Project 1037: C. R. Congreve, A. Z. Krug, M. E. Patzkowsky. 2015. Phylogenetic revision of the Strophomenida, a diverse and ecologically important Palaeozoic brachiopod order. Palaeontology. 58 (4):743-758.
Abstract
The order Strophomenida was an ecologically abundant and taxonomically diverse group of Palaeozoic brachiopods that originated in the earliest Ordovician and went extinct in the Carboniferous. During their long geologic range, the Strophomenida survived two of the “Big Five” mass extinction events, the Late Ordovician and the Late Devonian, suggesting that they are potentially informative taxa for studying the evolutionary effects of these two distinct mass extinctions, each with drastically different forcing mechanisms. However, while there have been previous phylogenetic studies on smaller groups within the Strophomenida, the phylogenetic relationships of the whole group are still largely unknown. The group has been divided into two major superfamilies, the Strophomenoidea (strophomenoids) and the Plectambonitoidea (plectambonitoids). Despite being treated as separate clades, the plectambonitoids may form a paraphyletic grade into the strophomenoids. We present a detailed higher level parsimony based phylogenetic analysis of the Strophomenida, consisting of sixty-nine characters and sixty-two exemplar species sampled from the majority of the taxonomically defined families/subfamilies. Several species of basal chonetids (strophochonetids) were also included in this analysis, since they may be closely related to the Strophomenida and share several characters with both the plectambonitoids and strophomenoids. The phylogenetic analysis suggests the plectambonitoids, as originally defined, are paraphyletic to the monophyletic strophomenoids. The basal chonetids are reconstructed as a monophyletic group that is sister to the strophomenoids, suggesting that their proper placement might be within the Strophomenida. The topology also suggests that at least seventeen of the taxonomically defined strophomenoid and plectambonitoid families are likely monophyletic. The Plectambonitidae and the Taffiidae as defined are paraphyletic, and the Grorudiidae and Leptostrophiidae are polyphyletic. Furthermore, subfamilies Leptodontellinae, Dicoelostrophiinae, Palaeostrophomeninae, and Aegiromeninae are raised to the level of the family. When analysed within this phylogenetic context, the Late Ordovician mass extinction event had little effect on the large-scale evolution of the group.Read the article »
Article DOI: 10.1111/pala.12177
Project DOI: 10.7934/P1037, http://dx.doi.org/10.7934/P1037
