Project 1013: I. Vea, D. A. Grimaldi. 2015. Isabelle M Vea
Diverse New Scale Insects (Hemiptera: Coccoidea) in Amber from the Cretaceous and Eocene with a Phylogenetic Framework for Fossil Coccoidea. American Museum Novitates. 3823:1-80.
Specimen: † Rosahendersonia prisca Vea and Grimaldi (AMNH/AMNH Bu-835)
View: Light compound microscopy
View: Light compound microscopy
Abstract
Coccoids are abundant and diverse in most amber deposits around the world, but largely as macropterous males. Based on a study of male coccoids in Lebanese amber (Early Cretaceous), Burmese amber (Albian-Cenomanian), Cambay amber from western India (Early Eocene), and Baltic amber (mid-Eocene), 16 new species, 11 new genera, and three new families are added to the coccoid fossil record: Apticoccidae, n. fam., based on Apticoccus Koteja and Azar, and including two new species A. fortis, n. sp., and A. longitenuis, n. sp.; the monotypic family Hodgsonicoccidae, n. fam., including Hodgsonicoccus patefactus, n. gen., n. sp.; Kozariidae, n. fam., including Kozarius achronus, n. gen., n. sp., and K. perpetuus, n. sp.; the first occurrence of a Coccidae in Burmese amber, Rosahendersonia prisca, n. gen., n. sp.; the first fossil record of a Margarodidae sensu stricto, Heteromargarodes hukamsinghi, n. sp.; a peculiar Diaspididae in Indian amber, Normarkicoccus cambayae, n. gen., n. sp.; a Pityococcidae from Baltic amber, Pityococcus moniliformalis, n. sp., two Pseudococcidae in Lebanese and Burmese ambers, Williamsicoccus megalops, n. gen., n. sp., and Gilderius eukrinops, n. gen., n. sp.; an Early Cretaceous Weitschatidae, Pseudoweitschatus audebertis, n. gen., n. sp.; four genera considered incertae sedis, Alacrena peculiaris, n. gen., n. sp., Magnilens glaesaria, n. gen., n. sp., and Pedicellicoccus marginatus, n. gen., n. sp., and Xiphos vani, n. gen., n. sp. Interpretation of fossil coccoids is supported by a parsimony phylogenetic analysis based on 174 morphological characters (both adult males and females) and 112 taxa (69 Recent and 43 extinct).Read the article »
Article DOI: 10.1206/3823.1
Project DOI: 10.7934/P1013, http://dx.doi.org/10.7934/P1013
| This project contains | Matrices |
|---|---|
Download Project SDD File  | Total scored cells: 14995 Total media associated with cells: 0 Total labels associated with cell media: 0 |
| Characters | |
| Total characters: 174 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 510 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:174/0 |
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MorphoBank Project 1013
MorphoBank Project 1013
- Creation Date:
02 September 2013 - Publication Date:
05 December 2016 - Project views: 59444
