Project 1238: K. T. Stilson, S. S. B. Hopkins, E. B. Davis. 2014. 50 Million Years of Severe Osteopathology in Rhinocerotidae, Meeting Abstract. Society for Integrative and Comparative Biology.
Specimen: † Menoceras arikarense (AMNH:144630)
View: Phalanges

Abstract

Skeletal pathologies in the fossil record are commonly considered indicators of an individual animal’s life history and development, and not reflections of processes related to all or part of a lineage’s history. Individual elements of many extinct and extant rhinocerotids display arthritis-like osteopathologies. The proportion of severe osteopathology increases over the last 50 million years from about 30% of all elements in a taxon from the Eocene showing some form of osteopathology to 100% of all elements in modern taxa. The Rhinocerotidae preserved in the fossil record do not represent a direct lineage, but a group of closely related organisms that present a study system for pathological evolution and development. Osteopathology is examined here in six extinct taxa spanning 50 Ma (Hyrachyus eximus, Diceratherium sp., Trigonias osborni, Menoceras arikarensus, Aphelops sp., and Teleoceras sp.) as well as the five living species of rhinoceroses. Seven pathological indicators (overgrowth, lipping, remodeling, erosions, worn articular surface, and variable foramen shape and size) were scored for each element on a scale of 1-4 and mapped on a phylogenetic tree using Mesquite to establish ancestral and lineage expression of osteopathology (i.e., that present in the common ancestor and also that present as successive points along the tree). We estimated mass from femur length and compared resulting values with the literature. We measured cursoriality using the established proxies of metatarsal length over femur length (MT/F), and total hind-leg length (femur + tibia + metatarsal). Estimated mass increases from 50 Ma to the present and correlates with the increased expression of osteopathology while cursoriality, the other trait examined as a possible contributing factor, did not. This degree of osteopatholgical expression may be an example of evolutionary compromise.


Read the article »

Project DOI: 10.7934/P1238, http://dx.doi.org/10.7934/P1238
This project containsMatrices
  • 1297 Media
  • 1 Matrix
  • 7 Documents
  • 51 Taxa
  • 322 Specimens
  • 72 Characters
Total size of project's media files: 2G

Download Project SDD File
Total scored cells: 2724
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 72
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 183
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:72/0
Currently Viewing:
MorphoBank Project 1238

    Authors' Institutions

    • University of Oregon

    • University of Texas at Austin



    Members

    member name taxa specimens media media
    notes
    chars character
    comms
       
    notes  media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
    comms
        
    notes  medialabels
    rules
    Kelsey Stilson
    Project Administrator
    Last logged in 05/05/15
    513221297777200002724
    (2724, 0, 0)
    02664000
    MorphoBank Curator
    Full membership
    Last logged in 05/08/15
    0000000000
    (0, 0, 0)
    00000


