Project 2084: J. D. Carrillo, R. J. Asher. 2017. An exceptionally well preserved skeleton Thomashuxleya externa (Mammalia, Notoungulata), from the Eocene of Patagonia, Argentina. Palaeontologia electronica. 20.2.34A:1-33.
Abstract
We describe one of the oldest notoungulate skeletons with associated craniodental and postcranial elements: Thomashuxleya externa (Isotemnidae) from Cañadón Vaca in Patagonia, Argentina (Vacan subage of the Casamayoran SALMA, middle Eocene). We provide body mass estimates given by different elements of the skeleton, describe the bone histology, and study its phylogenetic position. We note differences in the scapulae, humerii, ulnae and radii of the new specimen in comparison with other specimens previously referred to this taxon. We estimate a body mass of 84 ± 24.2 kg, showing that notoungulates had acquired a large body mass by the middle Eocene. Bone histology shows that the new specimen was skeletally mature. The new material supports the placement of Thomashuxleya as an early, divergent member of Toxodontia. Among placentals, our phylogenetic analysis of a combined DNA, collagen, and morphology matrix favor only a limited number of possible phylogenetic relationships, but cannot yet arbitrate between potential affinities with Afrotheria or Laurasiatheria. With no constraint, maximum parsimony supports Thomashuxleya and Carodnia with Afrotheria. With Notoungulata and Litopterna constrained as monophyletic (including Macrauchenia and Toxodon known for collagens), these clades are reconstructed on the stem to Euungulata (i.e., Perissodactyla and Artiodactyla). Unconstrained, Bayesian analysis weakly supports the possibility that Thomashuxleya is a stem xenarthran; with Notoungulata and Litopterna constrained as monophyletic, the two clades are recovered as sister to Perissodactyla. Anatomical data sampled thus far for Thomashuxleya, combined with collagen amino acids for Pleistocene meridiungulates, substantially limit the number of possible affinities for endemic South American species among mammals, although ambiguity still remains.Read the article »
Project DOI: 10.7934/P2084, http://dx.doi.org/10.7934/P2084
This project contains | Matrices |
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Download Project SDD File ![]() | Total scored cells: 257229 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 3806 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 8991 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:3792/14 |
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MorphoBank Project 2084
MorphoBank Project 2084
- Creation Date:
15 December 2014 - Publication Date:
19 July 2017 - Project views: 25567
- Media downloads: 2
- Matrix downloads: 79
Authors' Institutions
- Paläontologisches Museum, Zürich