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 11251 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 209 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 156 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:209/0 |
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MorphoBank Project 1037
MorphoBank Project 1037
- Creation Date:
08 October 2013 - Publication Date:
27 July 2015 - Project views: 33963
- Matrix downloads: 17
Authors' Institutions
- Pennsylvania State University
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Curtis Congreve Project Administrator | 65 | 1 | 1 | 209 | 0 | 0 | 11251 (9646, 0, 1605) | 0 | 0 | 57 | ||||
Andrew Krug Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Mark Patzkowsky Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Judi Sclafani Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M23282' (62 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Billingsella perfecta Last Modified in 07/27/15 | 3 | 53 | 53 | 0 | 15 | 0 | 0 | 4 |
[2] † Strophomena planumbona Last Modified in 07/27/15 | 4 | 60 | 60 | 0 | 7 | 0 | 0 | 4 |
[3] † Actinomena orta Last Modified in 07/27/15 | 3 | 65 | 65 | 0 | 3 | 0 | 0 | 4 |
[4] † Furcitella plicata Last Modified in 07/27/15 | 21 | 44 | 44 | 0 | 6 | 0 | 0 | 4 |
[5] † Bekkerina dorsata Last Modified in 07/27/15 | 5 | 61 | 61 | 0 | 5 | 0 | 0 | 4 |
[6] † Oepikina septata Last Modified in 07/27/15 | 3 | 61 | 61 | 0 | 7 | 0 | 0 | 4 |
[7] † Colaptomena leptostrophoidea Last Modified in 07/27/15 | 6 | 53 | 53 | 0 | 12 | 0 | 0 | 4 |
[8] † Kjerulfina trigonalis Last Modified in 07/27/15 | 10 | 49 | 49 | 0 | 12 | 0 | 0 | 4 |
[9] † Rafinesquina alternata Last Modified in 07/27/15 | 5 | 57 | 57 | 0 | 9 | 0 | 0 | 4 |
[10] † Kjaerina poljensis Last Modified in 07/27/15 | 10 | 54 | 54 | 0 | 7 | 0 | 0 | 4 |
[11] † Leptaena richmondensis Last Modified in 07/27/15 | 6 | 59 | 59 | 0 | 6 | 0 | 0 | 4 |
[12] † Kiaeromena estonensis Last Modified in 07/27/15 | 15 | 48 | 48 | 0 | 8 | 0 | 0 | 4 |
[13] † Glyptomena sculpturata Last Modified in 07/27/15 | 9 | 51 | 51 | 0 | 11 | 0 | 0 | 4 |
[14] † Platymena plana Last Modified in 07/27/15 | 7 | 58 | 58 | 0 | 6 | 0 | 0 | 4 |
[15] † Christiania subquadrata Last Modified in 07/27/15 | 28 | 37 | 37 | 0 | 6 | 0 | 0 | 4 |
[16] † Leptaenoidea rugata Last Modified in 07/27/15 | 6 | 59 | 59 | 0 | 6 | 0 | 0 | 4 |
[17] † Leptaenisca concava Last Modified in 07/27/15 | 7 | 56 | 56 | 0 | 8 | 0 | 0 | 4 |
[18] † Amphistrophia striata Last Modified in 07/27/15 | 7 | 58 | 58 | 0 | 6 | 0 | 0 | 4 |
[19] † Mesodouvillina subinterstrialis Last Modified in 07/27/15 | 7 | 58 | 58 | 0 | 6 | 0 | 0 | 4 |
[20] † Maoristrophia neozelanica Last Modified in 07/27/15 | 18 | 47 | 47 | 0 | 6 | 0 | 0 | 4 |
[21] † Douvillina dutertrei Last Modified in 07/27/15 | 7 | 53 | 53 | 0 | 11 | 0 | 0 | 4 |
