Morphobank

Project 1200: Alan Turner. 2015. A review of Shamosuchus and Paralligator (Crocodyliformes, Neosuchia) from the Cretaceous of Asia. PLOS ONE. 10 (2):e0118116.

Specimen: † Shamosuchus djadochtaensis (IGM:100/1195)

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Abstract

The crocodyliform Shamosuchus is known from numerous Late Cretaceous localities in southern and eastern Mongolia and fragmentary remains from Uzbekistan. Seven species of Shamosuchus have been named from six localities in Mongolia and three in Uzbekistan. Six species originally described as Paralligator were later referred to Shamosuchus. Only the type species, Shamosuchus djadochtaensis has been examined in detail. Many of the named species of Shamosuchus show striking similarity in size and cranial morphology but most are based on partial remains suggesting that the true species diversity is overestimated.
A review of all species referred to Shamosuchus recognizes three valid taxa: Shamosuchus djadochtaensis, S. gradilifrons, and S. major. Shamosuchus sungaricus, S. borealis, and S. karakalpakensis are nomena dubia, whereas S. ancestralis, S. ulgicus, S. tersus, and S. ulanicus are junior synonyms of S. gradilifrons. Phylogenetic analysis of 318 phenotypic characters recovers a Paralligatoridae clade consisting of Shamosuchus, Rugosuchus, Batrachomimus, Glen Rose Form, and Wannchampsus. Shamosuchus is non-monophyletic: S. djadochatensis is near the base of Paralligatoridae whereas S. gradilifrons + S. major are the most deeply nested. The name Paralligator is resurrected for this clade. Rugosuchus and Batrachomimus are sister taxa to Paralligator. Paralligatoridae is closely related to Theriosuchus, hylaeochampsids and a speciose Allodaposuchus clade, which together are the sister group of Borealosuchus plus Crocodylia. These results support the presence of a diverse clade in eastern Asia and western North America throughout the Cretaceous with origins in the Late Jurassic.

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Article DOI: 10.1371/journal.pone.0118116

Project DOI: 10.7934/P1200, http://dx.doi.org/10.7934/P1200

This project contains	Matrices
1 Media 1 Matrix 13 Documents 120 Taxa 1 Specimen 321 Characters Total size of project's media files: 5.66M Download Project SDD File	Total scored cells: 21071 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 321 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 731 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:321/0

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MorphoBank Project 1200

Creation Date:
03 June 2014
Publication Date:
18 February 2015
Project views: 44265
Media downloads: 3
Matrix downloads: 146