- University of Cambridge
- University of Zurich (Universität Zürich)
Members
member name | taxa ![]() |
specimens ![]() |
media ![]() | chars ![]() | character
| cell scorings (scored, NPA, "-") | cell
| rules ![]() | ||||||||
Juan David Carrillo Project Administrator | 154 | 1 | 1 | 3806 | 0 | 0 | 0 | 257229 (175656, 0, 81573) | 0 | 0 | 0 | |||||
Robert Asher Full membership ![]() | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M22531' (86 Taxa)
taxon | unscored cells |
scored cells ![]() |
no cell support ![]() |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Thomashuxleya externa Last Modified in 07/19/17 | 1653 | 1500 | 1486 | 0 | 521 | 0 | 0 | 2 ![]() |
[2] † Carodnia vieirai Last Modified in 07/19/17 | 1702 | 1464 | 464 | 0 | 494 | 0 | 0 | 2 ![]() |
[3] † Protolipterna ellipsodontoides Last Modified in 07/19/17 | 2366 | 981 | 332 | 0 | 314 | 0 | 0 | 2 ![]() |
[4] † Didolodus multicuspis Last Modified in 07/19/17 | 2482 | 891 | 891 | 0 | 287 | 0 | 0 | 2 ![]() |
[5] † Morganucodon oehleri Last Modified in 07/19/17 | 1303 | 1328 | 1285 | 0 | 1032 | 0 | 0 | 2 ![]() |
[6] † Morganucodon watsoni Last Modified in 07/19/17 | 747 | 1720 | 1716 | 0 | 1196 | 0 | 0 | 2 ![]() |
[7] † Haldanodon exspectatus Last Modified in 07/19/17 | 1073 | 1601 | 1573 | 0 | 992 | 0 | 0 | 2 ![]() |
[8] Ornithorhynchus anatinus Last Modified in 12/16/14 | 53 | 1930 | 1895 | 0 | 1682 | 0 | 0 | 2 ![]() |
[9] Tachyglossus aculeatus Last Modified in 12/16/14 | 19 | 1546 | 1344 | 0 | 2099 | 0 | 0 | 2 ![]() |
[10] † Zhangheotherium quinquecuspidens Last Modified in 07/19/17 | 816 | 1609 | 1586 | 0 | 1238 | 0 | 0 | 2 ![]() |
[11] † Henkelotherium guimarotae Last Modified in 07/19/17 | 1872 | 1083 | 1083 | 0 | 705 | 0 | 0 | 2 ![]() |
[12] † Pucadelphys andinus Last Modified in 07/19/17 | 800 | 2066 | 2030 | 0 | 801 | 0 | 0 | 2 ![]() |
[13] Didelphis virginiana Last Modified in 12/16/14 | 11 | 2743 | 593 | 0 | 922 | 0 | 0 | 2 ![]() |
[14] Dromiciops gliroides Last Modified in 12/16/14 | 302 | 2482 | 562 | 0 | 882 | 0 | 0 | 2 ![]() |
[15] † Eomaia scansoria Last Modified in 07/19/17 | 1835 | 1413 | 1410 | 0 | 412 | 0 | 0 | 2 ![]() |
[16] † Maelestes gobiensis Last Modified in 07/19/17 | 1690 | 1454 | 1229 | 0 | 518 | 0 | 0 | 2 ![]() |
[17] Dasypus novemcinctus Last Modified in 12/16/14 | 34 | 1836 | 600 | 0 | 1804 | 0 | 0 | 2 ![]() |
[18] † Hapalops elongatus Last Modified in 07/19/17 | 463 | 1519 | 10 | 0 | 1706 | 0 | 0 | 2 ![]() |
[19] Tamandua tetradactyla Last Modified in 12/16/14 | 24 | 1729 | 83 | 0 | 1917 | 0 | 0 | 2 ![]() |
[20] Manis pentadactyla Last Modified in 12/16/14 | 60 | 1590 | 785 | 0 | 2019 | 0 | 0 | 2 ![]() |
[21] † Metacheiromys marshi Last Modified in 07/19/17 | 1912 | 1424 | 2 | 0 | 324 | 0 | 0 | 2 ![]() |
