 | 86 | 0 | 0 | 0 | 0 | 1 |
[39] † Gladocephaloideus Last Modified in 03/18/24 | 25 | 111 | 111 | 0 | 0 | 0 | 0 | 1 |
[40] † Ctenochasma Last Modified in 03/18/24 | 18 | 120 | 118 | 0 | 0 | 0 | 0 | 1 |
[41] † Pterodaustro Last Modified in 03/18/24 | 6 | 130 | 130 | 0 | 0 | 0 | 0 | 1 |
[42] † Istiodactylus Last Modified in 03/18/24 | 27 | 110 | 109 | 0 | 0 | 0 | 0 | 1 |
[43] † Nurhachius Last Modified in 03/18/24 | 34 | 102 | 102 | 0 | 0 | 0 | 0 | 1 |
[44] † Haopterus Last Modified in 03/18/24 | 53 | 83 | 83 | 0 | 0 | 0 | 0 | 1 |
[45] † Ornithocheirus Last Modified in 03/18/24 | 73 | 63 | 63 | 0 | 0 | 0 | 0 | 1 |
[46] † Anhanguera Last Modified in 03/18/24 | 10 | 128 | 126 | 0 | 0 | 0 | 0 | 1 |
[47] † Coloborhynchus Last Modified in 03/18/24 | 12 | 124 | 124 | 0 | 0 | 0 | 0 | 1 |
[48] † Ludodactylus Last Modified in 03/18/24 | 74 | 62 | 62 | 0 | 0 | 0 | 0 | 1 |
[49] † Boreopterus Last Modified in 03/18/24 | 36 | 101 | 100 | 0 | 0 | 0 | 0 | 1 |
[50] † Nyctosaurus Last Modified in 03/18/24 | 24 | 113 | 112 | 0 | 0 | 0 | 0 | 1 |
[51] † Muzquizopteryx Last Modified in 03/18/24 | 68 | 68 | 68 | 0 | 0 | 0 | 0 | 1 |
[52] † Pteranodon Last Modified in 03/18/24 | 21 | 115 | 115 | 0 | 0 | 0 | 0 | 1 |
[53] † Germanodactyluscrist Last Modified in 03/18/24 | 21 | 116 | 115 | 0 | 0 | 0 | 0 | 1 |
[54] † Germanodactylusrhamph Last Modified in 03/18/24 | 17 | 119 | 119 | 0 | 0 | 0 | 0 | 1 |
[55] † Dsungaripterus Last Modified in 03/18/24 | 26 | 110 | 110 | 0 | 0 | 0 | 0 | 1 |
[56] † Tatalpterosaur Last Modified in 03/18/24 | 37 | 99 | 99 | 0 | 0 | 0 | 0 | 1 |
[57] † Noripterus Last Modified in 03/18/24 | 87 | 49 | 49 | 0 | 0 | 0 | 0 | 1 |
[58] † Lonchodectes Last Modified in 03/18/24 | 86 | 52 | 50 | 0 | 0 | 0 | 0 | 1 |
[59] † Tapejara Last Modified in 03/18/24 | 36 | 100 | 100 | 0 | 0 | 0 | 0 | 1 |
[60] † Tupandactylus Last Modified in 03/18/24 | 29 | 107 | 107 | 0 | 0 | 0 | 0 | 1 |
[61] † Sinopterus Last Modified in 03/18/24 | 37 | 100 | 99 | 0 | 0 | 0 | 0 | 1 |
[62] † Huaxiapterus Last Modified in 03/18/24 | 45 | 92 | 91 | 0 | 0 | 0 | 0 | 1 |
[63] † Shenzhoupterus Last Modified in 03/18/24 | 53 | 83 | 83 | 0 | 0 | 0 | 0 | 1 |
[64] † Chaoyangopterus Last Modified in 03/18/24 | 70 | 66 | 66 | 0 | 0 | 0 | 0 | 1 |
[65] † Tupuxuara Last Modified in 03/18/24 | 25 | 111 | 111 | 0 | 0 | 0 | 0 | 1 |
[66] † Thalassodromeus Last Modified in 03/18/24 | 89 | 47 | 47 | 0 | 0 | 0 | 0 | 1 |
[67] † Quetzalcoatlus Last Modified in 03/18/24 | 43 | 93 | 93 | 0 | 0 | 0 | 0 | 1 |
[68] † Zhejiangopterus Last Modified in 03/18/24 | 40 | 96 | 96 | 0 | 0 | 0 | 0 | 1 |
[69] † Azhdarcho Last Modified in 03/18/24 | 93 | 43 | 43 | 0 | 0 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 996 | |
Project overview | 71 | |
Matrix views | 50 | Matrix landing page (45 views); matrix (5 views); |
Media views | 177 | Media search (151 views); M902669 (26 views); |
Views for media list | 59 | |
Specimen list | 92 | |
Bibliography | 34 | |
Taxon list | 513 |