- Matrix downloads: 8
This research
supported by
Authors' Institutions
- American Museum of Natural History
Members
| member name | taxa |
specimens |
media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
| Isabelle Vea Project Administrator | 143 | 19 | 31 | 174 | 0 | 0 | 14995 (14990, 0, 5) | 0 | 0 | 0 | ||||
Taxonomic Overview for Matrix 'M2800' (112 Taxa)
| taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
| [1] Aclerda arundinariae Last Modified in 12/10/14 | 2 | 172 | 0 | 0 | 0 | 0 | 0 | 1 |
| [2] Acyrthosiphon pisum Last Modified in 12/05/13 | 7 | 167 | 0 | 0 | 0 | 0 | 0 | 1 |
| [3] Adelges tsugae Last Modified in 12/05/13 | 53 | 121 | 0 | 0 | 0 | 0 | 0 | 1 |
| [4] † Alacrena peculiaris Last Modified in 12/10/14 | 102 | 72 | 0 | 0 | 0 | 0 | 0 | 1 |
| [5] † Albicoccus dimai Last Modified in 12/10/14 | 112 | 62 | 0 | 0 | 0 | 0 | 0 | 1 |
| [6] Antonina graminis crawi Last Modified in 12/10/14 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [7] Aonidiella aurantii Last Modified in 12/10/14 | 0 | 174 | 0 | 0 | 0 | 0 | 0 | 1 |
| [8] † Apticoccus fortis Last Modified in 12/10/14 | 95 | 79 | 0 | 0 | 0 | 0 | 0 | 1 |
| [9] † Apticoccus longitenuis Last Modified in 12/10/14 | 98 | 76 | 0 | 0 | 0 | 0 | 0 | 1 |
| [10] † Apticoccus minutus Last Modified in 12/10/14 | 94 | 80 | 0 | 0 | 0 | 0 | 0 | 1 |
| [11] † Undescribed undescribed Last Modified in 12/10/14 | 125 | 49 | 0 | 0 | 0 | 0 | 0 | 1 |
| [12] † Arnoldus capitatus Last Modified in 12/10/14 | 98 | 76 | 0 | 0 | 0 | 0 | 0 | 1 |
| [13] Beesonia dipterocarpi Last Modified in 12/10/14 | 12 | 162 | 0 | 0 | 0 | 0 | 0 | 1 |
| [14] † Burmacoccus danyi Last Modified in 12/10/14 | 106 | 68 | 0 | 0 | 0 | 0 | 0 | 1 |
| [15] Callipappus Last Modified in 12/10/14 | 2 | 172 | 0 | 0 | 0 | 0 | 0 | 1 |
| [16] Cerococcus artemisiae Last Modified in 12/10/14 | 10 | 164 | 0 | 0 | 0 | 0 | 0 | 1 |
| [17] Ceroplastes japonicus Last Modified in 12/05/13 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [18] Ceroputo pilosellae Last Modified in 12/10/14 | 4 | 170 | 0 | 0 | 0 | 0 | 0 | 1 |
| [19] Chionaspis salicis Last Modified in 12/10/14 | 0 | 174 | 0 | 0 | 0 | 0 | 0 | 1 |
| [20] Coccidohystrix insolitus Last Modified in 12/10/14 | 4 | 170 | 0 | 0 | 0 | 0 | 0 | 1 |
| [21] Coccus hesperidum Last Modified in 12/05/13 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [22] Coelostomidia pilosa Last Modified in 12/05/13 | 2 | 172 | 0 | 0 | 0 | 0 | 0 | 1 |
| [23] Coelostomidia wairoensis Last Modified in 12/05/13 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [24] Conchaspis lata Last Modified in 12/05/13 | 2 | 172 | 0 | 0 | 0 | 0 | 0 | 1 |
| [25] † Cretorthezia hammanaica Last Modified in 12/10/14 | 112 | 62 | 0 | 0 | 0 | 0 | 0 | 1 |
