Project 5457: D. Geraads, C. Blondel, A. Likius, H. T. Mackaye, P. Vignaud, M. Brunet. 2008. New hippotragini (Bovidae, Mammalia) from the Late Miocene of Toros-Menalla (Chad). Journal of Vertebrate Paleontology. 28 (1):231-242.
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