Morphobank

Project 1200: Alan Turner. 2015. A review of Shamosuchus and Paralligator (Crocodyliformes, Neosuchia) from the Cretaceous of Asia. PLOS ONE. 10 (2):e0118116.

Specimen: † Shamosuchus djadochtaensis (IGM:100/1195)

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Abstract

The crocodyliform Shamosuchus is known from numerous Late Cretaceous localities in southern and eastern Mongolia and fragmentary remains from Uzbekistan. Seven species of Shamosuchus have been named from six localities in Mongolia and three in Uzbekistan. Six species originally described as Paralligator were later referred to Shamosuchus. Only the type species, Shamosuchus djadochtaensis has been examined in detail. Many of the named species of Shamosuchus show striking similarity in size and cranial morphology but most are based on partial remains suggesting that the true species diversity is overestimated.
A review of all species referred to Shamosuchus recognizes three valid taxa: Shamosuchus djadochtaensis, S. gradilifrons, and S. major. Shamosuchus sungaricus, S. borealis, and S. karakalpakensis are nomena dubia, whereas S. ancestralis, S. ulgicus, S. tersus, and S. ulanicus are junior synonyms of S. gradilifrons. Phylogenetic analysis of 318 phenotypic characters recovers a Paralligatoridae clade consisting of Shamosuchus, Rugosuchus, Batrachomimus, Glen Rose Form, and Wannchampsus. Shamosuchus is non-monophyletic: S. djadochatensis is near the base of Paralligatoridae whereas S. gradilifrons + S. major are the most deeply nested. The name Paralligator is resurrected for this clade. Rugosuchus and Batrachomimus are sister taxa to Paralligator. Paralligatoridae is closely related to Theriosuchus, hylaeochampsids and a speciose Allodaposuchus clade, which together are the sister group of Borealosuchus plus Crocodylia. These results support the presence of a diverse clade in eastern Asia and western North America throughout the Cretaceous with origins in the Late Jurassic.

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Article DOI: 10.1371/journal.pone.0118116

Project DOI: 10.7934/P1200, http://dx.doi.org/10.7934/P1200

This project contains	Matrices
1 Media 1 Matrix 13 Documents 120 Taxa 1 Specimen 321 Characters Total size of project's media files: 5.66M Download Project SDD File	Total scored cells: 21071 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 321 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 731 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:321/0

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MorphoBank Project 1200

Creation Date:
03 June 2014
Publication Date:
18 February 2015
Media downloads: 3
Matrix downloads: 147

