Project 5140: R. J. O’Flynn, M. Williams, M.-X. Yu, J. Guo, D. Audo, M. Schmidt, H.-J. Mai, Y. Liu, G. D. Edgecombe. 2024. The early Cambrian Bushizheia yangi and head segmentation in upper stem-group euarthropods. Papers in Palaeontology. (In Press)
Abstract
Bushizheia yangi O’Flynn & Liu, a euarthropod from the lower Cambrian (Series 2, Stage 3) Chengjiang Konservat-Lagerstätte, possesses both raptorial frontal appendages sharing traits with stem-group euarthropods and deuteropodan characters shared by all extant euarthropods (e.g. dorsal arthrodisation and arthropodisation of all limbs). Microtomography of new specimens of B. yangi reveals previously unknown details of head morphology — a six-segmented head composed of an ocular segment that carries diminutive paired stalked eyes, a post-ocular segment with deutocerebral frontal appendages, and four subsequent cephalic segments each bearing a pair of appendages. The phylogenetic position of B. yangi in the euarthropod stem-group adds support to a six-segmented head being an ancestral state for Deuteropoda. This study also reports the first occurrence of possible eggs closely associated with B. yangi. The homology of raptorial frontal appendages between lower stem-group euarthropod radiodonts and deuteropods (i.e., upper stem- and crown-group euarthropods) is strengthened by similarity and phylogenetic continuity between B. yangi, its likely sister taxon Kiisortoqia soperi Stein, from the Sirius Passet Konservat-Lagerstätte, and other euarthropod taxa.Project DOI: 10.7934/P5140, http://dx.doi.org/10.7934/P5140
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 22387 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 283 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 643 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:283/0 |
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MorphoBank Project 5140
MorphoBank Project 5140
- Creation Date:
21 March 2024 - Publication Date:
29 March 2024 - Project views: 1174
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Robert O'Flynn Project Administrator | 85 | 1 | 1 | 283 | 0 | 0 | 22387 (14403, 0, 7984) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29729' (85 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Priapulus caudatus Last Modified in 03/29/24 | 2 | 53 | 53 | 0 | 228 | 0 | 0 | 1 |
[2] † Cricocosmia jinningensis Last Modified in 03/29/24 | 7 | 54 | 54 | 0 | 222 | 0 | 0 | 1 |
[3] Tardigrada Last Modified in 03/22/24 | 6 | 107 | 107 | 0 | 170 | 0 | 0 | 1 |
[4] † Microdictyon sinicum Last Modified in 03/29/24 | 14 | 116 | 116 | 0 | 153 | 0 | 0 | 1 |
[5] † Paucipodia inermis Last Modified in 03/29/24 | 16 | 109 | 109 | 0 | 158 | 0 | 0 | 1 |
[6] † Cardiodictyon catenulum Last Modified in 03/29/24 | 17 | 114 | 114 | 0 | 152 | 0 | 0 | 1 |
[7] † Hallucigenia sparsa Last Modified in 03/29/24 | 11 | 120 | 120 | 0 | 152 | 0 | 0 | 1 |
[8] † Hallucigenia fortis Last Modified in 03/29/24 | 14 | 117 | 117 | 0 | 152 | 0 | 0 | 1 |
[9] † Collinsium ciliosum Last Modified in 03/29/24 | 18 | 114 | 114 | 0 | 151 | 0 | 0 | 1 |
[10] † Luolishania longicruris Last Modified in 03/29/24 | 14 | 118 | 118 | 0 | 151 | 0 | 0 | 1 |
[11] † Onychodictyon ferox Last Modified in 03/29/24 | 14 | 117 | 117 | 0 | 152 | 0 | 0 | 1 |
[12] † Aysheaia pedunculata Last Modified in 03/29/24 | 12 | 114 | 114 | 0 | 157 | 0 | 0 | 1 |
[13] † Antennacanthopodia gracilis Last Modified in 03/29/24 | 13 | 112 | 112 | 0 | 158 | 0 | 0 | 1 |