[22] † Ukhaatherium nessovi Last Modified in 07/19/17 | 1650 | 1481 | 1358 | 0 | 530 | 0 | 0 | 2 ![]() |
[23] † Zalambdalestes lechei Last Modified in 07/19/17 | 1463 | 1677 | 1637 | 0 | 526 | 0 | 0 | 2 ![]() |
[24] † Leptictis dakotensis Last Modified in 07/19/17 | 534 | 2494 | 2457 | 0 | 634 | 0 | 0 | 2 ![]() |
[25] Solenodon paradoxus Last Modified in 12/16/14 | 31 | 2758 | 2373 | 0 | 891 | 0 | 0 | 2 ![]() |
[26] Talpa europaea Last Modified in 12/16/14 | 89 | 2583 | 2445 | 0 | 1000 | 0 | 0 | 2 ![]() |
[27] Erinaceus europaeus Last Modified in 12/16/14 | 37 | 2665 | 2630 | 0 | 968 | 0 | 0 | 2 ![]() |
[28] Sorex araneus Last Modified in 12/16/14 | 525 | 1941 | 1896 | 0 | 1197 | 0 | 0 | 2 ![]() |
[29] Amblysomus hottentotus Last Modified in 12/16/14 | 354 | 2332 | 2004 | 0 | 987 | 0 | 0 | 2 ![]() |
[30] Echinops telfairi Last Modified in 12/16/14 | 314 | 2359 | 2209 | 0 | 993 | 0 | 0 | 2 ![]() |
[31] Rhynchocyon cirnei Last Modified in 12/16/14 | 50 | 2708 | 2542 | 0 | 910 | 0 | 0 | 2 ![]() |
[32] Procavia capensis Last Modified in 07/19/17 | 51 | 2807 | 2487 | 0 | 828 | 0 | 0 | 2 ![]() |
[33] Loxodonta africana Last Modified in 12/16/14 | 69 | 2194 | 2124 | 0 | 1398 | 0 | 0 | 2 ![]() |
[34] Trichechus manatus Last Modified in 12/16/14 | 193 | 1820 | 1765 | 0 | 1660 | 0 | 0 | 2 ![]() |
[35] † Moeritherium lyonsi Last Modified in 07/19/17 | 1453 | 1543 | 1452 | 0 | 665 | 0 | 0 | 2 ![]() |
[36] Galeopterus variegatus Last Modified in 12/16/14 | 348 | 2555 | 2319 | 0 | 762 | 0 | 0 | 2 ![]() |
[37] Cynocephalus volans Last Modified in 12/16/14 | 94 | 2780 | 15 | 0 | 796 | 0 | 0 | 2 ![]() |
[38] Ptilocercus lowii Last Modified in 12/16/14 | 330 | 2693 | 770 | 0 | 642 | 0 | 0 | 2 ![]() |
[39] Tupaia glis Last Modified in 12/16/14 | 41 | 2913 | 925 | 0 | 713 | 0 | 0 | 2 ![]() |
[40] Homo sapiens Last Modified in 12/16/14 | 29 | 2762 | 2509 | 0 | 932 | 0 | 0 | 2 ![]() |
[41] Lemur catta catta Last Modified in 12/16/14 | 115 | 2808 | 721 | 0 | 742 | 0 | 0 | 2 ![]() |
[42] Saimiri sciureus Last Modified in 12/16/14 | 88 | 2717 | 768 | 0 | 863 | 0 | 0 | 2 ![]() |
[43] Tarsius syrichta Last Modified in 12/16/14 | 277 | 2472 | 172 | 0 | 912 | 0 | 0 | 2 ![]() |
[44] † Notharctus tenebrosus Last Modified in 07/19/17 | 626 | 2420 | 1142 | 0 | 620 | 0 | 0 | 2 ![]() |
[45] † Protungulatum donnae Last Modified in 07/19/17 | 2345 | 1009 | 987 | 0 | 306 | 0 | 0 | 2 ![]() |
[46] † Apheliscus insidiosus Last Modified in 07/19/17 | 2229 | 1027 | 417 | 0 | 404 | 0 | 0 | 2 ![]() |
[47] † Phenacodus intermedius Last Modified in 07/19/17 | 502 | 2535 | 1420 | 0 | 624 | 0 | 0 | 2 ![]() |
[48] † Hyopsodus paulus Last Modified in 07/19/17 | 1018 | 1996 | 1992 | 0 | 650 | 0 | 0 | 2 ![]() |
[49] Felis silvestris Last Modified in 12/16/14 | 17 | 2521 | 2511 | 0 | 1127 | 0 | 0 | 2 ![]() |