[22] † Protodouvillina inaequistriata Last Modified in 07/27/15 | 16 | 49 | 49 | 0 | 6 | 0 | 0 | 4 |
[23] † Douvillinella filifer Last Modified in 07/27/15 | 6 | 59 | 59 | 0 | 6 | 0 | 0 | 4 |
[24] † Dicoelostrophia punctatum Last Modified in 07/27/15 | 13 | 52 | 52 | 0 | 6 | 0 | 0 | 4 |
[25] † Leptodontella caudata Last Modified in 07/27/15 | 14 | 50 | 50 | 0 | 7 | 0 | 0 | 4 |
[26] † Leptostrophia magniventra Last Modified in 07/27/15 | 11 | 52 | 52 | 0 | 8 | 0 | 0 | 4 |
[27] † Leptostrophia beckii Last Modified in 07/27/15 | 13 | 51 | 51 | 0 | 7 | 0 | 0 | 4 |
[28] † Brachyprion philomela Last Modified in 07/27/15 | 11 | 52 | 52 | 0 | 8 | 0 | 0 | 4 |
[29] † Brachyprion semiglobosa Last Modified in 07/27/15 | 6 | 59 | 59 | 0 | 6 | 0 | 0 | 4 |
[30] † Strophodonta demissa Last Modified in 07/27/15 | 7 | 57 | 57 | 0 | 7 | 0 | 0 | 4 |
[31] † Parapholidostrophia harperi Last Modified in 07/27/15 | 7 | 57 | 57 | 0 | 7 | 0 | 0 | 4 |
[32] † Shaleria ornatella Last Modified in 07/27/15 | 11 | 51 | 51 | 0 | 9 | 0 | 0 | 4 |
[33] † Eostrophonella eothen Last Modified in 07/27/15 | 16 | 42 | 42 | 0 | 13 | 0 | 0 | 4 |
[34] † Plectambonites planissimus Last Modified in 07/27/15 | 7 | 50 | 50 | 0 | 14 | 0 | 0 | 4 |
[35] † Isophragma ricevillense Last Modified in 07/27/15 | 2 | 53 | 53 | 0 | 16 | 0 | 0 | 4 |
[36] † Tafia planoconvexa Last Modified in 07/27/15 | 12 | 44 | 44 | 0 | 15 | 0 | 0 | 4 |
[37] † Ahtiella lirata Last Modified in 07/27/15 | 10 | 47 | 47 | 0 | 14 | 0 | 0 | 4 |
[38] † Leptella nevadensis Last Modified in 07/27/15 | 17 | 34 | 34 | 0 | 20 | 0 | 0 | 4 |
[39] † Bimuria superba Last Modified in 07/27/15 | 6 | 54 | 54 | 0 | 11 | 0 | 0 | 4 |
[40] † Sowerbyites hami Last Modified in 07/27/15 | 1 | 58 | 58 | 0 | 12 | 0 | 0 | 4 |
[41] † Leptellina? pulchra Last Modified in 07/27/15 | 10 | 54 | 54 | 0 | 7 | 0 | 0 | 4 |
[42] † Leptelloidea leptelloides Last Modified in 07/27/15 | 0 | 58 | 58 | 0 | 13 | 0 | 0 | 4 |
[43] † Palaeostrophomena concava Last Modified in 07/27/15 | 1 | 63 | 62 | 0 | 8 | 0 | 0 | 4 |
[44] † Alwynella ildjemensis Last Modified in 07/27/15 | 1 | 62 | 62 | 0 | 8 | 0 | 0 | 4 |
[45] † Railtonella scanloni Last Modified in 07/27/15 | 13 | 47 | 47 | 0 | 11 | 0 | 0 | 4 |
[46] † Leptestia musculosa Last Modified in 07/27/15 | 7 | 55 | 55 | 0 | 9 | 0 | 0 | 4 |
[47] † Sampo hiiuensis Last Modified in 07/27/15 | 3 | 62 | 62 | 0 | 6 | 0 | 0 | 4 |
[48] † Xenambonites undosus Last Modified in 07/27/15 | 4 | 58 | 58 | 0 | 9 | 0 | 0 | 4 |
[49] † Metambonites meritus Last Modified in 07/27/15 | 24 | 42 | 42 | 0 | 5 | 0 | 0 | 4 |
[50] † Aegiromena aquila Last Modified in 07/27/15 | 14 | 47 | 47 | 0 | 10 | 0 | 0 | 4 |
[51] † Epelidoaegiria chilidifera Last Modified in 07/27/15 | 13 | 49 | 49 | 0 | 9 | 0 | 0 | 4 |