    Taxonomic Overview for Matrix 'M2867' (45 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] † Hyrachyus
    Last Modified in 01/16/15
    072000002
    [2] † Egysodontinae
    Last Modified in 01/16/15
    1953000002
    [3] † Hyracodontinae
    Last Modified in 01/16/15
    369000002
    [4] † Teletaceras
    Last Modified in 01/16/15
    2151000002
    [5] † Penetrigonas
    Last Modified in 01/16/15
    2250000002
    [6] † Amphicaenopus
    Last Modified in 01/16/15
    2151000002
    [7] † Trigonias osborni
    Last Modified in 08/25/14
    468000002
    [8] † Subhyracodon
    Last Modified in 01/16/15
    864000002
    [9] † Diceratherium
    Last Modified in 01/16/15
    369000002
    [10] † Menoceras arikarense
    Last Modified in 01/16/15
    072000002
    [11] † Floridaceras
    Last Modified in 01/16/15
    567000002
    [13] † Peraceras
    Last Modified in 01/16/15
    1260000002
    [14] † Aphelops mutilis
    Last Modified in 08/25/14
    468000002
    [15] † Teleoceras hicksi
    Last Modified in 08/25/14
    567000002
    [16] † Ronzotherium
    Last Modified in 01/16/15
    1359000002
    [17] † Mesaceratherium
    Last Modified in 01/16/15
    3141000002
    [18] † Protaceratherium
    Last Modified in 01/16/15
    765000002
    [19] † Pleuroceros
    Last Modified in 01/16/15
    1458000002
    [20] † Diceratherium niobrarense
    Last Modified in 08/25/14
    765000002
    [21] † Brachypotherium
    Last Modified in 01/16/15
    765000002
    [22] † Prosantorhinus
    Last Modified in 01/16/15
    666000002
    [23] † Alicornops
    Last Modified in 01/16/15
    270000002
    [24] † Hoploaceratherium
    Last Modified in 01/16/15
    270000002
    [25] † Aceratjeroi
    Last Modified in 01/16/15
    666000002
    [26] † Lartetotherium
    Last Modified in 01/16/15
    468000002
    [27] † L.schleiermacheri
    Last Modified in 01/16/15
    369000002
    [28] † Stephanorhinus
    Last Modified in 01/16/15
    1062000002
    [29] † Coleodonta
    Last Modified in 01/16/15
    468000002
    [30] † Hispanotherium
    Last Modified in 01/16/15
    2052000002
    [31] † Begertherium
    Last Modified in 01/16/15
    1260000002
    [32] † Beliajevina
    Last Modified in 01/16/15
    1260000002
    [33] † Iranotherium
    Last Modified in 01/16/15
    2745000002
    [34] † Elasmotherium
    Last Modified in 01/16/15
    2151000002
    [35] † Ninxiatherium
    Last Modified in 01/16/15
    3933000002
    [36] † Acerorhinus
    Last Modified in 01/16/15
    963000002
    [37] † Chilotherium
    Last Modified in 01/16/15
    1062000002
    [38] † Chilotheridium
    Last Modified in 01/16/15
    1953000002
    [39] † Aprotodon
    Last Modified in 01/16/15
    1557000002
    [40] † Gaindatherium
    Last Modified in 01/16/15
    2448000002
    [41] † Punjabitherium
    Last Modified in 01/16/15
    3240000002
    [42] Rhinoceros
    Last Modified in 09/11/14
    369000002
    [43] Dicerorhinus sumatrensis
    Last Modified in 08/27/14
    567000002
    [44] Diceros bicornis
    Last Modified in 08/25/14
    171000002
    [45] Ceratotherium simum
    Last Modified in 08/25/14
    171000002
    [46] † Paradiceros
    Last Modified in 01/16/15
    2349000002