| [26] Dactylopius coccus Last Modified in 12/05/13 | 0 | 174 | 0 | 0 | 0 | 0 | 0 | 1 |
| [27] Dactylopius confusus Last Modified in 12/05/13 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [28] Dimargarodes meridionalis Last Modified in 12/05/13 | 4 | 170 | 0 | 0 | 0 | 0 | 0 | 1 |
| [29] Dimargarodes tanganyicius Last Modified in 12/05/13 | 3 | 171 | 0 | 0 | 0 | 0 | 0 | 1 |
| [30] Drosicha corpulenta Last Modified in 12/05/13 | 2 | 172 | 0 | 0 | 0 | 0 | 0 | 1 |
| [31] Drosicha dalbergiae Last Modified in 12/05/13 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [32] Drosicha pinicola Last Modified in 12/05/13 | 2 | 172 | 0 | 0 | 0 | 0 | 0 | 1 |
| [33] Dysmicoccus alazon Last Modified in 12/05/13 | 2 | 172 | 0 | 0 | 0 | 0 | 0 | 1 |
| [34] † Electrococcus canadensis Last Modified in 12/10/14 | 89 | 85 | 0 | 0 | 0 | 0 | 0 | 1 |
| [35] † Eomatsucoccus casei Last Modified in 12/10/14 | 110 | 64 | 0 | 0 | 0 | 0 | 0 | 1 |
| [36] Eriococcus buxi Last Modified in 12/05/13 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [37] Eriococcus coccineus Last Modified in 12/05/13 | 0 | 174 | 0 | 0 | 0 | 0 | 0 | 1 |
| [38] Eucallipterus tiliae Last Modified in 12/05/13 | 51 | 123 | 0 | 0 | 0 | 0 | 0 | 1 |
| [39] Eulecanium tiliae Last Modified in 12/05/13 | 2 | 172 | 0 | 0 | 0 | 0 | 0 | 1 |
| [40] Ferrisia virgata Last Modified in 12/05/13 | 2 | 172 | 0 | 0 | 0 | 0 | 0 | 1 |
| [41] Gallacoccus secundus Last Modified in 12/05/13 | 14 | 160 | 0 | 0 | 0 | 0 | 0 | 1 |
| [42] Gigantococcus maximus Last Modified in 12/05/13 | 2 | 172 | 0 | 0 | 0 | 0 | 0 | 1 |
| [43] † Grimaldiella gregaria Last Modified in 12/10/14 | 75 | 99 | 0 | 0 | 0 | 0 | 0 | 1 |
| [44] † Grohnus eichmanni Last Modified in 12/10/14 | 91 | 83 | 0 | 0 | 0 | 0 | 0 | 1 |
| [45] Heteromargarodes americanus Last Modified in 12/05/13 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [46] † Heteromargarodes hukamsinghi Last Modified in 12/10/14 | 101 | 73 | 0 | 0 | 0 | 0 | 0 | 1 |
| [47] † Hodgsonicoccus patefactus Last Modified in 12/10/14 | 85 | 89 | 0 | 0 | 0 | 0 | 0 | 1 |
| [48] Icerya purchasi Last Modified in 12/05/13 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [49] Icerya seychellarum Last Modified in 12/05/13 | 0 | 174 | 0 | 0 | 0 | 0 | 0 | 1 |
| [50] Inka minuta Last Modified in 12/05/13 | 74 | 100 | 0 | 0 | 0 | 0 | 0 | 1 |
| [51] Insignorthezia insignis Last Modified in 12/05/13 | 4 | 170 | 0 | 0 | 0 | 0 | 0 | 1 |
| [52] Jersicoccus kurthi Last Modified in 12/05/13 | 120 | 54 | 0 | 0 | 0 | 0 | 0 | 1 |
| [53] † Gilderius eukrinops Last Modified in 12/10/14 | 93 | 81 | 0 | 0 | 0 | 0 | 0 | 1 |
| [54] Kermes Last Modified in 12/05/13 | 56 | 118 | 0 | 0 | 0 | 0 | 0 | 1 |