[52] † Aegiria norvegica Last Modified in 07/27/15 | 7 | 54 | 54 | 0 | 10 | 0 | 0 | 4 |
[53] † Hesperomena leptellinoidea Last Modified in 07/27/15 | 24 | 42 | 42 | 0 | 5 | 0 | 0 | 4 |
[54] † Anoptambonites grayae Last Modified in 07/27/15 | 6 | 58 | 58 | 0 | 7 | 0 | 0 | 4 |
[55] † Sowerbyella punctostriata Last Modified in 07/27/15 | 7 | 61 | 61 | 0 | 3 | 0 | 0 | 4 |
[56] † Eochonetes clarksvillensis Last Modified in 07/27/15 | 16 | 49 | 49 | 0 | 6 | 0 | 0 | 4 |
[57] † Plectodonta mariae Last Modified in 07/27/15 | 21 | 45 | 45 | 0 | 5 | 0 | 0 | 4 |
[58] † Ptychoglyptus virginiensis Last Modified in 07/27/15 | 9 | 55 | 55 | 0 | 7 | 0 | 0 | 4 |
[59] † Strophochonetes cingulatus Last Modified in 06/03/15 | 25 | 38 | 38 | 0 | 8 | 0 | 0 | 4 |
[60] † Borealinetes comestus Last Modified in 07/27/15 | 7 | 55 | 55 | 0 | 9 | 0 | 0 | 4 |
[61] † Ctenochonetes tenuicostata Last Modified in 07/27/15 | 2 | 57 | 57 | 0 | 12 | 0 | 0 | 4 |
[62] † Archaeochonetes primigenius Last Modified in 07/27/15 | 14 | 50 | 50 | 0 | 7 | 0 | 0 | 4 |
Taxonomic Overview for Matrix 'M23367' (62 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Billingsella perfecta Last Modified in 07/27/15 | 3 | 51 | 51 | 0 | 15 | 0 | 0 | 4 |
[2] † Strophomena planumbona Last Modified in 07/27/15 | 4 | 58 | 58 | 0 | 7 | 0 | 0 | 4 |
[3] † Actinomena orta Last Modified in 07/27/15 | 3 | 63 | 63 | 0 | 3 | 0 | 0 | 4 |
[4] † Furcitella plicata Last Modified in 07/27/15 | 21 | 42 | 42 | 0 | 6 | 0 | 0 | 4 |
[5] † Bekkerina dorsata Last Modified in 07/27/15 | 5 | 59 | 59 | 0 | 5 | 0 | 0 | 4 |
[6] † Oepikina septata Last Modified in 07/27/15 | 3 | 59 | 59 | 0 | 7 | 0 | 0 | 4 |
[7] † Colaptomena leptostrophoidea Last Modified in 07/27/15 | 6 | 51 | 51 | 0 | 12 | 0 | 0 | 4 |
[8] † Kjerulfina trigonalis Last Modified in 07/27/15 | 10 | 47 | 47 | 0 | 12 | 0 | 0 | 4 |
[9] † Rafinesquina alternata Last Modified in 07/27/15 | 5 | 55 | 55 | 0 | 9 | 0 | 0 | 4 |
[10] † Kjaerina poljensis Last Modified in 07/27/15 | 10 | 52 | 52 | 0 | 7 | 0 | 0 | 4 |
[11] † Leptaena richmondensis Last Modified in 07/27/15 | 6 | 57 | 57 | 0 | 6 | 0 | 0 | 4 |
[12] † Kiaeromena estonensis Last Modified in 07/27/15 | 15 | 46 | 46 | 0 | 8 | 0 | 0 | 4 |
[13] † Glyptomena sculpturata Last Modified in 07/27/15 | 9 | 49 | 49 | 0 | 11 | 0 | 0 | 4 |
[14] † Platymena plana Last Modified in 07/27/15 | 7 | 56 | 56 | 0 | 6 | 0 | 0 | 4 |
[15] † Christiania subquadrata Last Modified in 07/27/15 | 26 | 37 | 37 | 0 | 6 | 0 | 0 | 4 |
[16] † Leptaenoidea rugata Last Modified in 07/27/15 | 6 | 57 | 57 | 0 | 6 | 0 | 0 | 4 |
[17] † Leptaenisca concava Last Modified in 07/27/15 | 7 | 54 | 54 | 0 | 8 | 0 | 0 | 4 |
