Project 3258: A. Takahashi, R. Hirayama, H. Otsuka. 2018. Systematic revision of Manouria oyamai (Testudines, Testudinidae), based on new material from the Upper Pleistocene of Okinawajima Island, the Ryukyu Archipelago, Japan, and its paleogeographic implications. Journal of Vertebrate Paleontology. 38 (2):e1427594.

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Original filename: Manouria-emys-asian-forest-tortoise-640x426-1.jpg

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Manouria emys

Manouria emys (unvouchered)

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whole animal

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