[50] Canis lupus familiaris Last Modified in 12/16/14 | 19 | 2879 | 2863 | 0 | 767 | 0 | 0 | 2 ![]() |
[51] † Vulpavus ovatus Last Modified in 07/19/17 | 1545 | 1625 | 2 | 0 | 493 | 0 | 0 | 2 ![]() |
[52] † Vulpavus profectus Last Modified in 07/19/17 | 1233 | 1890 | 2 | 0 | 540 | 0 | 0 | 2 ![]() |
[53] † Sinopa rapax Last Modified in 07/19/17 | 1318 | 1850 | 1842 | 0 | 496 | 0 | 0 | 2 ![]() |
[54] Oryctolagus cuniculus Last Modified in 12/16/14 | 128 | 2431 | 2425 | 0 | 1102 | 0 | 0 | 2 ![]() |
[55] † Dawsonolagus antiquus Last Modified in 07/19/17 | 2514 | 610 | 607 | 0 | 536 | 0 | 0 | 2 ![]() |
[56] † Gomphos elkema Last Modified in 07/19/17 | 910 | 1892 | 1 | 0 | 858 | 0 | 0 | 2 ![]() |
[57] † Rhombomylus turpanensis Last Modified in 07/19/17 | 550 | 2132 | 2103 | 0 | 996 | 0 | 0 | 2 ![]() |
[58] † Tribosphenomys minutus Last Modified in 07/19/17 | 1991 | 996 | 977 | 0 | 685 | 0 | 0 | 2 ![]() |
[59] † Paramys delicatus Last Modified in 07/19/17 | 676 | 1975 | 1665 | 0 | 1021 | 0 | 0 | 2 ![]() |
[60] † Cocomys lingchaensis Last Modified in 07/19/17 | 1376 | 1376 | 1353 | 0 | 917 | 0 | 0 | 2 ![]() |
[61] Ictidomys tridecemlineatus Last Modified in 12/16/14 | 300 | 2293 | 2211 | 0 | 1073 | 0 | 0 | 2 ![]() |
[62] Castor canadensis Last Modified in 12/16/14 | 79 | 2414 | 2319 | 0 | 1188 | 0 | 0 | 2 ![]() |
[63] Cavia porcellus Last Modified in 12/16/14 | 13 | 2246 | 2196 | 0 | 1420 | 0 | 0 | 2 ![]() |
[64] Rattus norvegicus Last Modified in 12/16/14 | 4 | 2201 | 1886 | 0 | 1466 | 0 | 0 | 2 ![]() |
[65] Orycteropus afer Last Modified in 12/16/14 | 296 | 1867 | 1400 | 0 | 1502 | 0 | 0 | 2 ![]() |
[66] Equus caballus Last Modified in 12/16/14 | 78 | 2709 | 2178 | 0 | 883 | 0 | 0 | 2 ![]() |
[67] Sus scrofa Last Modified in 12/16/14 | 204 | 2617 | 2017 | 0 | 854 | 0 | 0 | 2 ![]() |
[68] Tursiops truncatus Last Modified in 12/16/14 | 95 | 1163 | 836 | 0 | 2410 | 0 | 0 | 2 ![]() |
[69] Hippopotamus amphibius Last Modified in 12/16/14 | 174 | 2549 | 2181 | 0 | 942 | 0 | 0 | 2 ![]() |
[70] Lama glama Last Modified in 12/16/14 | 161 | 2428 | 2034 | 0 | 1074 | 0 | 0 | 2 ![]() |
[71] Bos taurus Last Modified in 12/16/14 | 77 | 2579 | 2173 | 0 | 1010 | 0 | 0 | 2 ![]() |
[72] † Mesohippus bairdi Last Modified in 07/19/17 | 740 | 2242 | 1939 | 0 | 682 | 0 | 0 | 2 ![]() |
[73] Caperea marginata Last Modified in 12/16/14 | 348 | 834 | 701 | 0 | 2486 | 0 | 0 | 2 ![]() |
[74] † Basilosaurus cetoides Last Modified in 07/19/17 | 1703 | 1040 | 815 | 0 | 917 | 0 | 0 | 2 ![]() |
[75] † Mesonyx obtusidens Last Modified in 07/19/17 | 1093 | 1729 | 1447 | 0 | 838 | 0 | 0 | 2 ![]() |
[76] † Archaeotherium mortoni Last Modified in 07/19/17 | 839 | 1959 | 1754 | 0 | 864 | 0 | 0 | 2 ![]() |
[77] † Artiocetus clavis Last Modified in 07/19/17 | 2450 | 750 | 679 | 0 | 461 | 0 | 0 | 2 ![]() |
