Project 5155: B. Holgado, R. Pegas. 2020. A taxonomic and phylogenetic review of the anhanguerid pterosaur group Coloborhynchinae and the new clade Tropeognathinae. Acta Palaeontologica Polonica. 65:null.
Abstract
Anhanguerids are a particular group of pterodactyloid pterosaurs, characterized mainly by their rostral sagittal crests, well laterally expanded jaw tips and enlarged anterior teeth. Due to the fragmentary nature of most known specimens, including holotypes, the taxonomy of the group has proved particularly difficult and controversial. Coloborhynchinae is a recently proposed clade within the Anhangueridae, and was defined as the most inclusive clade containing Coloborhynchus clavirostris but not Anhanguera or Ludodactylus. Coloborhynchinae was originally thought to include Coloborhynchus, Uktenadactylus, and Siroccopteryx. Here we present a reassessment of the taxonomy and phylogeny of all proposed members of the Coloborhynchinae and Coloborhynchus complex, with new anatomical comparisons and a novel phylogenetic analysis. Several features allow us to establish that coloborhynchines were much more diverse than previously thought, englobing four genera and seven species: Aerodraco sedgwickii gen. et comb. nov., Coloborhynchus clavirostris, Nicorhynchus capito gen. et comb. nov., Nicorhynchus fluviferox gen. et comb. nov., Uktenadactylus rodriguesae sp. nov., and Uktenadactylus wadleighi. Nicorhynchus and Uktenadactylus are considered sister taxa, being distinct on the basis of several rostral characters. Although with a homoplastic flat rostrum surface, Siroccopteryx was recovered out of the Coloborhynchinae, as sister taxon of Tropeognathus, due to similarities on the palatal ridge (which is broad and deep, and starting at the same level) and the relatively stout teeth compared to other anhanguerids. Tropeognathus and Siroccopteryx are further related to the Australian taxa Ferrodraco and Mythunga, which are all grouped in a new clade: the Tropeognathinae. Our analysis suggests that morphological evolution within anhanguerids was quite more complex than previously thought, with coloborhynchines representing the oldest recorded lineage of Anhangueridae, which achieved a worldwide distribution at least from the Aptian to the Cenomanian.Read the article »
Article DOI: 10.4202/app.00751.2020
Project DOI: 10.7934/P5155, http://dx.doi.org/10.7934/P5155
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 6886 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 179 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 460 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:179/0 |
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MorphoBank Project 5155
MorphoBank Project 5155
- Creation Date:
26 March 2024 - Publication Date:
27 March 2024 - Project views: 1454
- Matrix downloads: 1
Authors' Institutions
- Universidade Federal do Rio De Janeiro (UFRJ)
- Universitat Autònoma de Barcelona
- Alameda da Universidade
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Shreya Jariwala Project Administrator | 74 | 4 | 4 | 179 | 0 | 0 | 6886 (6056, 0, 830) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29741' (74 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Ikrandraco avatar Last Modified in 03/26/24 | 51 | 110 | 110 | 0 | 18 | 0 | 0 | 1 |
[2] † Lonchodraco giganteus Last Modified in 03/26/24 | 122 | 47 | 47 | 0 | 10 | 0 | 0 | 1 |
[3] † Haopterus gracilis Last Modified in 03/26/24 | 87 | 76 | 76 | 0 | 16 | 0 | 0 | 1 |
[4] † Hongshanopterus lacustris Last Modified in 03/26/24 | 111 | 56 | 56 | 0 | 12 | 0 | 0 | 1 |
