Project 1200: Alan Turner. 2015. A review of Shamosuchus and Paralligator (Crocodyliformes, Neosuchia) from the Cretaceous of Asia. PLOS ONE. 10 (2):e0118116.
Specimen: † Shamosuchus djadochtaensis (IGM:100/1195)

Abstract

The crocodyliform Shamosuchus is known from numerous Late Cretaceous localities in southern and eastern Mongolia and fragmentary remains from Uzbekistan. Seven species of Shamosuchus have been named from six localities in Mongolia and three in Uzbekistan. Six species originally described as Paralligator were later referred to Shamosuchus. Only the type species, Shamosuchus djadochtaensis has been examined in detail. Many of the named species of Shamosuchus show striking similarity in size and cranial morphology but most are based on partial remains suggesting that the true species diversity is overestimated.
A review of all species referred to Shamosuchus recognizes three valid taxa: Shamosuchus djadochtaensis, S. gradilifrons, and S. major. Shamosuchus sungaricus, S. borealis, and S. karakalpakensis are nomena dubia, whereas S. ancestralis, S. ulgicus, S. tersus, and S. ulanicus are junior synonyms of S. gradilifrons. Phylogenetic analysis of 318 phenotypic characters recovers a Paralligatoridae clade consisting of Shamosuchus, Rugosuchus, Batrachomimus, Glen Rose Form, and Wannchampsus. Shamosuchus is non-monophyletic: S. djadochatensis is near the base of Paralligatoridae whereas S. gradilifrons + S. major are the most deeply nested. The name Paralligator is resurrected for this clade. Rugosuchus and Batrachomimus are sister taxa to Paralligator. Paralligatoridae is closely related to Theriosuchus, hylaeochampsids and a speciose Allodaposuchus clade, which together are the sister group of Borealosuchus plus Crocodylia. These results support the presence of a diverse clade in eastern Asia and western North America throughout the Cretaceous with origins in the Late Jurassic.


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Article DOI: 10.1371/journal.pone.0118116

Project DOI: 10.7934/P1200, http://dx.doi.org/10.7934/P1200
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 13 Documents
  • 120 Taxa
  • 1 Specimen
  • 321 Characters
Total size of project's media files: 6 MB

Download Project SDD File
Total scored cells: 21071
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 321
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 731
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:321/0
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MorphoBank Project 1200

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    Alan Turner
    Project Administrator
    Last logged in 02/18/15
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    Taxonomic Overview for Matrix 'M2711' (109 Taxa)

