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Nesbitt, S. J., Denton, R. K., Loewen, M. A., Brusatte, S. L., Smith, N. D., Turner, A. H., Kirkland, J. I., McDonald, A. T. and Wolfe, D. G. 2019. A mid-Cretaceous tyrannosauroid and the origin of North American end-Cretaceous dinosaur assemblages. Nature Ecology & Evolution. Vol. 3(6), pp. 892-899.