Project 2099: Jonah N. Choiniere, James M. Clark, Catherine A. Forster, Mark A. Norell, David A.Eberth, Gregory M. Erickson, Hongjun Chu, Xing Xu. 2014. A juvenile specimen of a new coelurosaur (Dinosauria: Theropoda) from the Middle–Late Jurassic Shishugou Formation of Xinjiang, People's Republic of China. Journal of Systematic Palaeontology. 12 (2):177-215.
Specimen: † Aorun zhaoi (IVPP/V:15709)

Abstract

We describe the anatomy of a new coelurosaurian theropod Aorun zhaoi gen. et sp. nov., from the Middle–Late Jurassic of Xinjiang, China. Histological analysis of the holotype and only known specimen shows that the new taxon is represented by the skeleton of a juvenile individual aged no more than one year. A phylogenetic analysis of theropod relationships places Aorun as a basal member of the Coelurosauria. Although the sole use of a sub-adult ontogenetic exemplar is potentially problematic for phylogenetic reconstruction, we show that the phylogenetic position of Aorun as a member of Coelurosauria is robust to the exclusion of characters known to change during theropod ontogeny. Aorun is the seventh theropod taxon, and temporally oldest coelurosaur, known from the Shishugou Formation, which has one of the most taxonomically diverse Jurassic coelurosaurian theropod faunas in the world.


Read the article »

Article DOI: 10.1080/14772019.2013.781067

Project DOI: 10.7934/P2099, http://dx.doi.org/10.7934/P2099
This project containsMatrices
  • 1 Media
  • 2 Matrices
  • 228 Taxa
  • 1 Specimen
  • 556 Characters
Total size of project's media files: 2 MB

Download Project SDD File
Total scored cells: 54045
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 1110
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 1276
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:998/112
Currently Viewing:
MorphoBank Project 2099

    This research
    supported by



    Members

    member name taxa specimens media chars character
    media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
    medialabels
    rules
    Jonah Choiniere
    Project Administrator
    Last logged in 01/16/15
    2281111100054045
    (51167, 0, 2878)
    000


    Taxonomic Overview for Matrix 'M23207' (114 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Herrerasaurus ischigualastensi
    Last Modified in 01/16/15
    184346345025001
    [2] Abelisaurus comahuensis
    Last Modified in 01/16/15
    478767502001
    [3] Acrocanthosaurus atokensis
    Last Modified in 01/16/15
    25229729407001
    [4] Aerosteon riocoloradensis
    Last Modified in 01/16/15
    458949403001
    [5] Afrovenator abakensis
    Last Modified in 01/16/15
    30524424307001
    [6] Albertonykus borealis
    Last Modified in 01/16/15
    522323101001
    [7] Albertosaurus sarcophagus
    Last Modified in 01/16/15
    15539439406001
    [8] Allosaurus fragilis
    Last Modified in 01/16/15
    41502493014001
    [9] Alvarezsaurus calvoi
    Last Modified in 01/16/15
    467848304001
    [10] Alxasaurus elesitaiensis
    Last Modified in 01/16/15
    37417717604001
    [11] Anserimimus planinychus
    Last Modified in 01/16/15
    45010010005001
    [12] Apsaravis ukhaana
    Last Modified in 01/16/15
    337193193025001
    [13] Archaeopteryx lithographica
    Last Modified in 01/16/15
    193341340021001
    [14] Avimimus portentosus
    Last Modified in 01/16/15
    248295295012001
    [15] Beipiaosaurus inexpectus
    Last Modified in 01/16/15
    43511211208001
    [16] Buitreraptor gonzalezorum
    Last Modified in 01/16/15
    37917017006001
    [17] Byronosaurus jaffeei
    Last Modified in 01/16/15
    385157156013001
    [18] Carcharodontosaurus saharicus
    Last Modified in 01/16/15
    42811911808001
    [19] Carnotaurus sastrei
    Last Modified in 01/16/15
    210324323021001
    [20] Caudipteryx zhoui
    Last Modified in 01/16/15
    298246241012001
    [21] Ceratosaurus nasicornis
    Last Modified in 01/16/15
    159382381014001
    [22] Chirostenotes pergracilis
    Last Modified in 01/16/15
    370172172013001
    [23] Citipati osmolskae
    Last Modified in 01/16/15
    178329329048001
    [24] Coelophysis bauri
    Last Modified in 01/16/15
    222316314018001
    [25] Coelurus fragilis
    Last Modified in 01/16/15
    41314114101001
    [26] Compsognathus longipes
    Last Modified in 01/16/15
    224316311015001
    [27] Conchoraptor gracilis
    Last Modified in 01/16/15
    363175175017001
    [28] Condorraptor currumili
    Last Modified in 01/16/15
    41713613602001
    [29] Confuciusornis sanctus
    Last Modified in 01/16/15
    267240240048001
    [30] Cryolophosaurus ellioti
    Last Modified in 01/16/15
    40115014706001
    [31] Daspletosaurus torosus
    Last Modified in 01/16/15
    35119719607001
    [32] Deinonychus antirrhopus
    Last Modified in 01/16/15
    127415415013001
    [33] Dilong paradoxus
    Last Modified in 01/16/15
    222318315015001
    [34] Dilophosaurus wetherilli
    Last Modified in 01/16/15
    199343340015001
    [35] Dromaeosaurus albertensis
    Last Modified in 01/16/15
    39315315309001
    [36] Dubreuillosaurus valesdunensis
    Last Modified in 01/16/15
    41413413209001
    [37] Duriavenator hesperis
    Last Modified in 01/16/15
    489636303001
    [38] EK troodontid
    Last Modified in 01/16/15
    505494901001
    [39] Eocarcharia dinops
    Last Modified in 01/16/15
    497575701001
    [40] Eoraptor lunensis
    Last Modified in 01/16/15
    311221221023001
    [41] Epidexipteryx hui
    Last Modified in 01/16/15
    37816916908001
    [42] Erlikosaurus andrewsi
    Last Modified in 01/16/15
    359174173022001
    [43] Eustreptospondylus oxiensis
    Last Modified in 01/16/15
    190350348016001
    [44] Falcarius utahensis
    Last Modified in 01/16/15
    18336836605001
    [45] † