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Mattoni, C. I., Ochoa, J. A., Affilastro, A. A. O. and Prendini, L. 2012. Orobothriurus (Scorpiones: Bothriuridae) phylogeny, Andean biogeography, and the relative importance of genitalic and somatic characters. Zoologica Scripta. Vol. 41(2), p. 160–176.