[18] † Amphistrophia striata Last Modified in 07/27/15 | 7 | 56 | 56 | 0 | 6 | 0 | 0 | 4 |
[19] † Mesodouvillina subinterstrialis Last Modified in 07/27/15 | 7 | 56 | 56 | 0 | 6 | 0 | 0 | 4 |
[20] † Maoristrophia neozelanica Last Modified in 07/27/15 | 18 | 45 | 45 | 0 | 6 | 0 | 0 | 4 |
[21] † Douvillina dutertrei Last Modified in 07/27/15 | 7 | 51 | 51 | 0 | 11 | 0 | 0 | 4 |
[22] † Protodouvillina inaequistriata Last Modified in 07/27/15 | 16 | 47 | 47 | 0 | 6 | 0 | 0 | 4 |
[23] † Douvillinella filifer Last Modified in 07/27/15 | 6 | 57 | 57 | 0 | 6 | 0 | 0 | 4 |
[24] † Dicoelostrophia punctatum Last Modified in 07/27/15 | 13 | 50 | 50 | 0 | 6 | 0 | 0 | 4 |
[25] † Leptodontella caudata Last Modified in 07/27/15 | 14 | 48 | 48 | 0 | 7 | 0 | 0 | 4 |
[26] † Leptostrophia magniventra Last Modified in 07/27/15 | 11 | 50 | 50 | 0 | 8 | 0 | 0 | 4 |
[27] † Leptostrophia beckii Last Modified in 07/27/15 | 13 | 49 | 49 | 0 | 7 | 0 | 0 | 4 |
[28] † Brachyprion philomela Last Modified in 07/27/15 | 11 | 50 | 50 | 0 | 8 | 0 | 0 | 4 |
[29] † Brachyprion semiglobosa Last Modified in 07/27/15 | 6 | 57 | 57 | 0 | 6 | 0 | 0 | 4 |
[30] † Strophodonta demissa Last Modified in 07/27/15 | 7 | 55 | 55 | 0 | 7 | 0 | 0 | 4 |
[31] † Parapholidostrophia harperi Last Modified in 07/27/15 | 7 | 55 | 55 | 0 | 7 | 0 | 0 | 4 |
[32] † Shaleria ornatella Last Modified in 07/27/15 | 11 | 49 | 49 | 0 | 9 | 0 | 0 | 4 |
[33] † Eostrophonella eothen Last Modified in 07/27/15 | 16 | 40 | 40 | 0 | 13 | 0 | 0 | 4 |
[34] † Plectambonites planissimus Last Modified in 07/27/15 | 7 | 48 | 48 | 0 | 14 | 0 | 0 | 4 |
[35] † Isophragma ricevillense Last Modified in 07/27/15 | 2 | 51 | 51 | 0 | 16 | 0 | 0 | 4 |
[36] † Tafia planoconvexa Last Modified in 07/27/15 | 12 | 42 | 42 | 0 | 15 | 0 | 0 | 4 |
[37] † Ahtiella lirata Last Modified in 07/27/15 | 10 | 45 | 45 | 0 | 14 | 0 | 0 | 4 |
[38] † Leptella nevadensis Last Modified in 07/27/15 | 17 | 32 | 32 | 0 | 20 | 0 | 0 | 4 |
[39] † Bimuria superba Last Modified in 07/27/15 | 6 | 52 | 52 | 0 | 11 | 0 | 0 | 4 |
[40] † Sowerbyites hami Last Modified in 07/27/15 | 1 | 56 | 56 | 0 | 12 | 0 | 0 | 4 |
[41] † Leptellina? pulchra Last Modified in 07/27/15 | 10 | 52 | 52 | 0 | 7 | 0 | 0 | 4 |
[42] † Leptelloidea leptelloides Last Modified in 07/27/15 | 0 | 56 | 56 | 0 | 13 | 0 | 0 | 4 |
[43] † Palaeostrophomena concava Last Modified in 07/27/15 | 1 | 61 | 60 | 0 | 8 | 0 | 0 | 4 |
[44] † Alwynella ildjemensis Last Modified in 07/27/15 | 1 | 60 | 60 | 0 | 8 | 0 | 0 | 4 |
[45] † Railtonella scanloni Last Modified in 07/27/15 | 13 | 45 | 45 | 0 | 11 | 0 | 0 | 4 |
[46] † Leptestia musculosa Last Modified in 07/27/15 | 7 | 53 | 53 | 0 | 9 | 0 | 0 | 4 |
[47] † Sampo hiiuensis Last Modified in 07/27/15 | 3 | 60 | 60 | 0 | 6 | 0 | 0 | 4 |
