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Carvalho, A. P. S., Orr, A. G. and Kawahara, A. Y. 2017. A review of the occurrence and diversity of the sphragis in butterflies (Lepidoptera, Papilionoidea). ZooKeys. Vol. 694, pp. 41-70.
Carvalho, Ana Paula S, Mota, Luísa L. and Kawahara, Akito Y. 2019. Intersexual ‘Arms Race’ and the Evolution of the Sphragis in Pteronymia Butterflies. Insect Systematics and Diversity. Vol. 3(1).