[5] † Nurhachius ignaciobritoi Last Modified in 03/26/24 | 34 | 126 | 126 | 0 | 19 | 0 | 0 | 1 |
[6] † Nurhachius luei Last Modified in 03/26/24 | 75 | 89 | 89 | 0 | 15 | 0 | 0 | 1 |
[7] † Liaoxipterus brachyognathus Last Modified in 03/26/24 | 149 | 30 | 30 | 0 | 0 | 0 | 0 | 1 |
[8] † Istiodactylus latidens Last Modified in 03/26/24 | 45 | 114 | 114 | 0 | 20 | 0 | 0 | 1 |
[9] † Istiodactylus sinensis Last Modified in 03/26/24 | 57 | 102 | 102 | 0 | 20 | 0 | 0 | 1 |
[10] † Zhenyuanopterus longirostris Last Modified in 03/26/24 | 56 | 113 | 113 | 0 | 10 | 0 | 0 | 1 |
[11] † Boreopterus giganticus Last Modified in 03/26/24 | 100 | 65 | 65 | 0 | 14 | 0 | 0 | 1 |
[12] † Boreopterus cuiae Last Modified in 03/26/24 | 109 | 58 | 58 | 0 | 12 | 0 | 0 | 1 |
[13] † Ornithocheirus simus Last Modified in 03/26/24 | 139 | 37 | 37 | 0 | 3 | 0 | 0 | 1 |
[14] † Targaryendraco wiedenrothi Last Modified in 03/26/24 | 148 | 29 | 29 | 0 | 2 | 0 | 0 | 1 |
[15] † Aussiedraco molnari Last Modified in 03/26/24 | 158 | 19 | 19 | 0 | 2 | 0 | 0 | 1 |
[16] † Barbosania gracilirostris Last Modified in 03/26/24 | 73 | 88 | 88 | 0 | 18 | 0 | 0 | 1 |
[17] † Cimoliopterus cuvieri Last Modified in 03/26/24 | 132 | 45 | 45 | 0 | 2 | 0 | 0 | 1 |
[18] † Cimoliopterus dunni Last Modified in 03/26/24 | 143 | 35 | 35 | 0 | 1 | 0 | 0 | 1 |
[19] † Aetodactylus halli Last Modified in 03/26/24 | 142 | 29 | 29 | 0 | 8 | 0 | 0 | 1 |
[20] † Camposipterus nasutus Last Modified in 03/26/24 | 134 | 37 | 37 | 0 | 8 | 0 | 0 | 1 |
[21] † Hamipterus tianshanensis Last Modified in 03/26/24 | 33 | 140 | 140 | 0 | 6 | 0 | 0 | 1 |
[22] † Iberodactylus andreui Last Modified in 03/26/24 | 128 | 51 | 51 | 0 | 0 | 0 | 0 | 1 |
[23] † Anhanguera n. sp. Last Modified in 03/27/24 | 35 | 143 | 143 | 0 | 1 | 0 | 0 | 1 |
[24] † Anhanguera blittersdorffi Last Modified in 03/27/24 | 63 | 115 | 115 | 0 | 1 | 0 | 0 | 1 |
[25] † Anhanguera piscator Last Modified in 03/26/24 | 21 | 157 | 157 | 0 | 1 | 0 | 0 | 1 |
[26] † Liaoningopterus gui Last Modified in 03/26/24 | 96 | 83 | 83 | 0 | 0 | 0 | 0 | 1 |
[27] † Cearadactylus atrox Last Modified in 03/26/24 | 80 | 97 | 97 | 0 | 2 | 0 | 0 | 1 |
[28] † Maaradactylus kellneri Last Modified in 03/26/24 | 90 | 89 | 89 | 0 | 0 | 0 | 0 | 1 |
[29] † Guidraco venator Last Modified in 03/26/24 | 86 | 90 | 90 | 0 | 3 | 0 | 0 | 1 |
[30] † Ludodactylus sibbicki Last Modified in 03/26/24 | 74 | 104 | 104 | 0 | 1 | 0 | 0 | 1 |
[31] † Caulkicephalus trimicrodon Last Modified in 03/26/24 | 123 | 56 | 56 | 0 | 0 | 0 | 0 | 1 |
[32] † Ferrodraco lentoni Last Modified in 03/26/24 | 106 | 73 | 73 | 0 | 0 | 0 | 0 | 1 |
[33] † Mythunga camara Last Modified in 03/26/24 | 161 | 18 | 18 | 0 | 0 | 0 | 0 | 1 |
[34] † Tropeognathus mesembrinus Last Modified in 03/26/24 | 38 | 140 | 140 | 0 | 1 | 0 | 0 | 1 |
[35] † Siroccopteryx moroccensis Last Modified in 03/26/24 | 124 | 55 | 55 | 0 | 0 | 0 | 0 | 1 |
[36] † Aerodraco sedgwickii Last Modified in 03/26/24 | 139 | 40 | 40 | 0 | 0 | 0 | 0 | 1 |
[37] † Coloborhynchus clavirostris Last Modified in 03/26/24 | 133 | 46 | 46 | 0 | 0 | 0 | 0 | 1 |
[38] † Uktenadactylus wadleighi Last Modified in 03/26/24 | 133 | 46 | 46 | 0 | 0 | 0 | 0 | 1 |
[39] † Uktenadactylus rodriguesae Last Modified in 03/26/24 | 158 | 21 | 21 | 0 | 0 | 0 | 0 | 1 |
[40] † Nicorhynchus capito Last Modified in 03/26/24 | 150 | 29 | 29 | 0 | 0 | 0 | 0 | 1 |
[41] † Nicorhynchus fluviferox Last Modified in 03/26/24 | 155 | 24 | 24 | 0 | 0 | 0 | 0 | 1 |
[42] † Ornithosuchus woodwardi Last Modified in 03/26/24 | 17 | 145 | 145 | 0 | 17 | 0 | 0 | 1 |