    Project views

    type number of views Individual items viewed (where applicable)
    Total project views184554
    Project overview3635
    Media views66342Media search (4885 views); M336849 (404 views); M336850 (444 views); M336851 (535 views); M336852 (409 views); M336853 (455 views); M336854 (409 views); M336855 (407 views); M336856 (442 views); M336857 (400 views); M336858 (421 views); M336860 (405 views); M336859 (444 views); M336862 (425 views); M336861 (423 views); M336863 (428 views); M336864 (404 views); M336868 (418 views); M336865 (403 views); M336870 (482 views); M336866 (403 views); M336873 (417 views); M336869 (444 views); M336871 (423 views); M336872 (433 views); M336867 (385 views); M336874 (430 views); M336875 (435 views); M336876 (379 views); M336877 (444 views); M336878 (408 views); M336879 (437 views); M336880 (443 views); M336881 (452 views); M336882 (379 views); M336883 (373 views); M336885 (397 views); M336884 (416 views); M336886 (383 views); M336887 (433 views); M336888 (364 views); M336890 (436 views); M336891 (466 views); M336889 (418 views); M336892 (411 views); M336893 (339 views); M336894 (400 views); M336895 (392 views); M336896 (416 views); M336897 (414 views); M336898 (364 views); M336899 (376 views); M336900 (379 views); M336901 (365 views); M336902 (439 views); M336903 (371 views); M336904 (398 views); M336905 (383 views); M336906 (382 views); M336909 (417 views); M336907 (424 views); M336908 (374 views); M336910 (405 views); M336911 (368 views); M336912 (415 views); M336913 (389 views); M336917 (384 views); M336914 (377 views); M336915 (387 views); M336916 (376 views); M336918 (411 views); M336919 (410 views); M336920 (411 views); M336921 (560 views); M336923 (418 views); M336922 (399 views); M336924 (437 views); M336925 (384 views); M336926 (431 views); M336927 (414 views); M336928 (423 views); M336929 (370 views); M336930 (378 views); M336931 (392 views); M336932 (351 views); M336933 (429 views); M336934 (433 views); M336935 (393 views); M336936 (399 views); M336937 (392 views); M336938 (462 views); M336939 (400 views); M336940 (398 views); M336941 (370 views); M336942 (386 views); M336943 (402 views); M336944 (396 views); M336945 (398 views); M336946 (395 views); M336947 (441 views); M336948 (428 views); M336949 (137 views); M336950 (145 views); M336951 (113 views); M336952 (171 views); M336953 (142 views); M336954 (119 views); M336955 (136 views); M336956 (108 views); M336957 (104 views); M336958 (138 views); M336959 (145 views); M336960 (103 views); M336961 (105 views); M336962 (149 views); M336963 (117 views); M336964 (113 views); M336965 (143 views); M336966 (127 views); M336967 (154 views); M336968 (126 views); M336969 (152 views); M336970 (130 views); M336971 (157 views); M336972 (146 views); M336973 (164 views); M336974 (107 views); M336975 (140 views); M336976 (145 views); M336977 (139 views); M336978 (147 views); M336979 (148 views); M336980 (128 views); M336981 (146 views); M336982 (155 views); M336983 (150 views); M336984 (141 views); M336985 (145 views); M336986 (156 views); M336987 (113 views); M336988 (133 views); M336989 (196 views); M336990 (129 views); M336991 (171 views); M336992 (116 views); M336993 (116 views); M336994 (132 views); M336995 (160 views); M336996 (115 views); M336997 (107 views); M336998 (114 views); M336999 (160 views); M337000 (143 views); M337001 (148 views); M337002 (129 views); M337003 (103 views); M337004 (140 views); M337005 (144 views); M337006 (120 views); M337007 (147 views); M337008 (139 views); M337009 (142 views); M337010 (115 views); M337011 (157 views); M337012 (136 views); M337013 (136 views); M337014 (145 views); M337015 (119 views); M337016 (134 views); M337017 (120 views); M337021 (123 views); M337022 (151 views); M337023 (109 views); M337024 (147 views); M337025 (113 views); M337026 (147 views); M337027 (134 views); M337028 (148 views); M337029 (146 views); M337030 (147 views); M337035 (150 views); M337037 (145 views); M337038 (136 views); M337039 (146 views); M337040 (149 views); M337041 (105 views); M337042 (115 views); M337043 (154 views); M337044 (146 views); M337045 (105 views); M337046 (124 views); M337047 (140 views); M337048 (155 views); M337049 (141 views); M337050 (104 views); M337051 (111 views); M337052 (134 views); M337053 (155 