[14] Onychophora Last Modified in 03/29/24 | 5 | 117 | 117 | 0 | 161 | 0 | 0 | 1 |
[15] † Megadictyon haikouensis Last Modified in 03/29/24 | 17 | 108 | 108 | 0 | 158 | 0 | 0 | 1 |
[16] † Pambdelurion whittingtoni Last Modified in 03/29/24 | 40 | 109 | 109 | 0 | 134 | 0 | 0 | 1 |
[17] † Kerygmachela kierkegaardi Last Modified in 03/29/24 | 43 | 110 | 110 | 0 | 130 | 0 | 0 | 1 |
[18] † Opabinia regalis Last Modified in 03/29/24 | 43 | 107 | 107 | 0 | 133 | 0 | 0 | 1 |
[19] † Anomalocaris canadensis Last Modified in 03/29/24 | 23 | 142 | 142 | 0 | 118 | 0 | 0 | 1 |
[20] † Lyrarapax unguispinus Last Modified in 03/29/24 | 26 | 139 | 139 | 0 | 118 | 0 | 0 | 1 |
[21] † Amplectobelua symbrachiata Last Modified in 03/29/24 | 18 | 147 | 147 | 0 | 118 | 0 | 0 | 1 |
[22] † Peytoia nathorsti Last Modified in 03/29/24 | 25 | 140 | 140 | 0 | 118 | 0 | 0 | 1 |
[23] † Stanleycaris hirpex Last Modified in 03/29/24 | 29 | 137 | 137 | 0 | 117 | 0 | 0 | 1 |
[24] † Aegirocassis benmoulai Last Modified in 03/29/24 | 67 | 102 | 102 | 0 | 114 | 0 | 0 | 1 |
[25] † Hurdia victoria Last Modified in 03/29/24 | 31 | 134 | 134 | 0 | 118 | 0 | 0 | 1 |
[26] † Schinderhannes bartelsi Last Modified in 03/29/24 | 46 | 121 | 121 | 0 | 116 | 0 | 0 | 1 |
[27] † Fengzhengia mamingae Last Modified in 03/29/24 | 45 | 168 | 168 | 0 | 70 | 0 | 0 | 1 |
[28] † Kylinxia zhangi Last Modified in 03/29/24 | 12 | 203 | 203 | 0 | 68 | 0 | 0 | 1 |
[29] † Kiisortoqia soperi Last Modified in 03/29/24 | 32 | 172 | 172 | 0 | 79 | 0 | 0 | 1 |
[30] † Bushizheia yangi Last Modified in 03/29/24 | 32 | 175 | 175 | 0 | 76 | 0 | 0 | 1 |
[31] † Habelia optata Last Modified in 03/29/24 | 14 | 198 | 198 | 0 | 71 | 0 | 0 | 1 |
[32] † Mollisonia plenovenatrix Last Modified in 03/29/24 | 14 | 200 | 200 | 0 | 69 | 0 | 0 | 1 |
[33] Limulus polyphemus Last Modified in 03/29/24 | 1 | 210 | 210 | 0 | 72 | 0 | 0 | 1 |
[34] † Dibasterium durgae Last Modified in 03/29/24 | 24 | 187 | 187 | 0 | 72 | 0 | 0 | 1 |
[35] † Offacolus kingi Last Modified in 03/29/24 | 24 | 187 | 187 | 0 | 72 | 0 | 0 | 1 |
[36] † Haikoucaris ercaiensis Last Modified in 03/29/24 | 17 | 197 | 197 | 0 | 69 | 0 | 0 | 1 |
[37] † Yawunik kootenayi Last Modified in 03/29/24 | 7 | 208 | 208 | 0 | 68 | 0 | 0 | 1 |
[38] † Alalcomenaeus cambricus Last Modified in 03/29/24 | 6 | 209 | 209 | 0 | 68 | 0 | 0 | 1 |
[39] † Leanchoilia illecebrosa Last Modified in 03/29/24 | 6 | 209 | 209 | 0 | 68 | 0 | 0 | 1 |
[40] † Leanchoilia superlata Last Modified in 03/29/24 | 7 | 208 | 208 | 0 | 68 | 0 | 0 | 1 |
[41] † Fortiforceps foliosa Last Modified in 03/29/24 | 7 | 206 | 206 | 0 | 70 | 0 | 0 | 1 |
[42] † Tanglangia caudata Last Modified in 03/29/24 | 40 | 175 | 175 | 0 | 68 | 0 | 0 | 1 |
[43] † Yohoia tenuis Last Modified in 03/29/24 | 8 | 205 | 205 | 0 | 70 | 0 | 0 | 1 |
[44] † Sklerolibyon maomima Last Modified in 03/29/24 | 11 | 202 | 202 | 0 | 70 | 0 | 0 | 1 |
[45] † Jianfengia multisegmentalis Last Modified in 03/29/24 | 8 | 205 | 205 | 0 | 70 | 0 | 0 | 1 |
[46] † Forfexicaris valida Last Modified in 03/29/24 | 89 | 135 | 135 | 0 | 59 | 0 | 0 | 1 |
[47] † Occacaris oviformis Last Modified in 03/29/24 | 63 | 161 | 161 | 0 | 59 | 0 | 0 | 1 |
[48] † Surusicaris elegans Last Modified in 03/29/24 | 27 | 191 | 191 | 0 | 65 | 0 | 0 | 1 |
[49] † Isoxys auritus Last Modified in 03/29/24 | 20 | 198 | 198 | 0 | 65 | 0 | 0 | 1 |
[50] † Isoxys volucris Last Modified in 03/29/24 | 67 | 154 | 154 | 0 | 62 | 0 | 0 | 1 |
[51] † Isoxys curvirostratus Last Modified in 03/29/24 | 19 | 199 | 199 | 0 | 65 | 0 | 0 | 1 |