[43] † Herrerasaurus ischigualastensis Last Modified in 03/26/24 | 15 | 147 | 147 | 0 | 17 | 0 | 0 | 1 |
[44] † Scleromochlus taylori Last Modified in 03/26/24 | 62 | 102 | 102 | 0 | 15 | 0 | 0 | 1 |
[45] † Anurognathus ammoni Last Modified in 03/26/24 | 34 | 128 | 128 | 0 | 17 | 0 | 0 | 1 |
[46] † Rhamphorhynchus muensteri Last Modified in 03/26/24 | 16 | 146 | 146 | 0 | 17 | 0 | 0 | 1 |
[47] † Darwinopterus linglongtaensis Last Modified in 03/26/24 | 50 | 120 | 120 | 0 | 9 | 0 | 0 | 1 |
[48] † Darwinopterus robustodens Last Modified in 03/26/24 | 59 | 111 | 111 | 0 | 9 | 0 | 0 | 1 |
[49] † Kunpengopterus sinensis Last Modified in 03/26/24 | 25 | 133 | 133 | 0 | 21 | 0 | 0 | 1 |
[50] † Pterodactylus antiquus Last Modified in 03/26/24 | 28 | 136 | 136 | 0 | 15 | 0 | 0 | 1 |
[51] † Nyctosaurus gracilis Last Modified in 03/26/24 | 29 | 122 | 122 | 0 | 28 | 0 | 0 | 1 |
[52] † Muzquizopteryx coahuilensis Last Modified in 03/26/24 | 106 | 59 | 59 | 0 | 14 | 0 | 0 | 1 |
[53] † Pteranodon longiceps Last Modified in 03/26/24 | 17 | 136 | 136 | 0 | 26 | 0 | 0 | 1 |
[54] † Tethydraco regalis Last Modified in 03/26/24 | 171 | 6 | 6 | 0 | 2 | 0 | 0 | 1 |
[55] † Dsungaripterus weii Last Modified in 03/26/24 | 28 | 134 | 134 | 0 | 17 | 0 | 0 | 1 |
[56] † Noripterus parvus Last Modified in 03/26/24 | 70 | 92 | 92 | 0 | 17 | 0 | 0 | 1 |
[57] † Keresdrakon vilsoni Last Modified in 03/26/24 | 91 | 61 | 61 | 0 | 27 | 0 | 0 | 1 |
[58] † Tupuxuara leonardii Last Modified in 03/26/24 | 32 | 127 | 127 | 0 | 20 | 0 | 0 | 1 |
[59] † Thalassodromeus sethi Last Modified in 03/26/24 | 65 | 94 | 94 | 0 | 20 | 0 | 0 | 1 |
[60] † Aymberedactylus cearensis Last Modified in 03/26/24 | 139 | 24 | 24 | 0 | 16 | 0 | 0 | 1 |
[61] † Tupandactylus imperator Last Modified in 03/26/24 | 90 | 71 | 71 | 0 | 18 | 0 | 0 | 1 |
[62] † Caiuajara dobruski Last Modified in 03/26/24 | 54 | 105 | 105 | 0 | 20 | 0 | 0 | 1 |
[63] † Europejara olcadesorum Last Modified in 03/26/24 | 128 | 35 | 35 | 0 | 16 | 0 | 0 | 1 |
[64] † Tapejara wellnhoferi Last Modified in 03/26/24 | 34 | 125 | 125 | 0 | 20 | 0 | 0 | 1 |
[65] † Sinopterus dongi Last Modified in 03/26/24 | 44 | 115 | 115 | 0 | 20 | 0 | 0 | 1 |
[66] † Huaxiapterus corollatus Last Modified in 03/26/24 | 92 | 69 | 69 | 0 | 18 | 0 | 0 | 1 |
[67] † Huaxiapterus benxiensis Last Modified in 03/26/24 | 74 | 87 | 87 | 0 | 18 | 0 | 0 | 1 |
[68] † Eopteranodon lii Last Modified in 03/26/24 | 112 | 51 | 51 | 0 | 16 | 0 | 0 | 1 |
[69] † Chaoyangopterus zhangi Last Modified in 03/26/24 | 91 | 68 | 68 | 0 | 20 | 0 | 0 | 1 |
[70] † Shenzhoupterus chaoyangensis Last Modified in 03/26/24 | 72 | 89 | 89 | 0 | 18 | 0 | 0 | 1 |
[71] † Jidapterus edentus Last Modified in 03/26/24 | 89 | 66 | 66 | 0 | 24 | 0 | 0 | 1 |
[72] † Quetzalcoatlus sp. Last Modified in 03/26/24 | 48 | 104 | 104 | 0 | 27 | 0 | 0 | 1 |
[73] † Zhejiangopterus linhaiensis Last Modified in 03/26/24 | 58 | 91 | 91 | 0 | 30 | 0 | 0 | 1 |
[74] † Azhdarcho lancicollis Last Modified in 03/26/24 | 129 | 35 | 35 | 0 | 15 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 1454 | |
Matrix views | 63 | Matrix landing page (56 views); Holgado and Pegas 2020 (7 views); |
Project overview | 78 | |
Documents list | 50 | |
Media views | 284 | Media search (178 views); M902711 (27 views); M902710 (25 views); M902709 (27 views); M902708 (27 views); |
Specimen list | 214 | |
Taxon list | 664 | |
Bibliography | 36 | |
Views for media list | 65 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 1 | |
Matrix downloads | 1 | Holgado and Pegas 2020 (1 download); |