    taxon unscored
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    [1] † Gracilisuchus stipanicicorum
    Last Modified in 02/19/15
    114202005002
    [2] † Terrestrisuchus gracilis
    Last Modified in 02/19/15
    157159006002
    [3] † Dibothrosuchus elaphros
    Last Modified in 02/19/15
    79236006002
    [4] † Protosuchus richardsoni
    Last Modified in 02/19/15
    50273004002
    [5] † Hemiprotosuchus leali
    Last Modified in 02/19/15
    194126001002
    [6] † Orthosuchus stormbergi
    Last Modified in 02/19/15
    91226005002
    [7] † Kayenta form
    Last Modified in 02/19/15
    142174006002
    [8] † Edentosuchus tienshanensis
    Last Modified in 02/19/15
    193134000002
    [9] † Zaraasuchus shepardi
    Last Modified in 02/19/15
    24282000002
    [10] † Gobiosuchus kielanae
    Last Modified in 02/19/15
    88235003002
    [11] † Sichuanosuchus shuhanensis
    Last Modified in 02/19/15
    135188002002
    [12] † Shantungosuchus hangjinensis
    Last Modified in 02/19/15
    208113002002
    [13] † Zosuchus davidsoni
    Last Modified in 02/19/15
    109215001002
    [14] † Fruitachampsa callisoni
    Last Modified in 02/19/15
    120200002002
    [15] † Notosuchus terrestris
    Last Modified in 02/19/15
    25298002002
    [16] † Comahuesuchus brachybuccalis
    Last Modified in 02/19/15
    156167002002
    [17] † Mariliasuchus amarali
    Last Modified in 02/19/15
    108215002002
    [18] † Uruguaysuchus aznarezi
    Last Modified in 02/19/15
    162162001002
    [19] † Chimaeresuchus paradoxus
    Last Modified in 02/19/15
    218103001002
    [20] † Malawisuchus mwakasyungutiensis
    Last Modified in 02/19/15
    80246001002
    [21] † Candidodon itapecuruense
    Last Modified in 02/19/15
    30119001002
    [22] † Simosuchus clarki
    Last Modified in 02/19/15
    16311001002
    [23] † Yacarerani boliviensis
    Last Modified in 02/19/15
    95226002002
    [24] † Sphagesaurus hueni
    Last Modified in 02/19/15
    159160002002
    [25] † Sphagesaurus montealtensis
    Last Modified in 02/19/15
    105216002002
    [26] † Adamantinasuchus navae
    Last Modified in 02/19/15
    22398000002
    [27] † Armadillosuchus arrudai
    Last Modified in 02/19/15
    204117001002
    [28] † Bretesuchus bonapartei
    Last Modified in 02/19/15
    206119001002
    [29] † Baurusuchus pachecoi
    Last Modified in 02/19/15
    99223001002
    [30] † Iberosuchus macrodon
    Last Modified in 02/19/15
    132197000002
    [31] † Libycosuchus brevirostris
    Last Modified in 02/19/15
    132195000002
    [32] † Anatosuchus minor
    Last Modified in 02/19/15
    48275001002
    [33] † Araripesuchus gomesii
    Last Modified in 02/19/15
    19308001002
    [34] † Araripesuchus patagonicus
    Last Modified in 02/19/15
    80245001002
    [35] † Araripesuchus buitreraensis
    Last Modified in 02/19/15
    190138001002
    [36] † Araripesuchus wegeneri
    Last Modified in 02/19/15
    29293002002
    [37] † Araripesuchus tsangatsangana
    Last Modified in 02/19/15
    41284001002
    [38] † Lomasuchus palpebrosus
    Last Modified in 02/19/15
    122200001002
    [39] † Peirosaurus torminni
    Last Modified in 02/19/15
    207115002002
    [40] † Hamadasuchus rebouli
    Last Modified in 02/19/15
    78242001002
    [41] † Uberabasuchus terrificus
    Last Modified in 02/19/15
    151174000002
    [42] † Montealtosuchus arrudacamposi
    Last Modified in 02/19/15
    73246002002
    [43] † Trematochampsa
    Last Modified in 02/19/15
    217108000002
    [44] † Mahajangasuchus insignis
    Last Modified in 02/19/15
    43280000002
    [45] † Kaprosuchus saharicus
    Last Modified in 02/19/15
    79243000002
    [46] † Stolokrosuchus lapparenti
    Last Modified in 02/19/15
    93228001002
    [47] † Hsisosuchus chungkingensis
    Last Modified in 02/19/15
    121202003002
    [48] † Pelagosaurus typus
    Last Modified in 02/19/15
    62263001002
    [49] † Steneosaurus bollensis
    Last Modified in 02/19/15
    55274001002
    [50] † Metriorhynchus superciliosus
    Last Modified in 02/19/15
    87236000002
    [51] † Metriorhynchus casamequelai
    Last Modified in 02/19/15
    201120000002
    [52] † Cricosaurus araucanensis
    Last Modified in 02/19/15
    109214001002
    [53] † Cricosaurus suevicus
    Last Modified in 02/19/15
    199124000002
    [54] † Dakosaurus maximus
    Last Modified in 02/19/15
    23883000002
    [55] † Dakosaurus andiniensis
    Last Modified in 02/19/15
    148174001002
    [56] † Rhabdognathus
    Last Modified in 02/19/15
    132191001002
    [57] † Sokotosuchus ianwilsoni
    Last Modified in 02/19/15
    25170000002
    [58] † Dyrosaurus
    Last Modified in 02/19/15
    126194001002
    [59] † Hyposaurus rogersii
    Last Modified in 02/19/15
    203118000002
    [60] † Pholidosaurus purbeckensis
    Last Modified in 02/19/15
    174148000002
    [61] † Sarcosuchus imperator
    Last Modified in 02/19/15
    66261001002