[78] † Rodhocetus balochistanensis Last Modified in 07/19/17 | 3196 | 386 | 339 | 0 | 78 | 0 | 0 | 2 ![]() |
[79] Pteropus giganteus Last Modified in 12/16/14 | 51 | 2165 | 1072 | 0 | 1445 | 0 | 0 | 2 ![]() |
[80] Rhinopoma hardwickii Last Modified in 12/16/14 | 347 | 2342 | 1291 | 0 | 974 | 0 | 0 | 2 ![]() |
[81] Saccopteryx bilineata Last Modified in 12/16/14 | 356 | 2353 | 1399 | 0 | 952 | 0 | 0 | 2 ![]() |
[82] Nycteris thebaica Last Modified in 12/16/14 | 495 | 2163 | 1418 | 0 | 1002 | 0 | 0 | 2 ![]() |
[83] Myotis lucifugus Last Modified in 12/16/14 | 340 | 2257 | 1523 | 0 | 1065 | 0 | 0 | 2 ![]() |
[84] Pteronotus parnellii Last Modified in 12/16/14 | 445 | 2351 | 1687 | 0 | 868 | 0 | 0 | 2 ![]() |
[85] † Icaronycteris index Last Modified in 07/19/17 | 1297 | 1818 | 1135 | 0 | 549 | 0 | 0 | 2 ![]() |
[86] † Onychonycteris finneyi Last Modified in 07/19/17 | 1715 | 1471 | 1002 | 0 | 486 | 0 | 0 | 2 ![]() |
Taxonomic Overview for Matrix 'M24667' (68 Taxa)
taxon | unscored cells |
scored cells ![]() |
no cell support ![]() |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Leptictis Last Modified in 07/19/17 | 8 | 108 | 108 | 0 | 30 | 0 | 0 | 2 ![]() |
[2] † Phenacodus Last Modified in 07/19/17 | 7 | 102 | 102 | 0 | 37 | 0 | 0 | 2 ![]() |
[3] † Meniscotherium Last Modified in 07/19/17 | 5 | 105 | 105 | 0 | 36 | 0 | 0 | 2 ![]() |
[4] † Protolipterna Last Modified in 07/19/17 | 102 | 23 | 23 | 0 | 21 | 0 | 0 | 2 ![]() |
[5] † Proterotherium Last Modified in 07/19/17 | 9 | 93 | 93 | 0 | 44 | 0 | 0 | 2 ![]() |
[6] † Trigonostylops Last Modified in 07/19/17 | 19 | 95 | 95 | 0 | 32 | 0 | 0 | 2 ![]() |
[7] † Astrapotherium Last Modified in 07/19/17 | 9 | 99 | 98 | 0 | 39 | 0 | 0 | 2 ![]() |
[8] † Pyrotherium Last Modified in 07/19/17 | 42 | 79 | 79 | 0 | 25 | 0 | 0 | 2 ![]() |
[9] † Notostylops Last Modified in 07/19/17 | 4 | 138 | 138 | 0 | 4 | 0 | 0 | 2 ![]() |
[10] † Henricosbornia Last Modified in 07/19/17 | 101 | 43 | 43 | 0 | 2 | 0 | 0 | 2 ![]() |
[11] † Simpsonotus Last Modified in 07/19/17 | 25 | 119 | 119 | 0 | 2 | 0 | 0 | 2 ![]() |
[12] † Pleurostylodon Last Modified in 07/19/17 | 15 | 130 | 129 | 0 | 2 | 0 | 0 | 2 ![]() |
[13] † Thomashuxleya Last Modified in 07/19/17 | 35 | 109 | 109 | 0 | 2 | 0 | 0 | 2 ![]() |
[14] † Pampahippus Last Modified in 07/19/17 | 81 | 64 | 59 | 0 | 2 | 0 | 0 | 2 ![]() |
[15] † Periphragnis Last Modified in 07/19/17 | 9 | 136 | 135 | 0 | 2 | 0 | 0 | 2 ![]() |
[16] † Ryphodon Last Modified in 07/19/17 | 54 | 89 | 88 | 0 | 3 | 0 | 0 | 2 ![]() |
[17] † Puelia Last Modified in 07/19/17 | 54 | 91 | 90 | 0 | 1 | 0 | 0 | 2 ![]() |
[18] † Leontinia Last Modified in 07/19/17 | 16 | 127 | 127 | 0 | 3 | 0 | 0 | 2 ![]() |
[19] † Colpodon Last Modified in 07/19/17 | 24 | 119 | 119 | 0 | 3 | 0 | 0 | 2 ![]() |