[52] † Isoxys acutangulus Last Modified in 03/29/24 | 23 | 195 | 195 | 0 | 65 | 0 | 0 | 1 |
[53] † Retifacies abnormalis Last Modified in 03/29/24 | 7 | 199 | 199 | 0 | 77 | 0 | 0 | 1 |
[54] † Sinoburius lunaris Last Modified in 03/29/24 | 15 | 192 | 192 | 0 | 76 | 0 | 0 | 1 |
[55] † Cindarella eucalla Last Modified in 03/29/24 | 5 | 202 | 202 | 0 | 76 | 0 | 0 | 1 |
[56] † Xandarella spectaculum Last Modified in 03/29/24 | 5 | 202 | 202 | 0 | 76 | 0 | 0 | 1 |
[57] † Olenoides serratus Last Modified in 03/29/24 | 4 | 203 | 203 | 0 | 76 | 0 | 0 | 1 |
[58] † Hongshiyanaspis yiliangensis Last Modified in 03/29/24 | 3 | 204 | 204 | 0 | 76 | 0 | 0 | 1 |
[59] † Eoredlichia intermedia Last Modified in 03/29/24 | 4 | 203 | 203 | 0 | 76 | 0 | 0 | 1 |
[60] † Misszhouia longicaudata Last Modified in 03/29/24 | 15 | 192 | 192 | 0 | 76 | 0 | 0 | 1 |
[61] † Naraoia compacta Last Modified in 03/29/24 | 15 | 192 | 192 | 0 | 76 | 0 | 0 | 1 |
[62] † Naraoia spinosa Last Modified in 03/29/24 | 15 | 192 | 192 | 0 | 76 | 0 | 0 | 1 |
[63] † Arthroaspis bergstroemi Last Modified in 03/29/24 | 13 | 195 | 195 | 0 | 75 | 0 | 0 | 1 |
[64] † Tegopelte gigas Last Modified in 03/29/24 | 10 | 197 | 197 | 0 | 76 | 0 | 0 | 1 |
[65] † Saperion glumaceum Last Modified in 03/29/24 | 6 | 201 | 201 | 0 | 76 | 0 | 0 | 1 |
[66] † Helmetia expansa Last Modified in 03/29/24 | 73 | 139 | 139 | 0 | 71 | 0 | 0 | 1 |
[67] † Kuamaia lata Last Modified in 03/29/24 | 6 | 203 | 203 | 0 | 74 | 0 | 0 | 1 |
[68] † Chengjiangocaris kunmingensis Last Modified in 03/29/24 | 6 | 204 | 204 | 0 | 73 | 0 | 0 | 1 |
[69] † Alacaris mirabilis Last Modified in 03/29/24 | 7 | 203 | 203 | 0 | 73 | 0 | 0 | 1 |
[70] † Liangwangshania biloba Last Modified in 03/29/24 | 10 | 199 | 199 | 0 | 74 | 0 | 0 | 1 |
[71] † Guangweicaris spinatus Last Modified in 03/29/24 | 12 | 198 | 198 | 0 | 73 | 0 | 0 | 1 |
[72] † Fuxianhuia protensa Last Modified in 03/29/24 | 3 | 207 | 207 | 0 | 73 | 0 | 0 | 1 |
[73] Narceus americanus Last Modified in 03/22/24 | 1 | 192 | 192 | 0 | 90 | 0 | 0 | 1 |
[74] Scutigera coleoptrata Last Modified in 03/22/24 | 3 | 190 | 190 | 0 | 90 | 0 | 0 | 1 |
[75] Artemia salina Last Modified in 03/22/24 | 2 | 212 | 212 | 0 | 69 | 0 | 0 | 1 |
[76] † Yicaris dianensis Last Modified in 03/29/24 | 7 | 207 | 207 | 0 | 69 | 0 | 0 | 1 |
[77] † Hesslandona unisulcata Last Modified in 03/29/24 | 17 | 197 | 197 | 0 | 69 | 0 | 0 | 1 |
[78] † Martinssonia elongata Last Modified in 03/29/24 | 10 | 204 | 204 | 0 | 69 | 0 | 0 | 1 |
[79] † Waptia fieldensis Last Modified in 03/29/24 | 7 | 211 | 211 | 0 | 65 | 0 | 0 | 1 |
[80] † Clypecaris serrata Last Modified in 03/29/24 | 48 | 173 | 173 | 0 | 62 | 0 | 0 | 1 |
[81] † Canadaspis perfecta Last Modified in 03/29/24 | 12 | 206 | 206 | 0 | 65 | 0 | 0 | 1 |
[82] † Tokummia katalepsis Last Modified in 03/29/24 | 16 | 203 | 203 | 0 | 64 | 0 | 0 | 1 |
[83] † Branchiocaris pretiosa Last Modified in 03/29/24 | 23 | 196 | 196 | 0 | 64 | 0 | 0 | 1 |
[84] † Odaraia alata Last Modified in 03/29/24 | 64 | 159 | 159 | 0 | 60 | 0 | 0 | 1 |
[85] † Jugatacaris agilis Last Modified in 03/29/24 | 30 | 191 | 191 | 0 | 62 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 1174 | |
Project overview | 95 | |
Specimen list | 90 | |
Bibliography | 44 | |
Views for media list | 56 | |
Matrix views | 66 | Matrix landing page (59 views); Appendix_P5140 (7 views); |
Media views | 196 | Media search (170 views); M902797 (26 views); |
Taxon list | 625 | |
Documents list | 2 |