[48] † Xenambonites undosus Last Modified in 07/27/15 | 4 | 56 | 56 | 0 | 9 | 0 | 0 | 4 |
[49] † Metambonites meritus Last Modified in 07/27/15 | 24 | 40 | 40 | 0 | 5 | 0 | 0 | 4 |
[50] † Aegiromena aquila Last Modified in 07/27/15 | 14 | 45 | 45 | 0 | 10 | 0 | 0 | 4 |
[51] † Epelidoaegiria chilidifera Last Modified in 07/27/15 | 13 | 47 | 47 | 0 | 9 | 0 | 0 | 4 |
[52] † Aegiria norvegica Last Modified in 07/27/15 | 7 | 52 | 52 | 0 | 10 | 0 | 0 | 4 |
[53] † Hesperomena leptellinoidea Last Modified in 07/27/15 | 24 | 40 | 40 | 0 | 5 | 0 | 0 | 4 |
[54] † Anoptambonites grayae Last Modified in 07/27/15 | 6 | 56 | 56 | 0 | 7 | 0 | 0 | 4 |
[55] † Sowerbyella punctostriata Last Modified in 07/27/15 | 7 | 59 | 59 | 0 | 3 | 0 | 0 | 4 |
[56] † Eochonetes clarksvillensis Last Modified in 07/27/15 | 16 | 47 | 47 | 0 | 6 | 0 | 0 | 4 |
[57] † Plectodonta mariae Last Modified in 07/27/15 | 21 | 43 | 43 | 0 | 5 | 0 | 0 | 4 |
[58] † Ptychoglyptus virginiensis Last Modified in 07/27/15 | 9 | 53 | 53 | 0 | 7 | 0 | 0 | 4 |
[59] † Strophochonetes cingulatus Last Modified in 06/03/15 | 25 | 36 | 36 | 0 | 8 | 0 | 0 | 4 |
[60] † Borealinetes comestus Last Modified in 07/27/15 | 7 | 53 | 53 | 0 | 9 | 0 | 0 | 4 |
[61] † Ctenochonetes tenuicostata Last Modified in 07/27/15 | 2 | 55 | 55 | 0 | 12 | 0 | 0 | 4 |
[62] † Archaeochonetes primigenius Last Modified in 07/27/15 | 14 | 48 | 48 | 0 | 7 | 0 | 0 | 4 |
Taxonomic Overview for Matrix 'M23368' (64 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Billingsella perfecta Last Modified in 07/27/15 | 3 | 51 | 51 | 0 | 15 | 0 | 0 | 4 |
[2] † Strophomena planumbona Last Modified in 07/27/15 | 4 | 58 | 58 | 0 | 7 | 0 | 0 | 4 |
[3] † Actinomena orta Last Modified in 07/27/15 | 3 | 63 | 63 | 0 | 3 | 0 | 0 | 4 |
[4] † Furcitella plicata Last Modified in 07/27/15 | 21 | 42 | 42 | 0 | 6 | 0 | 0 | 4 |
[5] † Bekkerina dorsata Last Modified in 07/27/15 | 5 | 59 | 59 | 0 | 5 | 0 | 0 | 4 |
[6] † Oepikina septata Last Modified in 07/27/15 | 3 | 59 | 59 | 0 | 7 | 0 | 0 | 4 |
[7] † Foliomena folium Last Modified in 07/27/15 | 22 | 37 | 37 | 0 | 10 | 0 | 0 | 4 |
[8] † Colaptomena leptostrophoidea Last Modified in 07/27/15 | 6 | 51 | 51 | 0 | 12 | 0 | 0 | 4 |
[9] † Kjerulfina trigonalis Last Modified in 07/27/15 | 10 | 47 | 47 | 0 | 12 | 0 | 0 | 4 |
[10] † Rafinesquina alternata Last Modified in 07/27/15 | 5 | 55 | 55 | 0 | 9 | 0 | 0 | 4 |
[11] † Kjaerina poljensis Last Modified in 07/27/15 | 10 | 52 | 52 | 0 | 7 | 0 | 0 | 4 |
[12] † Leptaena richmondensis Last Modified in 07/27/15 | 6 | 57 | 57 | 0 | 6 | 0 | 0 | 4 |
[13] † Kiaeromena estonensis Last Modified in 07/27/15 | 15 | 46 | 46 | 0 | 8 | 0 | 0 | 4 |
[14] † Glyptomena sculpturata Last Modified in 07/27/15 | 9 | 49 | 49 | 0 | 11 | 0 | 0 | 4 |