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Stony Brook University

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Alan Turner
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Taxonomic Overview for Matrix 'M2711' (109 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Gracilisuchus stipanicicorum Last Modified in 02/19/15	114	202	0	0	5	0	0	2
[2] † Terrestrisuchus gracilis Last Modified in 02/19/15	157	159	0	0	6	0	0	2
[3] † Dibothrosuchus elaphros Last Modified in 02/19/15	79	236	0	0	6	0	0	2
[4] † Protosuchus richardsoni Last Modified in 02/19/15	50	273	0	0	4	0	0	2
[5] † Hemiprotosuchus leali Last Modified in 02/19/15	194	126	0	0	1	0	0	2
[6] † Orthosuchus stormbergi Last Modified in 02/19/15	91	226	0	0	5	0	0	2
[7] † Kayenta form Last Modified in 02/19/15	142	174	0	0	6	0	0	2
[8] † Edentosuchus tienshanensis Last Modified in 02/19/15	193	134	0	0	0	0	0	2
[9] † Zaraasuchus shepardi Last Modified in 02/19/15	242	82	0	0	0	0	0	2
[10] † Gobiosuchus kielanae Last Modified in 02/19/15	88	235	0	0	3	0	0	2
[11] † Sichuanosuchus shuhanensis Last Modified in 02/19/15	135	188	0	0	2	0	0	2
[12] † Shantungosuchus hangjinensis Last Modified in 02/19/15	208	113	0	0	2	0	0	2
[13] † Zosuchus davidsoni Last Modified in 02/19/15	109	215	0	0	1	0	0	2
[14] † Fruitachampsa callisoni Last Modified in 02/19/15	120	200	0	0	2	0	0	2
[15] † Notosuchus terrestris Last Modified in 02/19/15	25	298	0	0	2	0	0	2
[16] † Comahuesuchus brachybuccalis Last Modified in 02/19/15	156	167	0	0	2	0	0	2
[17] † Mariliasuchus amarali Last Modified in 02/19/15	108	215	0	0	2	0	0	2
[18] † Uruguaysuchus aznarezi Last Modified in 02/19/15	162	162	0	0	1	0	0	2
[19] † Chimaeresuchus paradoxus Last Modified in 02/19/15	218	103	0	0	1	0	0	2
[20] † Malawisuchus mwakasyungutiensis Last Modified in 02/19/15	80	246	0	0	1	0	0	2
[21] † Candidodon itapecuruense Last Modified in 02/19/15	301	19	0	0	1	0	0	2
[22] † Simosuchus clarki Last Modified in 02/19/15	16	311	0	0	1	0	0	2
[23] † Yacarerani boliviensis Last Modified in 02/19/15	95	226	0	0	2	0	0	2
[24] † Sphagesaurus hueni Last Modified in 02/19/15	159	160	0	0	2	0	0	2
[25] † Sphagesaurus montealtensis Last Modified in 02/19/15	105	216	0	0	2	0	0	2
[26] † Adamantinasuchus navae Last Modified in 02/19/15	223	98	0	0	0	0	0	2
[27] † Armadillosuchus arrudai Last Modified in 02/19/15	204	117	0	0	1	0	0	2
[28] † Bretesuchus bonapartei Last Modified in 02/19/15	206	119	0	0	1	0	0	2
[29] † Baurusuchus pachecoi Last Modified in 02/19/15	99	223	0	0	1	0	0	2
[30] † Iberosuchus macrodon Last Modified in 02/19/15	132	197	0	0	0	0	0	2
[31] † Libycosuchus brevirostris Last Modified in 02/19/15	132	195	0	0	0	0	0	2
[32] † Anatosuchus minor Last Modified in 02/19/15	48	275	0	0	1	0	0	2
[33] † Araripesuchus gomesii Last Modified in 02/19/15	19	308	0	0	1	0	0	2
[34] † Araripesuchus patagonicus Last Modified in 02/19/15	80	245	0	0	1	0	0	2
[35] † Araripesuchus buitreraensis Last Modified in 02/19/15	190	138	0	0	1	0	0	2
[36] † Araripesuchus wegeneri Last Modified in 02/19/15	29	293	0	0	2	0	0	2
[37] † Araripesuchus tsangatsangana Last Modified in 02/19/15	41	284	0	0	1	0	0	2
[38] † Lomasuchus palpebrosus Last Modified in 02/19/15	122	200	0	0	1	0	0	2
[39] † Peirosaurus torminni Last Modified in 02/19/15	207	115	0	0	2	0	0	2