views); M337054 (138 views); M337055 (108 views); M337056 (165 views); M337057 (137 views); M337058 (111 views); M337059 (134 views); M337060 (100 views); M337061 (135 views); M337062 (144 views); M337063 (177 views); M337064 (138 views); M337065 (146 views); M337066 (139 views); M337067 (156 views); M337068 (140 views); M337069 (153 views); M337070 (146 views); M337071 (145 views); M337072 (144 views); M337073 (148 views); M337020 (115 views); M337033 (142 views); M337036 (107 views); M337019 (158 views); M337031 (135 views); M337018 (128 views); M337032 (114 views); M337034 (288 views); M337362 (4 views); M337364 (2 views); M337347 (4 views); M365765 (3 views); M337146 (9 views); M337567 (2 views); M337532 (2 views); M337508 (3 views); M337515 (2 views); M338018 (2 views); M338137 (2 views); M365724 (2 views); M337087 (11 views); M337094 (9 views); M337109 (19 views); M337583 (3 views); M337197 (3 views); M337548 (2 views); M337142 (7 views); M337089 (8 views); M337542 (2 views); M337494 (2 views); M337718 (3 views); M337141 (8 views); M337124 (7 views); M337076 (10 views); M337106 (10 views); M337571 (2 views); M337572 (2 views); M337770 (3 views); M337704 (3 views); M337758 (3 views); M337765 (3 views); M337683 (3 views); M337745 (2 views); M337751 (3 views); M337757 (2 views); M337533 (2 views); M337744 (3 views); M337130 (8 views); M337693 (3 views); M337122 (7 views); M337507 (9 views); M337527 (2 views); M337131 (7 views); M337511 (12 views); M337495 (2 views); M337117 (11 views); M337502 (2 views); M337681 (3 views); M337088 (8 views); M337086 (6 views); M337145 (8 views); M337712 (3 views); M337561 (1 view); M337090 (6 views); M337739 (3 views); M337526 (2 views); M337706 (3 views); M337498 (2 views); M337738 (4 views); M337110 (9 views); M337749 (3 views); M337539 (3 views); M337102 (8 views); M337125 (9 views); M337535 (2 views); M337563 (2 views); M337582 (3 views); M337725 (3 views); M337760 (3 views); M337574 (2 views); M337148 (8 views); M337084 (8 views); M337524 (2 views); M337759 (2 views); M337496 (2 views); M337578 (2 views); M337697 (14 views); M337529 (2 views); M337707 (3 views); M337111 (7 views); M337675 (2 views); M337082 (6 views); M337116 (7 views); M337555 (1 view); M337702 (3 views); M337147 (6 views); M337724 (3 views); M337699 (3 views); M337105 (6 views); M337521 (3 views); M337505 (1 view); M337575 (7 views); M337119 (5 views); M337576 (2 views); M337676 (3 views); M337198 (3 views); M337501 (2 views); M337091 (7 views); M337761 (2 views); M337547 (2 views); M337771 (3 views); M337537 (2 views); M337126 (7 views); M365890 (10 views); M337715 (2 views); M337859 (3 views); M337314 (1 view); M337270 (2 views); M337731 (4 views); M337753 (3 views); M337588 (2 views); M337769 (2 views); M337736 (2 views); M337136 (11 views); M337137 (7 views); M337764 (2 views); M337756 (3 views); M337554 (2 views); M337536 (11 views); M337763 (3 views); M337589 (3 views); M337080 (10 views); M337552 (2 views); M337549 (2 views); M337689 (3 views); M337590 (1 view); M337113 (6 views); M337557 (5 views); M337682 (2 views); M337244 (6 views); M337519 (3 views); M337565 (2 views); M337695 (3 views); M337558 (3 views); M337684 (3 views); M337743 (3 views); M337512 (3 views); M337083 (13 views); M337701 (3 views); M337509 (2 views); M337592 (2 views); M337545 (2 views); M337543 (2 views); M337556 (2 views); M337135 (10 views); M337742 (3 views); M337107 (7 views); M337553 (1 view); M337133 (6 views); M337499 (2 views); M337138 (6 views); M337674 (3 views); M337092 (10 views); M337688 (2 views); M337514 (2 views); M337560 (10 views); M337685 (3 views); M337139 (8 views); M337692 (3 views); M337680 (3 views); M337129 (7 views); M337691 (3 views); M337734 (2 views); M337144 (8 views); M337534 (1 view); M337540 (1 view); M337108 (7 views); M338123 (2 views); M337726 (3 views); M337677 (3 views); M337520 (2 views); M337115 (8 views); M337748 (3 views); M337705 (3 views); M337732 (3 views); M337584 (2 views); M337750 (3 views); M337586 (2 views); M337120 (7 views); M337735 (3 views); M337755 (3 views); M337104 (7 views); M337506 (2 views); M337546 (3 views); M337708 (3 views); M337700 (3 views); M337577 (2 views); M337531 (3 views); M337077 (6 views); M337746 (3 views); M337747 (4 views); M337528 (2 views); M337772 (4 views); M337716 (2 