Project 1013: I. Vea, D. A. Grimaldi. 2015. Isabelle M Vea Diverse New Scale Insects (Hemiptera: Coccoidea) in Amber from the Cretaceous and Eocene with a Phylogenetic Framework for Fossil Coccoidea. American Museum Novitates. 3823:1-80.
Specimen: † Rosahendersonia prisca (AMNH/AMNH Bu-835)
View: Light compound microscopy

Abstract

Coccoids are abundant and diverse in most amber deposits around the world, but largely as macropterous males. Based on a study of male coccoids in Lebanese amber (Early Cretaceous), Burmese amber (Albian-Cenomanian), Cambay amber from western India (Early Eocene), and Baltic amber (mid-Eocene), 16 new species, 11 new genera, and three new families are added to the coccoid fossil record: Apticoccidae, n. fam., based on Apticoccus Koteja and Azar, and including two new species A. fortis, n. sp., and A. longitenuis, n. sp.; the monotypic family Hodgsonicoccidae, n. fam., including Hodgsonicoccus patefactus, n. gen., n. sp.; Kozariidae, n. fam., including Kozarius achronus, n. gen., n. sp., and K. perpetuus, n. sp.; the first occurrence of a Coccidae in Burmese amber, Rosahendersonia prisca, n. gen., n. sp.; the first fossil record of a Margarodidae sensu stricto, Heteromargarodes hukamsinghi, n. sp.; a peculiar Diaspididae in Indian amber, Normarkicoccus cambayae, n. gen., n. sp.; a Pityococcidae from Baltic amber, Pityococcus moniliformalis, n. sp., two Pseudococcidae in Lebanese and Burmese ambers, Williamsicoccus megalops, n. gen., n. sp., and Gilderius eukrinops, n. gen., n. sp.; an Early Cretaceous Weitschatidae, Pseudoweitschatus audebertis, n. gen., n. sp.; four genera considered incertae sedis, Alacrena peculiaris, n. gen., n. sp., Magnilens glaesaria, n. gen., n. sp., and Pedicellicoccus marginatus, n. gen., n. sp., and Xiphos vani, n. gen., n. sp. Interpretation of fossil coccoids is supported by a parsimony phylogenetic analysis based on 174 morphological characters (both adult males and females) and 112 taxa (69 Recent and 43 extinct).


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Article DOI: 10.1206/3823.1

Project DOI: 10.7934/P1013, http://dx.doi.org/10.7934/P1013
This project containsMatrices
  • 31 Media
  • 1 Matrix
  • 143 Taxa
  • 19 Specimens
  • 343 Characters
Total size of project's media files: 31 MB

Download Project SDD File
Total scored cells: 14995
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 174
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 510
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:174/0
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MorphoBank Project 1013

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    Members

    member name taxa specimens media chars character
    media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
    medialabels
    rules
    Isabelle Vea
    Project Administrator
    Last logged in 12/05/16
    14319311740014995
    (14990, 0, 5)
    000