| [55] Kermes quercus Last Modified in 12/05/13 | 5 | 169 | 0 | 0 | 0 | 0 | 0 | 1 |
| [56] † Kozarius achronus Last Modified in 12/10/14 | 79 | 95 | 0 | 0 | 0 | 0 | 0 | 1 |
| [57] † Kozarius perpetuus Last Modified in 12/10/14 | 78 | 96 | 0 | 0 | 0 | 0 | 0 | 1 |
| [58] † Kuenowicoccus pietrzeniukae Last Modified in 12/10/14 | 91 | 83 | 0 | 0 | 0 | 0 | 0 | 1 |
| [59] † Kukaspis usingeri Last Modified in 12/10/14 | 92 | 82 | 0 | 0 | 0 | 0 | 0 | 1 |
| [60] † Lebanococcus longiventris Last Modified in 12/10/14 | 96 | 78 | 0 | 0 | 0 | 0 | 0 | 1 |
| [61] Lecanodiaspis baculifera Last Modified in 12/05/13 | 11 | 163 | 0 | 0 | 0 | 0 | 0 | 1 |
| [62] † Magnilens glaesaria Last Modified in 12/10/14 | 102 | 72 | 0 | 0 | 0 | 0 | 0 | 1 |
| [63] Matsucoccus bisetosus Last Modified in 12/05/13 | 23 | 151 | 0 | 0 | 0 | 0 | 0 | 1 |
| [64] Matsucoccus feytaudi Last Modified in 12/05/13 | 3 | 171 | 0 | 0 | 0 | 0 | 0 | 1 |
| [65] Matsucoccus josephi Last Modified in 12/05/13 | 0 | 174 | 0 | 0 | 0 | 0 | 0 | 1 |
| [66] Matsucoccus matsumurae Last Modified in 12/05/13 | 5 | 169 | 0 | 0 | 0 | 0 | 0 | 1 |
| [67] † Marmyan barbarae Last Modified in 12/10/14 | 122 | 52 | 0 | 0 | 0 | 0 | 0 | 1 |
| [68] Neosteingelia texana Last Modified in 12/05/13 | 0 | 174 | 0 | 0 | 0 | 0 | 0 | 1 |
| [69] Newsteadia floccosa Last Modified in 12/05/13 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [70] † Normarkicoccus cambayae Last Modified in 12/10/14 | 107 | 67 | 0 | 0 | 0 | 0 | 0 | 1 |
| [71] Orthezia urticae Last Modified in 12/05/13 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [72] Ovaticoccus agavium Last Modified in 12/05/13 | 0 | 174 | 0 | 0 | 0 | 0 | 0 | 1 |
| [73] † Palaeosteingelia acrai Last Modified in 12/10/14 | 79 | 95 | 0 | 0 | 0 | 0 | 0 | 1 |
| [74] † Palaeotupo danieleae Last Modified in 12/10/14 | 95 | 79 | 0 | 0 | 0 | 0 | 0 | 1 |
| [75] Paleococcus Last Modified in 12/05/13 | 4 | 170 | 0 | 0 | 0 | 0 | 0 | 1 |
| [76] † Paleonewsteadia huaniae Last Modified in 12/10/14 | 84 | 90 | 0 | 0 | 0 | 0 | 0 | 1 |
| [77] Parlatoria oleae Last Modified in 12/05/13 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [78] † Pedicellicoccus marginatus Last Modified in 12/10/14 | 98 | 76 | 0 | 0 | 0 | 0 | 0 | 1 |
| [79] Pennygullania electrina Last Modified in 12/05/13 | 99 | 75 | 0 | 0 | 0 | 0 | 0 | 1 |
| [80] Phenacoleachia species a Last Modified in 12/05/13 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [81] Phenacoleachia species b Last Modified in 12/05/13 | 53 | 121 | 0 | 0 | 0 | 0 | 0 | 1 |
| [82] Pineus strobi Last Modified in 12/05/13 | 12 | 162 | 0 | 0 | 0 | 0 | 0 | 1 |
| [83] † Pityococcus moniliformalis Last Modified in 12/10/14 | 62 | 112 | 0 | 0 | 0 | 0 | 0 | 1 |