views); M337721 (3 views); M337118 (5 views); M337754 (3 views); M337711 (4 views); M337112 (5 views); M337694 (2 views); M337513 (2 views); M337551 (1 view); M337733 (2 views); M337143 (8 views); M337566 (2 views); M337768 (4 views); M337767 (3 views); M337562 (3 views); M337510 (2 views); M337581 (1 view); M337709 (3 views); M337679 (4 views); M337703 (2 views); M337569 (3 views); M337591 (2 views); M337079 (6 views); M337687 (3 views); M337714 (3 views); M337696 (2 views); M337766 (3 views); M337134 (7 views); M337723 (3 views); M337101 (7 views); M337580 (2 views); M337698 (3 views); M337097 (5 views); M337713 (3 views); M337081 (5 views); M337538 (3 views); M337095 (7 views); M337752 (3 views); M337500 (1 view); M337690 (3 views); M337103 (6 views); M337140 (5 views); M337530 (2 views); M337570 (3 views); M337132 (9 views); M337078 (5 views); M337121 (9 views); M337722 (2 views); M337100 (6 views); M337123 (9 views); M337127 (5 views); M337085 (6 views); M337504 (3 views); M337516 (2 views); M337564 (2 views); M337522 (2 views); M337550 (3 views); M337099 (5 views); M337593 (3 views); M337585 (3 views); M337573 (2 views); M337544 (2 views); M337525 (2 views); M337717 (3 views); M337727 (3 views); M337114 (5 views); M337719 (3 views); M337098 (6 views); M337686 (3 views); M337720 (3 views); M337729 (3 views); M337503 (3 views); M337517 (1 view); M337728 (3 views); M337741 (3 views); M337762 (3 views); M337096 (6 views); M337497 (4 views); M337587 (1 view); M337710 (3 views); M337518 (2 views); M337075 (6 views); M337493 (1 view); M337737 (3 views); M337579 (3 views); M337093 (7 views); M337740 (2 views); M337730 (3 views); M337523 (2 views); M337074 (7 views); M337541 (3 views); M337559 (3 views); M337678 (3 views); M337568 (3 views); M337128 (9 views); M337159 (6 views); M337241 (5 views); M337223 (5 views); M337813 (3 views); M337851 (3 views); M337224 (7 views); M337825 (5 views); M337839 (3 views); M337840 (3 views); M337191 (7 views); M337234 (3 views); M337774 (4 views); M337247 (5 views); M337177 (3 views); M337812 (4 views); M337841 (5 views); M337775 (3 views); M337790 (6 views); M337831 (3 views); M337168 (5 views); M337196 (4 views); M337218 (4 views); M337818 (4 views); M337819 (4 views); M337836 (5 views); M337180 (4 views); M337842 (5 views); M337781 (6 views); M337821 (3 views); M337233 (3 views); M337170 (4 views); M337209 (3 views); M337808 (3 views); M337183 (7 views); M337175 (3 views); M337810 (4 views); M337184 (3 views); M337230 (4 views); M337221 (3 views); M337829 (3 views); M337163 (3 views); M337227 (5 views); M337199 (4 views); M337158 (5 views); M337780 (4 views); M337187 (4 views); M337868 (4 views); M337174 (3 views); M337212 (6 views); M337181 (3 views); M337235 (3 views); M337166 (4 views); M337161 (4 views); M337822 (4 views); M337172 (3 views); M337160 (4 views); M337205 (2 views); M337213 (3 views); M337857 (4 views); M337182 (5 views); M337794 (3 views); M337176 (3 views); M337844 (5 views); M337860 (5 views); M337777 (4 views); M337834 (3 views); M337200 (4 views); M337806 (18 views); M337867 (3 views); M337815 (4 views); M337204 (2 views); M337850 (5 views); M337150 (4 views); M337835 (4 views); M337206 (6 views); M337797 (3 views); M337802 (4 views); M337217 (4 views); M337248 (4 views); M337171 (4 views); M337236 (2 views); M337789 (3 views); M337210 (2 views); M337186 (3 views); M337189 (2 views); M337238 (6 views); M337833 (3 views); M337228 (5 views); M337871 (3 views); M337805 (3 views); M337215 (5 views); M337231 (3 views); M337820 (8 views); M337157 (3 views); M337786 (5 views); M337796 (4 views); M337214 (3 views); M337848 (3 views); M337870 (5 views); M337219 (3 views); M337865 (5 views); M337852 (4 views); M337847 (4 views); M337787 (5 views); M337245 (3 views); M337827 (3 views); M337156 (4 views); M337192 (3 views); M337801 (4 views); M337778 (4 views); M337800 (3 views); M337866 (3 views); M337222 (6 views); M337803 (4 views); M337843 (3 views); M337856 (4 views); M337791 (4 views); M337167 (2 views); M337773 (3 views); M337798 (5 views); M337165 (3 views); M337809 (3 views); M337152 (2 views); M337782 (4 views); M337832 (3 views); M337788 (3 views); M337830 (4 views); M337854 (4 views); M337826 (3 views); M337845 (5 views); M337164 (3 views); M337816 (4 views); M337194 (3 views); M337784 (3 views); M337162 (2 views); M337793 (4 views); M337799 (4 views); M337225 (3 views); M337785 (5 views); M337838 (4 views); M337804 (4 views); M337193 (3 views); M337837 (3 views); M337811 (5 views); M337216 (4 views); M337173 (2 views); M337792 (6 views); M337849 (4 views); M337211 (3 views); M337858 (4 views); M337226 (4 views); M337190 (6 views); M337828 (4 views); M337864 (4 views); M337779 (4 views); M337169 (3 views); M337861 (5 views); M337855 (5 views); M337202 (3 views); M337229 (3 views); M337201 (3 views); M337237 (3 views); M337149 (3 views); M337240 (3 views); M337207 (3 views); M337185 (4 views); M337239 (3 views); M337179 (4 views); M337862 (4 views); M337151 (2 views); M337783 (4 views); M337155 (2 views); M337246 (4 views); M337178 (2 views); M337188 (5 views); M337208 (3 views); M337195 (3 views); M337823 (3 views); M337242 (6 views); M337824 (4 views); M337872 (4 views); M337203 (5 views); M337153 (7 views); M337846 (5 views); M337776 (5 views); M337817 (6 views); M337863 (3 views); M337154 (3 views); M337807 (4 views); M337220 (12 views); M337853 (4 views); M337814 (4 views); M337795 (4 views); M337232 (3 views); M337869 (3 views); M337243 (2 views); M337313 (3 views); M337892 (4 views); M337907 (2 views); M337271 (2 views); M337965 (2 views); M337946 (2 views); M337945 (1 view); M337905 (5 views); M337280 (2 views); M337943 (4 views); M337253 (1 view); M337286 (2 views); M337931 (2 views); M337256 (2 views); M337958 (2 views); M337932 (2 views); M337267 (2 views); M339644 (4 views); M365720 (5 views); M365723 (3 views); M365736 (3 views); M365831 (2 views); M365783 (4 views); M365878 (2 views); M365944 (3 views); M365952 (3 views); M337896 (2 views); M337249 (3 views); M337885 (4 views); M337310 (3 views); M337890 (3 views); M337260 (2 views); M337926 (2 views); M337341 (8 views); M337351 (2 views); M337353 (1 view); M337936 (2 views); M337912 (4 views); M337875 (3 views); M337290 (2 views); M337961 (2 views); M337354 (2 views); M337275 (2 views); M337261 (2 views); M337324 (1 view); M337937 (1 view); M337901 (1 view); M337315 (3 views); M337340 (2 views); M337951 (2 views); M337305 (4 views); M337923 (3 views); M337955 (2 views); M337970 (1 view); M337346 (2 views); M337884 (1 view); M337915 (1 view); M337880 (1 view); M337281 (2 views); M337924 (1 view); M337917 (2 views); M337956 (3 views); M337278 (3 views); M337909 (3 views); M337287 (2 views); M337952 (2 views); M337877 (3 views); M337333 (1 view); M337913 (1 view); M337322 (1 view); M337971 (4 views); M337891 (7 views); M337895 (3 views); M337360 (2 views); M337361 (2 views); M338034 (2 views); M338130 (2 views); M337409 (2 views); M337405 (2 views); M338122 (2 views); M337484 (2 views); M337372 (2 views); M338013 (2 views); M338104 (2 views); M338014 (2 views); M338129 (2 views); M337406 (2 views); M338121 (1 view); M338011 (2 views); M337444 (3 views); M337438 (3 views); M337380 (3 views); M337441 (2 views); M338023 (2 views); M337374 (3 views); M337442 (3 views); M337998 (3 views); M337366 (3 views); M337485 (2 views); M338001 (2 views); M337982 (2 views); M337378 (2 views); M337413 (2 views); M337369 (2 views); M338125 (1 view); M337449 (2 views); M337985 (1 view); M337483 (2 views); M338128 (3 views); M337480 (1 view); M337446 (3 views); M337914 (3 views); M337911 (5 views); M337960 (3 views); M337894 (2 views); M337276 (1 view); M337967 (5 views); M337332 (1 view); M337950 (2 views); M337250 (4 views); M337356 (3 views); M337920 (4 views); M337969 (1 view); M337882 (3 views); M337328 (1 view); M337874 (1 view); M337925 (1 view); M337282 (1 view); M337929 (2 views); M337294 (2 views); M337264 (2 views); M337296 (2 views); M337268 (1 view); M337283 (4 views); M337292 (1 view); M337908 (6 views); M337269 (9 views); M337357 (2 views); M337942 (3 views); M337878 (11 views); M337949 (2 views); M337919 (4 views); M337255 (2 views); M337957 (3 views); M337953 (3 views); M337966 (2 views); M337289 (4 views); M337338 (3 views); M337265 (4 views); M337930 (2 views); M337291 (2 views); M337972 (2 views); M337910 (2 views); M337330 (3 views); M337337 (4 views); M337940 (3 views); M337879 (7 views); M337887 (2 views); M337904 (2 views); M337889 (3 views); M337954 (7 views); M337343 (2 views); M337964 (3 views); M337308 (2 views); M337881 (3 views); M337345 (2 