[15] † Platymena plana Last Modified in 07/27/15 | 7 | 56 | 56 | 0 | 6 | 0 | 0 | 4 |
[16] † Christiania subquadrata Last Modified in 07/27/15 | 26 | 37 | 37 | 0 | 6 | 0 | 0 | 4 |
[17] † Leptaenoidea rugata Last Modified in 07/27/15 | 6 | 57 | 57 | 0 | 6 | 0 | 0 | 4 |
[18] † Leptaenisca concava Last Modified in 07/27/15 | 7 | 54 | 54 | 0 | 8 | 0 | 0 | 4 |
[19] † Amphistrophia striata Last Modified in 07/27/15 | 7 | 56 | 56 | 0 | 6 | 0 | 0 | 4 |
[20] † Mesodouvillina subinterstrialis Last Modified in 07/27/15 | 7 | 56 | 56 | 0 | 6 | 0 | 0 | 4 |
[21] † Maoristrophia neozelanica Last Modified in 07/27/15 | 18 | 45 | 45 | 0 | 6 | 0 | 0 | 4 |
[22] † Douvillina dutertrei Last Modified in 07/27/15 | 7 | 51 | 51 | 0 | 11 | 0 | 0 | 4 |
[23] † Protodouvillina inaequistriata Last Modified in 07/27/15 | 16 | 47 | 47 | 0 | 6 | 0 | 0 | 4 |
[24] † Douvillinella filifer Last Modified in 07/27/15 | 6 | 57 | 57 | 0 | 6 | 0 | 0 | 4 |
[25] † Dicoelostrophia punctatum Last Modified in 07/27/15 | 13 | 50 | 50 | 0 | 6 | 0 | 0 | 4 |
[26] † Leptodontella caudata Last Modified in 07/27/15 | 14 | 48 | 48 | 0 | 7 | 0 | 0 | 4 |
[27] † Leptostrophia magniventra Last Modified in 07/27/15 | 11 | 50 | 50 | 0 | 8 | 0 | 0 | 4 |
[28] † Leptostrophia beckii Last Modified in 07/27/15 | 13 | 49 | 49 | 0 | 7 | 0 | 0 | 4 |
[29] † Brachyprion philomela Last Modified in 07/27/15 | 11 | 50 | 50 | 0 | 8 | 0 | 0 | 4 |
[30] † Brachyprion semiglobosa Last Modified in 07/27/15 | 6 | 57 | 57 | 0 | 6 | 0 | 0 | 4 |
[31] † Strophodonta demissa Last Modified in 07/27/15 | 7 | 55 | 55 | 0 | 7 | 0 | 0 | 4 |
[32] † Parapholidostrophia harperi Last Modified in 07/27/15 | 7 | 55 | 55 | 0 | 7 | 0 | 0 | 4 |
[33] † Shaleria ornatella Last Modified in 07/27/15 | 11 | 49 | 49 | 0 | 9 | 0 | 0 | 4 |
[34] † Eostrophonella eothen Last Modified in 07/27/15 | 16 | 40 | 40 | 0 | 13 | 0 | 0 | 4 |
[35] † Plectambonites planissimus Last Modified in 07/27/15 | 7 | 48 | 48 | 0 | 14 | 0 | 0 | 4 |
[36] † Isophragma ricevillense Last Modified in 07/27/15 | 2 | 51 | 51 | 0 | 16 | 0 | 0 | 4 |
[37] † Tafia planoconvexa Last Modified in 07/27/15 | 12 | 42 | 42 | 0 | 15 | 0 | 0 | 4 |
[38] † Ahtiella lirata Last Modified in 07/27/15 | 10 | 45 | 45 | 0 | 14 | 0 | 0 | 4 |
[39] † Leptella nevadensis Last Modified in 07/27/15 | 17 | 32 | 32 | 0 | 20 | 0 | 0 | 4 |
[40] † Bimuria superba Last Modified in 07/27/15 | 6 | 52 | 52 | 0 | 11 | 0 | 0 | 4 |
[41] † Sowerbyites hami Last Modified in 07/27/15 | 1 | 56 | 56 | 0 | 12 | 0 | 0 | 4 |
[42] † Leptellina? pulchra Last Modified in 07/27/15 | 10 | 52 | 52 | 0 | 7 | 0 | 0 | 4 |
[43] † Leptelloidea leptelloides Last Modified in 07/27/15 | 0 | 56 | 56 | 0 | 13 | 0 | 0 | 4 |