[20] † Ancylocoelus Last Modified in 07/19/17 | 42 | 101 | 100 | 0 | 4 | 0 | 0 | 2 ![]() |
[21] † Scarittia Last Modified in 07/19/17 | 34 | 110 | 110 | 0 | 2 | 0 | 0 | 2 ![]() |
[22] † Homalodotherium Last Modified in 07/19/17 | 15 | 128 | 128 | 0 | 3 | 0 | 0 | 2 ![]() |
[23] † Asmodeus Last Modified in 07/19/17 | 68 | 75 | 75 | 0 | 3 | 0 | 0 | 2 ![]() |
[24] † Rhynchippus Last Modified in 07/19/17 | 16 | 129 | 129 | 0 | 1 | 0 | 0 | 2 ![]() |
[25] † Eurygenium Last Modified in 07/19/17 | 60 | 85 | 84 | 0 | 1 | 0 | 0 | 2 ![]() |
[26] † Pascualihippus Last Modified in 07/19/17 | 63 | 83 | 82 | 0 | 1 | 0 | 0 | 2 ![]() |
[27] † Eomorphippus Last Modified in 07/19/17 | 73 | 74 | 72 | 0 | 1 | 0 | 0 | 2 ![]() |
[28] † Morphippus Last Modified in 07/19/17 | 35 | 110 | 110 | 0 | 1 | 0 | 0 | 2 ![]() |
[29] † Argyrohippus Last Modified in 07/19/17 | 33 | 113 | 112 | 0 | 1 | 0 | 0 | 2 ![]() |
[30] † Nesodon Last Modified in 07/19/17 | 13 | 131 | 129 | 0 | 3 | 0 | 0 | 2 ![]() |
[31] † Adinotherium Last Modified in 07/19/17 | 11 | 133 | 132 | 0 | 3 | 0 | 0 | 2 ![]() |
[32] † Hoffstetterius Last Modified in 07/19/17 | 41 | 98 | 98 | 0 | 7 | 0 | 0 | 2 ![]() |
[33] † Ponanskytherium Last Modified in 07/19/17 | 38 | 100 | 100 | 0 | 8 | 0 | 0 | 2 ![]() |
[34] † Toxodon Last Modified in 07/19/17 | 40 | 98 | 98 | 0 | 8 | 0 | 0 | 2 ![]() |
[35] † Colbertia Last Modified in 07/19/17 | 29 | 117 | 116 | 0 | 1 | 0 | 0 | 2 ![]() |
[36] † Oldfieldthomasia Last Modified in 07/19/17 | 16 | 128 | 128 | 0 | 2 | 0 | 0 | 2 ![]() |
[37] † Campanorco Last Modified in 07/19/17 | 50 | 94 | 94 | 0 | 2 | 0 | 0 | 2 ![]() |
[38] † Acropithecus Last Modified in 07/19/17 | 73 | 71 | 71 | 0 | 2 | 0 | 0 | 2 ![]() |
[39] † Ultrapithecus Last Modified in 07/19/17 | 101 | 43 | 43 | 0 | 2 | 0 | 0 | 2 ![]() |
[40] † Notopithecus Last Modified in 07/19/17 | 9 | 135 | 133 | 0 | 4 | 0 | 0 | 2 ![]() |
[41] † Archaeophylus Last Modified in 07/19/17 | 125 | 18 | 18 | 0 | 3 | 0 | 0 | 2 ![]() |
[42] † Cochilius Last Modified in 07/19/17 | 46 | 97 | 97 | 0 | 3 | 0 | 0 | 2 ![]() |
[43] † Plagiarthrus Last Modified in 07/19/17 | 54 | 87 | 87 | 0 | 5 | 0 | 0 | 2 ![]() |
[44] † Protypotherium Last Modified in 07/19/17 | 22 | 120 | 119 | 0 | 5 | 0 | 0 | 2 ![]() |
[45] † Miocochilius Last Modified in 07/19/17 | 52 | 89 | 89 | 0 | 5 | 0 | 0 | 2 ![]() |
[46] † Interatherium Last Modified in 07/19/17 | 36 | 104 | 104 | 0 | 6 | 0 | 0 | 2 ![]() |
[47] † Federicoanaya Last Modified in 07/19/17 | 41 | 100 | 100 | 0 | 5 | 0 | 0 | 2 ![]() |
[48] † Trachytherus Last Modified in 07/19/17 | 16 | 128 | 128 | 0 | 2 | 0 | 0 | 2 ![]() |
[49] † Plesiotypotherium Last Modified in 07/19/17 | 29 | 104 | 104 | 0 | 13 | 0 | 0 | 2 ![]() |