views); M337306 (1 view); M337295 (2 views); M337934 (6 views); M337257 (1 view); M337344 (1 view); M337317 (2 views); M337947 (5 views); M337968 (2 views); M337906 (2 views); M337873 (2 views); M337285 (3 views); M337251 (3 views); M337938 (3 views); M337903 (3 views); M337918 (2 views); M337352 (3 views); M337886 (3 views); M337325 (1 view); M337334 (2 views); M337876 (3 views); M337331 (3 views); M337272 (2 views); M337277 (2 views); M337318 (8 views); M337312 (2 views); M337893 (2 views); M337254 (1 view); M337900 (3 views); M337303 (4 views); M337898 (3 views); M337311 (3 views); M337284 (2 views); M337336 (2 views); M337933 (2 views); M337320 (3 views); M337941 (4 views); M337962 (2 views); M337327 (4 views); M337916 (3 views); M337319 (2 views); M337963 (1 view); M337342 (2 views); M337321 (2 views); M337928 (3 views); M337316 (1 view); M337355 (4 views); M337922 (2 views); M337329 (2 views); M337263 (3 views); M337326 (4 views); M337902 (4 views); M337298 (2 views); M337944 (1 view); M337888 (2 views); M337921 (2 views); M337897 (7 views); M337309 (2 views); M337899 (2 views); M337307 (1 view); M337927 (2 views); M337939 (1 view); M337339 (4 views); M337252 (2 views); M337948 (2 views); M337288 (3 views); M337293 (2 views); M337297 (2 views); M337959 (3 views); M337883 (2 views); M337935 (1 view); M337258 (3 views); M337304 (2 views); M337279 (3 views); M337262 (2 views); M337266 (1 view); M337259 (4 views); M337335 (1 view); M337299 (4 views); M337978 (2 views); M337386 (3 views); M338005 (2 views); M337997 (3 views); M337407 (2 views); M337440 (2 views); M337973 (2 views); M337384 (3 views); M338124 (1 view); M337993 (1 view); M338035 (1 view); M337481 (2 views); M337425 (3 views); M337427 (3 views); M365744 (2 views); M365771 (1 view); M338162 (1 view); M338171 (2 views); M365766 (2 views); M338165 (2 views); M365832 (2 views); M338164 (2 views); M365786 (2 views); M338006 (3 views); M338139 (3 views); M337423 (3 views); M338000 (3 views); M337426 (2 views); M337391 (2 views); M337367 (2 views); M338140 (2 views); M338103 (22 views); M338111 (3 views); M338132 (1 view); M338107 (3 views); M338110 (2 views); M338109 (4 views); M338098 (2 views); M337489 (1 view); M338008 (4 views); M338138 (2 views); M338012 (2 views); M338021 (3 views); M337431 (2 views); M337376 (3 views); M337447 (3 views); M338114 (2 views); M337387 (16 views); M337365 (2 views); M337486 (2 views); M337389 (2 views); M337382 (2 views); M337479 (2 views); M337488 (3 views); M337411 (3 views); M338015 (2 views); M338108 (2 views); M338010 (2 views); M337992 (1 view); M337412 (3 views); M337383 (2 views); M337986 (2 views); M338117 (2 views); M338113 (1 view); M337363 (2 views); M338126 (2 views); M337995 (2 views); M337989 (2 views); M337368 (2 views); M337428 (2 views); M337377 (2 views); M337477 (3 views); M337439 (3 views); M338102 (4 views); M337420 (3 views); M338020 (2 views); M338116 (3 views); M338037 (3 views); M337415 (7 views); M338017 (2 views); M337429 (2 views); M337437 (2 views); M337433 (3 views); M337443 (3 views); M337975 (1 view); M338115 (1 view); M337404 (5 views); M337410 (2 views); M337492 (2 views); M337371 (2 views); M337981 (2 views); M338112 (1 view); M337453 (3 views); M338022 (1 view); M337370 (3 views); M337451 (2 views); M337487 (2 views); M337381 (2 views); M338131 (2 views); M337984 (2 views); M338133 (2 views); M337388 (3 views); M337977 (2 views); M338009 (2 views); M337988 (1 view); M337996 (1 view); M337359 (2 views); M337390 (2 views); M337490 (3 views); M337421 (2 views); M337445 (2 views); M337434 (2 views); M338100 (2 views); M337450 (2 views); M337419 (2 views); M338016 (2 views); M337414 (2 views); M338026 (2 views); M337408 (2 views); M338003 (1 view); M337991 (1 view); M337373 (2 views); M337448 (2 views); M338025 (4 views); M337358 (3 views); M338127 (2 views); M338019 (2 views); M337478 (2 views); M337424 (2 views); M337416 (2 views); M338007 (4 views); M337491 (3 views); M337385 (3 views); M337375 (3 views); M338183 (2 views); M365760 (3 views); M365788 (1 view); M365774 (1 view); M365754 (1 view); M365728 (2 views); M338028 (3 views); M338106 (2 views); M338101 (2 views); M337418 (3 views); M337974 (2 views); M337990 (2 views); M337452 (2 views); M337987 (2 views); M337430 (2 views); M338120 (2 views); M337482 (2 