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Stony Brook University

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Alan Turner
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Taxonomic Overview for Matrix 'M2711' (109 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Gracilisuchus stipanicicorum Taxon name last Modified on 02/19/15	114	202	0	0	5	0	0	2
[2] † Terrestrisuchus gracilis Taxon name last Modified on 02/19/15	157	159	0	0	6	0	0	2
[3] † Dibothrosuchus elaphros Taxon name last Modified on 02/19/15	79	236	0	0	6	0	0	2
[4] † Protosuchus richardsoni Taxon name last Modified on 02/19/15	50	273	0	0	4	0	0	2
[5] † Hemiprotosuchus leali Taxon name last Modified on 02/19/15	194	126	0	0	1	0	0	2
[6] † Orthosuchus stormbergi Taxon name last Modified on 02/19/15	91	226	0	0	5	0	0	2
[7] † Kayenta form Taxon name last Modified on 02/19/15	142	174	0	0	6	0	0	2
[8] † Edentosuchus tienshanensis Taxon name last Modified on 02/19/15	193	134	0	0	0	0	0	2
[9] † Zaraasuchus shepardi Taxon name last Modified on 02/19/15	242	82	0	0	0	0	0	2
[10] † Gobiosuchus kielanae Taxon name last Modified on 02/19/15	88	235	0	0	3	0	0	2
[11] † Sichuanosuchus shuhanensis Taxon name last Modified on 02/19/15	135	188	0	0	2	0	0	2
[12] † Shantungosuchus hangjinensis Taxon name last Modified on 02/19/15	208	113	0	0	2	0	0	2
[13] † Zosuchus davidsoni Taxon name last Modified on 02/19/15	109	215	0	0	1	0	0	2
[14] † Fruitachampsa callisoni Taxon name last Modified on 02/19/15	120	200	0	0	2	0	0	2
[15] † Notosuchus terrestris Taxon name last Modified on 02/19/15	25	298	0	0	2	0	0	2
[16] † Comahuesuchus brachybuccalis Taxon name last Modified on 02/19/15	156	167	0	0	2	0	0	2
[17] † Mariliasuchus amarali Taxon name last Modified on 02/19/15	108	215	0	0	2	0	0	2
[18] † Uruguaysuchus aznarezi Taxon name last Modified on 02/19/15	162	162	0	0	1	0	0	2
[19] † Chimaeresuchus paradoxus Taxon name last Modified on 02/19/15	218	103	0	0	1	0	0	2
[20] † Malawisuchus mwakasyungutiensis Taxon name last Modified on 02/19/15	80	246	0	0	1	0	0	2
[21] † Candidodon itapecuruense Taxon name last Modified on 02/19/15	301	19	0	0	1	0	0	2
[22] † Simosuchus clarki Taxon name last Modified on 02/19/15	16	311	0	0	1	0	0	2
[23] † Yacarerani boliviensis Taxon name last Modified on 02/19/15	95	226	0	0	2	0	0	2
[24] † Sphagesaurus hueni Taxon name last Modified on 02/19/15	159	160	0	0	2	0	0	2
[25] † Sphagesaurus montealtensis Taxon name last Modified on 02/19/15	105	216	0	0	2	0	0	2
[26] † Adamantinasuchus navae Taxon name last Modified on 02/19/15	223	98	0	0	0	0	0	2
[27] † Armadillosuchus arrudai Taxon name last Modified on 02/19/15	204	117	0	0	1	0	0	2
[28] † Bretesuchus bonapartei Taxon name last Modified on 02/19/15	206	119	0	0	1	0	0	2
[29] † Baurusuchus pachecoi Taxon name last Modified on 02/19/15	99	223	0	0	1	0	0	2
[30] † Iberosuchus macrodon Taxon name last Modified on 02/19/15	132	197	0	0	0	0	0	2
[31] † Libycosuchus brevirostris Taxon name last Modified on 02/19/15	132	195	0	0	0	0	0	2
[32] † Anatosuchus minor Taxon name last Modified on 02/19/15	48	275	0	0	1	0	0	2
[33] † Araripesuchus gomesii Taxon name last Modified on 02/19/15	19	308	0	0	1	0	0	2
[34] † Araripesuchus patagonicus Taxon name last Modified on 02/19/15	80	245	0	0	1	0	0	2
[35] † Araripesuchus buitreraensis Taxon name last Modified on 02/19/15	190	138	0	0	1	0	0	2
[36] † Araripesuchus wegeneri Taxon name last Modified on 02/19/15	29	293	0	0	2	0	0	2
[37] † Araripesuchus tsangatsangana Taxon name last Modified on 02/19/15	41	284	0	0	1	0	0	2