    Taxonomic Overview for Matrix 'M2800' (112 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Aclerda arundinariae
    Last Modified in 12/10/14
    2172000001
    [2] Acyrthosiphon pisum
    Last Modified in 12/05/13
    7167000001
    [3] Adelges tsugae
    Last Modified in 12/05/13
    53121000001
    [4] † Alacrena peculiaris
    Last Modified in 12/10/14
    10272000001
    [5] † Albicoccus dimai
    Last Modified in 12/10/14
    11262000001
    [6] Antonina graminis crawi
    Last Modified in 12/10/14
    1173000001
    [7] Aonidiella aurantii
    Last Modified in 12/10/14
    0174000001
    [8] † Apticoccus fortis
    Last Modified in 12/10/14
    9579000001
    [9] † Apticoccus longitenuis
    Last Modified in 12/10/14
    9876000001
    [10] † Apticoccus minutus
    Last Modified in 12/10/14
    9480000001
    [11] † Undescribed undescribed
    Last Modified in 12/10/14
    12549000001
    [12] † Arnoldus capitatus
    Last Modified in 12/10/14
    9876000001
    [13] Beesonia dipterocarpi
    Last Modified in 12/10/14
    12162000001
    [14] † Burmacoccus danyi
    Last Modified in 12/10/14
    10668000001
    [15] Callipappus
    Last Modified in 12/10/14
    2172000001
    [16] Cerococcus artemisiae
    Last Modified in 12/10/14
    10164000001
    [17] Ceroplastes japonicus
    Last Modified in 12/05/13
    1173000001
    [18] Ceroputo pilosellae
    Last Modified in 12/10/14
    4170000001
    [19] Chionaspis salicis
    Last Modified in 12/10/14
    0174000001
    [20] Coccidohystrix insolitus
    Last Modified in 12/10/14
    4170000001
    [21] Coccus hesperidum
    Last Modified in 12/05/13
    1173000001
    [22] Coelostomidia pilosa
    Last Modified in 12/05/13
    2172000001
    [23] Coelostomidia wairoensis
    Last Modified in 12/05/13
    1173000001
    [24] Conchaspis lata
    Last Modified in 12/05/13
    2172000001
    [25] † Cretorthezia hammanaica
    Last Modified in 12/10/14
    11262000001
    [26] Dactylopius coccus
    Last Modified in 12/05/13
    0174000001
    [27] Dactylopius confusus
    Last Modified in 12/05/13
    1173000001
    [28] Dimargarodes meridionalis
    Last Modified in 12/05/13
    4170000001
    [29] Dimargarodes tanganyicius
    Last Modified in 12/05/13
    3171000001
    [30] Drosicha corpulenta
    Last Modified in 12/05/13
    2172000001
    [31] Drosicha dalbergiae
    Last Modified in 12/05/13
    1173000001
    [32] Drosicha pinicola
    Last Modified in 12/05/13
    2172000001
    [33] Dysmicoccus alazon
    Last Modified in 12/05/13
    2172000001
    [34] † Electrococcus canadensis
    Last Modified in 12/10/14
    8985000001
    [35] † Eomatsucoccus casei
    Last Modified in 12/10/14
    11064000001
    [36] Eriococcus buxi
    Last Modified in 12/05/13
    1173000001
    [37] Eriococcus coccineus
    Last Modified in 12/05/13
    0174000001
    [38] Eucallipterus tiliae
    Last Modified in 12/05/13
    51123000001
    [39] Eulecanium tiliae
    Last Modified in 12/05/13
    2172000001
    [40] Ferrisia virgata
    Last Modified in 12/05/13
    2172000001
    [41] Gallacoccus secundus
    Last Modified in 12/05/13
    14160000001
    [42] Gigantococcus maximus
    Last Modified in 12/05/13
    2172000001
    [43] † Grimaldiella gregaria
    Last Modified in 12/10/14
    7599000001
    [44] † Grohnus eichmanni
    Last Modified in 12/10/14
    9183000001
    [45] Heteromargarodes americanus
    Last Modified in 12/05/13
    1173000001
    [46] † Heteromargarodes hukamsinghi
    Last Modified in 12/10/14
    10173000001
    [47] † Hodgsonicoccus patefactus
    Last Modified in 12/10/14
    8589000001
    [48] Icerya purchasi
    Last Modified in 12/05/13
    1173000001
    [49] Icerya seychellarum
    Last Modified in 12/05/13
    0174000001
    [50] Inka minuta
    Last Modified in 12/05/13
    74100000001
    [51] Insignorthezia insignis
    Last Modified in 12/05/13
    4170000001
    [52] Jersicoccus kurthi
    Last Modified in 12/05/13
    12054000001
    [53] † Gilderius eukrinops
    Last Modified in 12/10/14
    9381000001
    [54] Kermes
    Last Modified in 12/05/13
    56118000001
    [55] Kermes quercus
    Last Modified in 12/05/13
    5169000001
    [56] † Kozarius achronus
    Last Modified in 12/10/14
    7995000001
    [57] † Kozarius perpetuus