    [62] † Terminonaris robusta
    Last Modified in 02/19/15
    93233001002
    [63] † Elosuchus cherifiensis
    Last Modified in 02/19/15
    217103001002
    [64] † Meridiosaurus vallisparadisi
    Last Modified in 02/19/15
    26456001002
    [65] † Oceanosuchus boecensis
    Last Modified in 02/19/15
    23685000002
    [66] † Paluxysuchus newmani
    Last Modified in 02/19/15
    112208001002
    [67] † Goniopholis simus
    Last Modified in 02/19/15
    68255001002
    [68] † Amphicotylus lucasii
    Last Modified in 02/19/15
    123196002002
    [69] † Calsoyasuchus valliceps
    Last Modified in 02/19/15
    161159002002
    [70] † Sunosuchus junggarensis
    Last Modified in 02/19/15
    192127002002
    [71] EutreALL
    Last Modified in 06/03/14
    502610010002
    [72] † Goniopholis baryglyphaeus
    Last Modified in 02/19/15
    196125000002
    [73] † Alligatorium
    Last Modified in 02/19/15
    24477000002
    [74] † Theriosuchus pusillus
    Last Modified in 02/19/15
    59264003002
    [75] † Theriosuchus sympiestodon
    Last Modified in 02/19/15
    24275004002
    [76] † Theriosuchus guimarotae
    Last Modified in 02/19/15
    171151001002
    [77] † Wannchampsus kirpachi
    Last Modified in 02/19/15
    88232001002
    [78] † Shamosuchus major
    Last Modified in 02/19/15
    164152007002
    [79] † Shamosuchus djadochtaensis
    Last Modified in 02/19/15
    76252001002
    [80] † Shamosuchus gradilifrons
    Last Modified in 02/19/15
    40278008002
    [81] † Shamosuchus gradilifrons holotype
    Last Modified in 02/19/15
    105218004002
    [82] † Shamosuchus tersus
    Last Modified in 02/19/15
    138182004002
    [83] † Shamosuchus ancestralis
    Last Modified in 02/19/15
    23091001002
    [84] † Shamosuchus ulanicus
    Last Modified in 02/19/15
    89228009002
    [85] † Shamosuchus ulgicus
    Last Modified in 02/19/15
    3137001002
    [86] † Rugosuchus nonganensis
    Last Modified in 02/19/15
    133191001002
    [87] † Batrachomimus pastosbonensis
    Last Modified in 02/19/15
    139173009002
    [88] Glen rose form
    Last Modified in 06/03/14
    119201001002
    [89] † Bernissartia fagesii
    Last Modified in 02/19/15
    125195001002
    [90] † Hylaeochampsa vectiana
    Last Modified in 02/19/15
    23486001002
    [91] † Iharkutosuchus makadii
    Last Modified in 02/19/15
    94226001002
    [92] † Acynodon iberoccitanus
    Last Modified in 02/19/15
    952080018002
    [93] † Acynodon adriaticus
    Last Modified in 02/19/15
    1052060012002
    [94] † Allodaposuchus precedens
    Last Modified in 02/19/15
    179144001002
    [95] † Allodaposuchus subjuniperus
    Last Modified in 02/19/15
    130190001002
    [96] † Allodaposuchus cf. precedens
    Last Modified in 02/19/15
    1111980012002
    [97] † Pachycheilosuchus trinquei
    Last Modified in 02/19/15
    22698000002
    [98] † Susisuchus anatoceps
    Last Modified in 02/19/15
    157164000002
    [99] † Isisfordia duncani
    Last Modified in 02/19/15
    442670011002
    [100] † Borealosuchus formidabilis
    Last Modified in 02/19/15
    46275001002
    [101] † Argochampsa krebsi
    Last Modified in 02/19/15
    147175001002
    [102] † Pristichampsus vorax
    Last Modified in 02/19/15
    76246001002
    [103] † Eothoracosaurus mississippiensis
    Last Modified in 02/19/15
    99227001002
    [104] Gavialis gangeticus
    Last Modified in 06/03/14
    23300001002
    [105] † Leidyosuchus canadensis
    Last Modified in 02/19/15
    68254001002
    [106] † Asiatosuchus germanicus
    Last Modified in 02/19/15
    93228001002
    [107] Crocodylus niloticus
    Last Modified in 06/03/14
    18303001002
    [108] † Diplocynodon hantonensis
    Last Modified in 02/19/15
    41280001002
    [109] Alligator mississippiensis
    Last Modified in 06/03/14
    17304001002


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    Project overview3497
    Media views3880Media search (3437 views); M333576 (443 views);
    Matrix views2383Matrix landing page (2000 views); A review of Shamosuchus and Paralligator (Crocodyliformes, Neosuchia) from the Cretaceous of Asia (383 views);
    Specimen list1746
    Taxon list9377
    Documents list1699
    Bibliography627




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    Matrix downloads125A review of Shamosuchus and Paralligator (Crocodyliformes, Neosuchia) from the Cretaceous of Asia (125 downloads);
    Document downloads61MrBayes Results Treefile (10 downloads); MrBayes analysis file (9 downloads); TNT file (15 downloads); Isisfordia Primary Analysis TNT file (8 downloads); TNT constraint file (9 downloads); Incipient procoely Tree File (1 download); Derive palate and procoely TNT file (1 download); Derived palate TNT file (2 downloads); Derived procoely TNT file (2 downloads); Incipient procoely TNT file (3 downloads); Derived palate strict consensus tree file (1 download);
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