views); M337422 (2 views); M338004 (2 views); M338036 (2 views); M338105 (2 views); M337983 (2 views); M337432 (3 views); M337976 (2 views); M337436 (3 views); M337417 (3 views); M338119 (2 views); M338118 (1 view); M337979 (2 views); M338134 (3 views); M337999 (8 views); M338024 (2 views); M337435 (3 views); M338002 (2 views); M338099 (2 views); M337994 (2 views); M337379 (3 views); M337980 (2 views); M338143 (2 views); M365721 (2 views); M338168 (2 views); M338169 (1 view); M365787 (2 views); M338146 (2 views); M365781 (2 views); M338159 (2 views); M365729 (2 views); M338167 (2 views); M365755 (2 views); M365725 (1 view); M365756 (2 views); M365746 (1 view); M338158 (1 view); M365757 (4 views); M365769 (3 views); M338151 (1 view); M365780 (2 views); M365784 (2 views); M365777 (2 views); M338172 (1 view); M338179 (2 views); M365775 (2 views); M365763 (2 views); M338166 (2 views); M338153 (1 view); M338154 (2 views); M338155 (1 view); M338150 (2 views); M365770 (2 views); M338157 (2 views); M338170 (3 views); M365745 (2 views); M365772 (2 views); M365752 (2 views); M365790 (2 views); M365726 (2 views); M365739 (2 views); M365753 (1 view); M365773 (2 views); M365747 (2 views); M338147 (2 views); M365737 (1 view); M338141 (2 views); M365768 (2 views); M365778 (2 views); M338160 (1 view); M365751 (2 views); M365782 (1 view); M338176 (2 views); M338174 (2 views); M365740 (2 views); M338178 (2 views); M365793 (2 views); M338161 (1 view); M365759 (2 views); M338180 (2 views); M365748 (1 view); M365762 (2 views); M365789 (2 views); M365750 (1 view); M338177 (2 views); M365764 (2 views); M365785 (2 views); M338152 (2 views); M365743 (3 views); M365761 (4 views); M365742 (3 views); M365730 (1 view); M365767 (1 view); M338175 (2 views); M338163 (3 views); M338173 (1 view); M365776 (3 views); M338156 (3 views); M338182 (3 views); M365889 (2 views); M365948 (2 views); M365962 (3 views); M365918 (3 views); M365895 (3 views); M365874 (2 views); M365869 (2 views); M365906 (2 views); M365901 (1 view); M365905 (1 view); M365894 (2 views); M365945 (2 views); M365833 (2 views); M365891 (2 views); M365882 (2 views); M365923 (2 views); M365955 (2 views); M365953 (2 views); M365885 (2 views); M365958 (2 views); M365920 (1 view); M365922 (2 views); M365907 (3 views); M365954 (2 views); M365877 (2 views); M365902 (2 views); M365868 (2 views); M365883 (2 views); M365926 (2 views); M365854 (2 views); M365896 (2 views); M365909 (2 views); M365947 (2 views); M365853 (3 views); M365884 (2 views); M365888 (2 views); M365951 (2 views); M365899 (2 views); M365927 (2 views); M365857 (2 views); M365893 (2 views); M365921 (2 views); M365897 (2 views); M365886 (3 views); M365879 (2 views); M365866 (2 views); M365880 (1 view); M365898 (1 view); M365861 (2 views); M365862 (2 views); M365912 (2 views); M365834 (2 views); M365855 (2 views); M365856 (1 view); M365881 (2 views); M365892 (2 views); M365928 (2 views); M365887 (2 views); M365858 (2 views); M365961 (1 view); M365916 (2 views); M365867 (3 views); M365871 (2 views); M365956 (2 views); M365860 (2 views); M365900 (2 views); M365873 (3 views); M365949 (3 views); M365864 (2 views); M365950 (2 views); M365960 (1 view); M365915 (1 view); M365957 (1 view); M365872 (1 view); M365919 (2 views); M365924 (1 view); M365946 (1 view); M365859 (1 view); M365870 (2 views); M365863 (1 view); M365904 (1 view); M365908 (1 view); M365876 (1 view); M365917 (2 views); M365925 (1 view); M365911 (1 view); M365914 (2 views); M365913 (2 views); M365910 (1 view); M365875 (1 view); M365959 (1 view); M365865 (1 view); M365903 (1 view);
    Views for media list5330
    Documents list1599
    Matrix views2012Matrix landing page (1823 views); Character Data from Cerdeño 1995 (184 views); (5 views);
    Specimen list94326
    Taxon list10643
    Bibliography667




    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project354
    Document downloads7Unpruned Rhino Tree from Cerdeño 1995 for R (1 download); Mass Estimates for Extinct and Extinct Rhinocerotidae (3 downloads); Osteopathology Scores for R (2 downloads); R Code for Rhino Osteopathology Project (1 download);
    Matrix downloads8Character Data from Cerdeño 1995 (8 downloads);
    Project downloads336
    Media downloads3M365878 (1 download); M337080 (1 download); M336850 (1 download);