[44] † Palaeostrophomena concava Last Modified in 07/27/15 | 1 | 61 | 60 | 0 | 8 | 0 | 0 | 4 |
[45] † Grorudia grorudi Last Modified in 07/27/15 | 27 | 34 | 34 | 0 | 8 | 0 | 0 | 4 |
[46] † Alwynella ildjemensis Last Modified in 07/27/15 | 1 | 60 | 60 | 0 | 8 | 0 | 0 | 4 |
[47] † Railtonella scanloni Last Modified in 07/27/15 | 13 | 45 | 45 | 0 | 11 | 0 | 0 | 4 |
[48] † Leptestia musculosa Last Modified in 07/27/15 | 7 | 53 | 53 | 0 | 9 | 0 | 0 | 4 |
[49] † Sampo hiiuensis Last Modified in 07/27/15 | 3 | 60 | 60 | 0 | 6 | 0 | 0 | 4 |
[50] † Xenambonites undosus Last Modified in 07/27/15 | 4 | 56 | 56 | 0 | 9 | 0 | 0 | 4 |
[51] † Metambonites meritus Last Modified in 07/27/15 | 24 | 40 | 40 | 0 | 5 | 0 | 0 | 4 |
[52] † Aegiromena aquila Last Modified in 07/27/15 | 14 | 45 | 45 | 0 | 10 | 0 | 0 | 4 |
[53] † Epelidoaegiria chilidifera Last Modified in 07/27/15 | 13 | 47 | 47 | 0 | 9 | 0 | 0 | 4 |
[54] † Aegiria norvegica Last Modified in 07/27/15 | 7 | 52 | 52 | 0 | 10 | 0 | 0 | 4 |
[55] † Hesperomena leptellinoidea Last Modified in 07/27/15 | 24 | 40 | 40 | 0 | 5 | 0 | 0 | 4 |
[56] † Anoptambonites grayae Last Modified in 07/27/15 | 6 | 56 | 56 | 0 | 7 | 0 | 0 | 4 |
[57] † Sowerbyella punctostriata Last Modified in 07/27/15 | 7 | 59 | 59 | 0 | 3 | 0 | 0 | 4 |
[58] † Eochonetes clarksvillensis Last Modified in 07/27/15 | 16 | 47 | 47 | 0 | 6 | 0 | 0 | 4 |
[59] † Plectodonta mariae Last Modified in 07/27/15 | 21 | 43 | 43 | 0 | 5 | 0 | 0 | 4 |
[60] † Ptychoglyptus virginiensis Last Modified in 07/27/15 | 9 | 53 | 53 | 0 | 7 | 0 | 0 | 4 |
[61] † Strophochonetes cingulatus Last Modified in 06/03/15 | 25 | 36 | 36 | 0 | 8 | 0 | 0 | 4 |
[62] † Borealinetes comestus Last Modified in 07/27/15 | 7 | 53 | 53 | 0 | 9 | 0 | 0 | 4 |
[63] † Ctenochonetes tenuicostata Last Modified in 07/27/15 | 2 | 55 | 55 | 0 | 12 | 0 | 0 | 4 |
[64] † Archaeochonetes primigenius Last Modified in 07/27/15 | 14 | 48 | 48 | 0 | 7 | 0 | 0 | 4 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 33963 | |
Project overview | 3586 | |
Matrix views | 2998 | Matrix landing page (2163 views); Strophomenide phylogeny (280 views); Strophomenide phylogeny with microstructure characters included (246 views); Strophomenide phylogeny including Grorudia and Foliomena (309 views); |
Media views | 5790 | Media search (4954 views); M382061 (836 views); |
Specimen list | 2736 | |
Taxon list | 17711 | |
Bibliography | 1137 | |
Documents list | 5 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 330 | |
Project downloads | 313 | |
Matrix downloads | 17 | Strophomenide phylogeny with microstructure characters included (13 downloads); Strophomenide phylogeny (2 downloads); Strophomenide phylogeny including Grorudia and Foliomena (2 downloads); |