    Last Modified in 12/10/14
    7896000001
    [58] † Kuenowicoccus pietrzeniukae
    Last Modified in 12/10/14
    9183000001
    [59] † Kukaspis usingeri
    Last Modified in 12/10/14
    9282000001
    [60] † Lebanococcus longiventris
    Last Modified in 12/10/14
    9678000001
    [61] Lecanodiaspis baculifera
    Last Modified in 12/05/13
    11163000001
    [62] † Magnilens glaesaria
    Last Modified in 12/10/14
    10272000001
    [63] Matsucoccus bisetosus
    Last Modified in 12/05/13
    23151000001
    [64] Matsucoccus feytaudi
    Last Modified in 12/05/13
    3171000001
    [65] Matsucoccus josephi
    Last Modified in 12/05/13
    0174000001
    [66] Matsucoccus matsumurae
    Last Modified in 12/05/13
    5169000001
    [67] † Marmyan barbarae
    Last Modified in 12/10/14
    12252000001
    [68] Neosteingelia texana
    Last Modified in 12/05/13
    0174000001
    [69] Newsteadia floccosa
    Last Modified in 12/05/13
    1173000001
    [70] † Normarkicoccus cambayae
    Last Modified in 12/10/14
    10767000001
    [71] Orthezia urticae
    Last Modified in 12/05/13
    1173000001
    [72] Ovaticoccus agavium
    Last Modified in 12/05/13
    0174000001
    [73] † Palaeosteingelia acrai
    Last Modified in 12/10/14
    7995000001
    [74] † Palaeotupo danieleae
    Last Modified in 12/10/14
    9579000001
    [75] Paleococcus
    Last Modified in 12/05/13
    4170000001
    [76] † Paleonewsteadia huaniae
    Last Modified in 12/10/14
    8490000001
    [77] Parlatoria oleae
    Last Modified in 12/05/13
    1173000001
    [78] † Pedicellicoccus marginatus
    Last Modified in 12/10/14
    9876000001
    [79] Pennygullania electrina
    Last Modified in 12/05/13
    9975000001
    [80] Phenacoleachia species a
    Last Modified in 12/05/13
    1173000001
    [81] Phenacoleachia species b
    Last Modified in 12/05/13
    53121000001
    [82] Pineus strobi
    Last Modified in 12/05/13
    12162000001
    [83] † Pityococcus moniliformalis
    Last Modified in 12/10/14
    62112000001
    [84] Pityococcus sp.
    Last Modified in 08/04/14
    1173000001
    [85] Planococcus citri
    Last Modified in 12/05/13
    0174000001
    [86] Porphyrophora hamelii
    Last Modified in 12/05/13
    0173001001
    [87] Praelongorthezia praelonga
    Last Modified in 12/05/13
    5169000001
    [88] † Protorthezia aurea
    Last Modified in 12/10/14
    9678000001
    [89] Pseudococcus longispinus
    Last Modified in 12/05/13
    2172000001
    [90] Pseudococcus maritimus
    Last Modified in 12/05/13
    29145000001
    [91] † Pseudoweitschatus audebertis
    Last Modified in 12/10/14
    8391000001
    [92] Puto kosztarabi
    Last Modified in 12/05/13
    2172000001
    [93] Puto mexicanus
    Last Modified in 12/05/13
    5169000001
    [94] Puto yuccae
    Last Modified in 12/05/13
    0174000001
    [95] Rhopalosiphum padi
    Last Modified in 12/05/13
    7165002001
    [96] † Rosahendersonia prisca
    Last Modified in 12/10/14
    9678000001
    [97] † Serafinus acutipterus
    Last Modified in 12/10/14
    74100000001
    [98] † Solicoccus nascimbenei
    Last Modified in 12/10/14
    9282000001
    [99] Steingelia cretacea
    Last Modified in 12/05/13
    8391000001
    [100] Steingelia gorodetskia
    Last Modified in 12/05/13
    1172001001
    [101] Stictococcus vayssievei
    Last Modified in 12/05/13
    15158001001
    [102] Stigmacoccus asper
    Last Modified in 12/05/13
    2172000001
    [103] Tachardina aurantiaca
    Last Modified in 12/05/13
    0174000001
    [104] Tanyscelis verrucula
    Last Modified in 08/04/14
    16158000001
    [105] † Turonicoccus beardsleyi
    Last Modified in 12/10/14
    11856000001
    [106] Ultracoelostoma assimile
    Last Modified in 12/05/13
    3171000001
    [107] † Weitschatus stigmatus
    Last Modified in 12/10/14
    61113000001
    [108] † Williamsicoccus megalops
    Last Modified in 12/10/14
    9876000001
    [109] † Xiphos vani
    Last Modified in 12/10/14
    8490000001
    [110] Xylococculus betulae
    Last Modified in 12/05/13
    1173000001
    [111] † Xylococcus granbenhorsti
    Last Modified in 12/10/14
    8292000001
    [112] Xylococcus japonicus
    Last Modified in 12/05/13
    2172000001


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