[38] † Lomasuchus palpebrosus Taxon name last Modified on 02/19/15	122	200	0	0	1	0	0	2
[39] † Peirosaurus torminni Taxon name last Modified on 02/19/15	207	115	0	0	2	0	0	2
[40] † Hamadasuchus rebouli Taxon name last Modified on 02/19/15	78	242	0	0	1	0	0	2
[41] † Uberabasuchus terrificus Taxon name last Modified on 02/19/15	151	174	0	0	0	0	0	2
[42] † Montealtosuchus arrudacamposi Taxon name last Modified on 02/19/15	73	246	0	0	2	0	0	2
[43] † Trematochampsa Taxon name last Modified on 02/19/15	217	108	0	0	0	0	0	2
[44] † Mahajangasuchus insignis Taxon name last Modified on 02/19/15	43	280	0	0	0	0	0	2
[45] † Kaprosuchus saharicus Taxon name last Modified on 02/19/15	79	243	0	0	0	0	0	2
[46] † Stolokrosuchus lapparenti Taxon name last Modified on 02/19/15	93	228	0	0	1	0	0	2
[47] † Hsisosuchus chungkingensis Taxon name last Modified on 02/19/15	121	202	0	0	3	0	0	2
[48] † Pelagosaurus typus Taxon name last Modified on 02/19/15	62	263	0	0	1	0	0	2
[49] † Steneosaurus bollensis Taxon name last Modified on 02/19/15	55	274	0	0	1	0	0	2
[50] † Metriorhynchus superciliosus Taxon name last Modified on 02/19/15	87	236	0	0	0	0	0	2
[51] † Metriorhynchus casamequelai Taxon name last Modified on 02/19/15	201	120	0	0	0	0	0	2
[52] † Cricosaurus araucanensis Taxon name last Modified on 02/19/15	109	214	0	0	1	0	0	2
[53] † Cricosaurus suevicus Taxon name last Modified on 02/19/15	199	124	0	0	0	0	0	2
[54] † Dakosaurus maximus Taxon name last Modified on 02/19/15	238	83	0	0	0	0	0	2
[55] † Dakosaurus andiniensis Taxon name last Modified on 02/19/15	148	174	0	0	1	0	0	2
[56] † Rhabdognathus Taxon name last Modified on 02/19/15	132	191	0	0	1	0	0	2
[57] † Sokotosuchus ianwilsoni Taxon name last Modified on 02/19/15	251	70	0	0	0	0	0	2
[58] † Dyrosaurus Taxon name last Modified on 02/19/15	126	194	0	0	1	0	0	2
[59] † Hyposaurus rogersii Taxon name last Modified on 02/19/15	203	118	0	0	0	0	0	2
[60] † Pholidosaurus purbeckensis Taxon name last Modified on 02/19/15	174	148	0	0	0	0	0	2
[61] † Sarcosuchus imperator Taxon name last Modified on 02/19/15	66	261	0	0	1	0	0	2
[62] † Terminonaris robusta Taxon name last Modified on 02/19/15	93	233	0	0	1	0	0	2
[63] † Elosuchus cherifiensis Taxon name last Modified on 02/19/15	217	103	0	0	1	0	0	2
[64] † Meridiosaurus vallisparadisi Taxon name last Modified on 02/19/15	264	56	0	0	1	0	0	2
[65] † Oceanosuchus boecensis Taxon name last Modified on 02/19/15	236	85	0	0	0	0	0	2
[66] † Paluxysuchus newmani Taxon name last Modified on 02/19/15	112	208	0	0	1	0	0	2
[67] † Goniopholis simus Taxon name last Modified on 02/19/15	68	255	0	0	1	0	0	2
[68] † Amphicotylus lucasii Taxon name last Modified on 02/19/15	123	196	0	0	2	0	0	2
[69] † Calsoyasuchus valliceps Taxon name last Modified on 02/19/15	161	159	0	0	2	0	0	2
[70] † Sunosuchus junggarensis Taxon name last Modified on 02/19/15	192	127	0	0	2	0	0	2
[71] EutreALL Taxon name last Modified on 06/03/14	50	261	0	0	10	0	0	2
[72] † Goniopholis baryglyphaeus Taxon name last Modified on 02/19/15	196	125	0	0	0	0	0	2
[73] † Alligatorium Taxon name last Modified on 02/19/15	244	77	0	0	0	0	0	2
[74] † Theriosuchus pusillus Taxon name last Modified on 02/19/15	59	264	0	0	3	0	0	2
[75] † Theriosuchus sympiestodon Taxon name last Modified on 02/19/15	242	75	0	0	4	0	0	2
[76] † Theriosuchus guimarotae Taxon name last Modified on 02/19/15	171	151	0	0	1	0	0	2
[77] † Wannchampsus kirpachi Taxon name last Modified on 02/19/15	88	232	0	0	1	0	0	2
[78] † Shamosuchus major Taxon name last Modified on 02/19/15	164	152	0	0	7	0	0	2