[50] † Mesotherium Last Modified in 07/19/17 | 33 | 100 | 100 | 0 | 13 | 0 | 0 | 2 ![]() |
[51] † Eohyrax Last Modified in 07/19/17 | 92 | 52 | 52 | 0 | 2 | 0 | 0 | 2 ![]() |
[52] † Pseudhyrax Last Modified in 07/19/17 | 77 | 67 | 67 | 0 | 2 | 0 | 0 | 2 ![]() |
[53] † Archaeotypotherium Last Modified in 07/19/17 | 64 | 80 | 80 | 0 | 2 | 0 | 0 | 2 ![]() |
[54] † Archaeohyrax Last Modified in 07/19/17 | 14 | 131 | 130 | 0 | 2 | 0 | 0 | 2 ![]() |
[55] † Prohegetotherium Last Modified in 07/19/17 | 58 | 86 | 86 | 0 | 2 | 0 | 0 | 2 ![]() |
[56] † Hegetotherium Last Modified in 07/19/17 | 32 | 113 | 112 | 0 | 2 | 0 | 0 | 2 ![]() |
[57] † Prosotherium Last Modified in 07/19/17 | 56 | 80 | 80 | 0 | 10 | 0 | 0 | 2 ![]() |
[58] † Paedotherium Last Modified in 07/19/17 | 43 | 90 | 90 | 0 | 13 | 0 | 0 | 2 ![]() |
[59] † Zalambdalestes Last Modified in 07/19/17 | 2 | 103 | 103 | 0 | 41 | 0 | 0 | 2 ![]() |
[60] † Macrauchenia Last Modified in 07/19/17 | 13 | 101 | 101 | 0 | 32 | 0 | 0 | 2 ![]() |
[61] † Pampahippus secundus Last Modified in 07/19/17 | 71 | 76 | 74 | 0 | 1 | 0 | 0 | 2 ![]() |
[62] † Plexotemnus Last Modified in 07/19/17 | 101 | 45 | 44 | 0 | 1 | 0 | 0 | 2 ![]() |
[63] † Coquenia Last Modified in 07/19/17 | 10 | 135 | 135 | 0 | 1 | 0 | 0 | 2 ![]() |
[64] † Griphotherion Last Modified in 07/19/17 | 21 | 123 | 123 | 0 | 2 | 0 | 0 | 2 ![]() |
[65] † Boreastylops Last Modified in 07/19/17 | 22 | 120 | 120 | 0 | 4 | 0 | 0 | 2 ![]() |
[66] † Pampatemnus deuteros Last Modified in 07/19/17 | 39 | 107 | 107 | 0 | 0 | 0 | 0 | 2 ![]() |
[67] † Pampatemnus infernalis Last Modified in 07/19/17 | 76 | 69 | 69 | 0 | 1 | 0 | 0 | 2 ![]() |
[68] † Martinmiguelia fernandezi Last Modified in 07/19/17 | 101 | 45 | 45 | 0 | 0 | 0 | 0 | 2 ![]() |
Project views 
type | number of views | Individual items viewed (where applicable) |
Total project views | 25567 | |
Media views | 3373 | Media search (2888 views); M425185 (485 views); |
Project overview | 3255 | |
Documents list | 931 | |
Views for media list | 1063 | |
Specimen list | 1693 | |
Taxon list | 12618 | |
Bibliography | 820 | |
Matrix views | 1814 | Matrix landing page (1283 views); Relationships of Thomashuxleya within Notoungulata (149 views); Relationships of Thomashuxleya within Placentals (382 views); |
Project downloads 
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 431 | |
Document downloads | 8 | Matrix with constraint (1 download); Matrix with no constraints for Notoungulata (3 downloads); Matrix with constraint for Notoungulata and Litopterna (3 downloads); Notoungulata data matrix (1 download); |
Matrix downloads | 79 | Relationships of Thomashuxleya within Placentals (60 downloads); Relationships of Thomashuxleya within Notoungulata (19 downloads); |
Project downloads | 342 | |
Media downloads | 2 | M425185 (2 downloads); |