| [84] Pityococcus sp. Last Modified in 08/04/14 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [85] Planococcus citri Last Modified in 12/05/13 | 0 | 174 | 0 | 0 | 0 | 0 | 0 | 1 |
| [86] Porphyrophora hamelii Last Modified in 12/05/13 | 0 | 173 | 0 | 0 | 1 | 0 | 0 | 1 |
| [87] Praelongorthezia praelonga Last Modified in 12/05/13 | 5 | 169 | 0 | 0 | 0 | 0 | 0 | 1 |
| [88] † Protorthezia aurea Last Modified in 12/10/14 | 96 | 78 | 0 | 0 | 0 | 0 | 0 | 1 |
| [89] Pseudococcus longispinus Last Modified in 12/05/13 | 2 | 172 | 0 | 0 | 0 | 0 | 0 | 1 |
| [90] Pseudococcus maritimus Last Modified in 12/05/13 | 29 | 145 | 0 | 0 | 0 | 0 | 0 | 1 |
| [91] † Pseudoweitschatus audebertis Last Modified in 12/10/14 | 83 | 91 | 0 | 0 | 0 | 0 | 0 | 1 |
| [92] Puto kosztarabi Last Modified in 12/05/13 | 2 | 172 | 0 | 0 | 0 | 0 | 0 | 1 |
| [93] Puto mexicanus Last Modified in 12/05/13 | 5 | 169 | 0 | 0 | 0 | 0 | 0 | 1 |
| [94] Puto yuccae Last Modified in 12/05/13 | 0 | 174 | 0 | 0 | 0 | 0 | 0 | 1 |
| [95] Rhopalosiphum padi Last Modified in 12/05/13 | 7 | 165 | 0 | 0 | 2 | 0 | 0 | 1 |
| [96] † Rosahendersonia prisca Last Modified in 12/10/14 | 96 | 78 | 0 | 0 | 0 | 0 | 0 | 1 |
| [97] † Serafinus acutipterus Last Modified in 12/10/14 | 74 | 100 | 0 | 0 | 0 | 0 | 0 | 1 |
| [98] † Solicoccus nascimbenei Last Modified in 12/10/14 | 92 | 82 | 0 | 0 | 0 | 0 | 0 | 1 |
| [99] Steingelia cretacea Last Modified in 12/05/13 | 83 | 91 | 0 | 0 | 0 | 0 | 0 | 1 |
| [100] Steingelia gorodetskia Last Modified in 12/05/13 | 1 | 172 | 0 | 0 | 1 | 0 | 0 | 1 |
| [101] Stictococcus vayssievei Last Modified in 12/05/13 | 15 | 158 | 0 | 0 | 1 | 0 | 0 | 1 |
| [102] Stigmacoccus asper Last Modified in 12/05/13 | 2 | 172 | 0 | 0 | 0 | 0 | 0 | 1 |
| [103] Tachardina aurantiaca Last Modified in 12/05/13 | 0 | 174 | 0 | 0 | 0 | 0 | 0 | 1 |
| [104] Tanyscelis verrucula Last Modified in 08/04/14 | 16 | 158 | 0 | 0 | 0 | 0 | 0 | 1 |
| [105] † Turonicoccus beardsleyi Last Modified in 12/10/14 | 118 | 56 | 0 | 0 | 0 | 0 | 0 | 1 |
| [106] Ultracoelostoma assimile Last Modified in 12/05/13 | 3 | 171 | 0 | 0 | 0 | 0 | 0 | 1 |
| [107] † Weitschatus stigmatus Last Modified in 12/10/14 | 61 | 113 | 0 | 0 | 0 | 0 | 0 | 1 |
| [108] † Williamsicoccus megalops Last Modified in 12/10/14 | 98 | 76 | 0 | 0 | 0 | 0 | 0 | 1 |
| [109] † Xiphos vani Last Modified in 12/10/14 | 84 | 90 | 0 | 0 | 0 | 0 | 0 | 1 |
| [110] Xylococculus betulae Last Modified in 12/05/13 | 1 | 173 | 0 | 0 | 0 | 0 | 0 | 1 |
| [111] † Xylococcus granbenhorsti Last Modified in 12/10/14 | 82 | 92 | 0 | 0 | 0 | 0 | 0 | 1 |
| [112] Xylococcus japonicus Last Modified in 12/05/13 | 2 | 172 | 0 | 0 | 0 | 0 | 0 | 1 |
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