[40] † Hamadasuchus rebouli Last Modified in 02/19/15	78	242	0	0	1	0	0	2
[41] † Uberabasuchus terrificus Last Modified in 02/19/15	151	174	0	0	0	0	0	2
[42] † Montealtosuchus arrudacamposi Last Modified in 02/19/15	73	246	0	0	2	0	0	2
[43] † Trematochampsa Last Modified in 02/19/15	217	108	0	0	0	0	0	2
[44] † Mahajangasuchus insignis Last Modified in 02/19/15	43	280	0	0	0	0	0	2
[45] † Kaprosuchus saharicus Last Modified in 02/19/15	79	243	0	0	0	0	0	2
[46] † Stolokrosuchus lapparenti Last Modified in 02/19/15	93	228	0	0	1	0	0	2
[47] † Hsisosuchus chungkingensis Last Modified in 02/19/15	121	202	0	0	3	0	0	2
[48] † Pelagosaurus typus Last Modified in 02/19/15	62	263	0	0	1	0	0	2
[49] † Steneosaurus bollensis Last Modified in 02/19/15	55	274	0	0	1	0	0	2
[50] † Metriorhynchus superciliosus Last Modified in 02/19/15	87	236	0	0	0	0	0	2
[51] † Metriorhynchus casamequelai Last Modified in 02/19/15	201	120	0	0	0	0	0	2
[52] † Cricosaurus araucanensis Last Modified in 02/19/15	109	214	0	0	1	0	0	2
[53] † Cricosaurus suevicus Last Modified in 02/19/15	199	124	0	0	0	0	0	2
[54] † Dakosaurus maximus Last Modified in 02/19/15	238	83	0	0	0	0	0	2
[55] † Dakosaurus andiniensis Last Modified in 02/19/15	148	174	0	0	1	0	0	2
[56] † Rhabdognathus Last Modified in 02/19/15	132	191	0	0	1	0	0	2
[57] † Sokotosuchus ianwilsoni Last Modified in 02/19/15	251	70	0	0	0	0	0	2
[58] † Dyrosaurus Last Modified in 02/19/15	126	194	0	0	1	0	0	2
[59] † Hyposaurus rogersii Last Modified in 02/19/15	203	118	0	0	0	0	0	2
[60] † Pholidosaurus purbeckensis Last Modified in 02/19/15	174	148	0	0	0	0	0	2
[61] † Sarcosuchus imperator Last Modified in 02/19/15	66	261	0	0	1	0	0	2
[62] † Terminonaris robusta Last Modified in 02/19/15	93	233	0	0	1	0	0	2
[63] † Elosuchus cherifiensis Last Modified in 02/19/15	217	103	0	0	1	0	0	2
[64] † Meridiosaurus vallisparadisi Last Modified in 02/19/15	264	56	0	0	1	0	0	2
[65] † Oceanosuchus boecensis Last Modified in 02/19/15	236	85	0	0	0	0	0	2
[66] † Paluxysuchus newmani Last Modified in 02/19/15	112	208	0	0	1	0	0	2
[67] † Goniopholis simus Last Modified in 02/19/15	68	255	0	0	1	0	0	2
[68] † Amphicotylus lucasii Last Modified in 02/19/15	123	196	0	0	2	0	0	2
[69] † Calsoyasuchus valliceps Last Modified in 02/19/15	161	159	0	0	2	0	0	2
[70] † Sunosuchus junggarensis Last Modified in 02/19/15	192	127	0	0	2	0	0	2
[71] EutreALL Last Modified in 06/03/14	50	261	0	0	10	0	0	2
[72] † Goniopholis baryglyphaeus Last Modified in 02/19/15	196	125	0	0	0	0	0	2
[73] † Alligatorium Last Modified in 02/19/15	244	77	0	0	0	0	0	2
[74] † Theriosuchus pusillus Last Modified in 02/19/15	59	264	0	0	3	0	0	2
[75] † Theriosuchus sympiestodon Last Modified in 02/19/15	242	75	0	0	4	0	0	2
[76] † Theriosuchus guimarotae Last Modified in 02/19/15	171	151	0	0	1	0	0	2
[77] † Wannchampsus kirpachi Last Modified in 02/19/15	88	232	0	0	1	0	0	2
[78] † Shamosuchus major Last Modified in 02/19/15	164	152	0	0	7	0	0	2
[79] † Shamosuchus djadochtaensis Last Modified in 02/19/15	76	252	0	0	1	0	0	2
[80] † Shamosuchus gradilifrons Last Modified in 02/19/15	40	278	0	0	8	0	0	2
[81] † Shamosuchus gradilifrons holotype Last Modified in 02/19/15	105	218	0	0	4	0	0	2
[82] † Shamosuchus tersus Last Modified in 02/19/15	138	182	0	0	4	0	0	2