[79] † Shamosuchus djadochtaensis Taxon name last Modified on 02/19/15	76	252	0	0	1	0	0	2
[80] † Shamosuchus gradilifrons Taxon name last Modified on 02/19/15	40	278	0	0	8	0	0	2
[81] † Shamosuchus gradilifrons holotype Taxon name last Modified on 02/19/15	105	218	0	0	4	0	0	2
[82] † Shamosuchus tersus Taxon name last Modified on 02/19/15	138	182	0	0	4	0	0	2
[83] † Shamosuchus ancestralis Taxon name last Modified on 02/19/15	230	91	0	0	1	0	0	2
[84] † Shamosuchus ulanicus Taxon name last Modified on 02/19/15	89	228	0	0	9	0	0	2
[85] † Shamosuchus ulgicus Taxon name last Modified on 02/19/15	313	7	0	0	1	0	0	2
[86] † Rugosuchus nonganensis Taxon name last Modified on 02/19/15	133	191	0	0	1	0	0	2
[87] † Batrachomimus pastosbonensis Taxon name last Modified on 02/19/15	139	173	0	0	9	0	0	2
[88] Glen rose form Taxon name last Modified on 06/03/14	119	201	0	0	1	0	0	2
[89] † Bernissartia fagesii Taxon name last Modified on 02/19/15	125	195	0	0	1	0	0	2
[90] † Hylaeochampsa vectiana Taxon name last Modified on 02/19/15	234	86	0	0	1	0	0	2
[91] † Iharkutosuchus makadii Taxon name last Modified on 02/19/15	94	226	0	0	1	0	0	2
[92] † Acynodon iberoccitanus Taxon name last Modified on 02/19/15	95	208	0	0	18	0	0	2
[93] † Acynodon adriaticus Taxon name last Modified on 02/19/15	105	206	0	0	12	0	0	2
[94] † Allodaposuchus precedens Taxon name last Modified on 02/19/15	179	144	0	0	1	0	0	2
[95] † Allodaposuchus subjuniperus Taxon name last Modified on 02/19/15	130	190	0	0	1	0	0	2
[96] † Allodaposuchus cf. precedens Taxon name last Modified on 02/19/15	111	198	0	0	12	0	0	2
[97] † Pachycheilosuchus trinquei Taxon name last Modified on 02/19/15	226	98	0	0	0	0	0	2
[98] † Susisuchus anatoceps Taxon name last Modified on 02/19/15	157	164	0	0	0	0	0	2
[99] † Isisfordia duncani Taxon name last Modified on 02/19/15	44	267	0	0	11	0	0	2
[100] † Borealosuchus formidabilis Taxon name last Modified on 02/19/15	46	275	0	0	1	0	0	2
[101] † Argochampsa krebsi Taxon name last Modified on 02/19/15	147	175	0	0	1	0	0	2
[102] † Pristichampsus vorax Taxon name last Modified on 02/19/15	76	246	0	0	1	0	0	2
[103] † Eothoracosaurus mississippiensis Taxon name last Modified on 02/19/15	99	227	0	0	1	0	0	2
[104] Gavialis gangeticus Taxon name last Modified on 06/03/14	23	300	0	0	1	0	0	2
[105] † Leidyosuchus canadensis Taxon name last Modified on 02/19/15	68	254	0	0	1	0	0	2
[106] † Asiatosuchus germanicus Taxon name last Modified on 02/19/15	93	228	0	0	1	0	0	2
[107] Crocodylus niloticus Taxon name last Modified on 06/03/14	18	303	0	0	1	0	0	2
[108] † Diplocynodon hantonensis Taxon name last Modified on 02/19/15	41	280	0	0	1	0	0	2
[109] Alligator mississippiensis Taxon name last Modified on 06/03/14	17	304	0	0	1	0	0	2

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	679
Matrix downloads	147	A review of Shamosuchus and Paralligator (Crocodyliformes, Neosuchia) from the Cretaceous of Asia (147 downloads);
Document downloads	67	MrBayes Results Treefile (10 downloads); MrBayes analysis file (9 downloads); TNT file (18 downloads); Isisfordia Primary Analysis TNT file (8 downloads); TNT constraint file (12 downloads); Incipient procoely Tree File (1 download); Derive palate and procoely TNT file (1 download); Derived palate TNT file (2 downloads); Derived procoely TNT file (2 downloads); Incipient procoely TNT file (3 downloads); Derived palate strict consensus tree file (1 download);
Project downloads	462
Media downloads	3	M333576 (3 downloads);