[83] † Shamosuchus ancestralis Last Modified in 02/19/15	230	91	0	0	1	0	0	2
[84] † Shamosuchus ulanicus Last Modified in 02/19/15	89	228	0	0	9	0	0	2
[85] † Shamosuchus ulgicus Last Modified in 02/19/15	313	7	0	0	1	0	0	2
[86] † Rugosuchus nonganensis Last Modified in 02/19/15	133	191	0	0	1	0	0	2
[87] † Batrachomimus pastosbonensis Last Modified in 02/19/15	139	173	0	0	9	0	0	2
[88] Glen rose form Last Modified in 06/03/14	119	201	0	0	1	0	0	2
[89] † Bernissartia fagesii Last Modified in 02/19/15	125	195	0	0	1	0	0	2
[90] † Hylaeochampsa vectiana Last Modified in 02/19/15	234	86	0	0	1	0	0	2
[91] † Iharkutosuchus makadii Last Modified in 02/19/15	94	226	0	0	1	0	0	2
[92] † Acynodon iberoccitanus Last Modified in 02/19/15	95	208	0	0	18	0	0	2
[93] † Acynodon adriaticus Last Modified in 02/19/15	105	206	0	0	12	0	0	2
[94] † Allodaposuchus precedens Last Modified in 02/19/15	179	144	0	0	1	0	0	2
[95] † Allodaposuchus subjuniperus Last Modified in 02/19/15	130	190	0	0	1	0	0	2
[96] † Allodaposuchus cf. precedens Last Modified in 02/19/15	111	198	0	0	12	0	0	2
[97] † Pachycheilosuchus trinquei Last Modified in 02/19/15	226	98	0	0	0	0	0	2
[98] † Susisuchus anatoceps Last Modified in 02/19/15	157	164	0	0	0	0	0	2
[99] † Isisfordia duncani Last Modified in 02/19/15	44	267	0	0	11	0	0	2
[100] † Borealosuchus formidabilis Last Modified in 02/19/15	46	275	0	0	1	0	0	2
[101] † Argochampsa krebsi Last Modified in 02/19/15	147	175	0	0	1	0	0	2
[102] † Pristichampsus vorax Last Modified in 02/19/15	76	246	0	0	1	0	0	2
[103] † Eothoracosaurus mississippiensis Last Modified in 02/19/15	99	227	0	0	1	0	0	2
[104] Gavialis gangeticus Last Modified in 06/03/14	23	300	0	0	1	0	0	2
[105] † Leidyosuchus canadensis Last Modified in 02/19/15	68	254	0	0	1	0	0	2
[106] † Asiatosuchus germanicus Last Modified in 02/19/15	93	228	0	0	1	0	0	2
[107] Crocodylus niloticus Last Modified in 06/03/14	18	303	0	0	1	0	0	2
[108] † Diplocynodon hantonensis Last Modified in 02/19/15	41	280	0	0	1	0	0	2
[109] Alligator mississippiensis Last Modified in 06/03/14	17	304	0	0	1	0	0	2

Project views

type	number of views	Individual items viewed (where applicable)
Total project views	44265
Project overview	5149
Media views	6409	Media search (5591 views); M333576 (818 views);
Matrix views	3545	Matrix landing page (2981 views); A review of Shamosuchus and Paralligator (Crocodyliformes, Neosuchia) from the Cretaceous of Asia (564 views);
Specimen list	3161
Taxon list	22416
Documents list	2404
Bibliography	1181

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	678
Matrix downloads	146	A review of Shamosuchus and Paralligator (Crocodyliformes, Neosuchia) from the Cretaceous of Asia (146 downloads);
Document downloads	67	MrBayes Results Treefile (10 downloads); MrBayes analysis file (9 downloads); TNT file (18 downloads); Isisfordia Primary Analysis TNT file (8 downloads); TNT constraint file (12 downloads); Incipient procoely Tree File (1 download); Derive palate and procoely TNT file (1 download); Derived palate TNT file (2 downloads); Derived procoely TNT file (2 downloads); Incipient procoely TNT file (3 downloads); Derived palate strict consensus tree file (1 download);
Project downloads	462
Media downloads	3	M333576 (3 downloads);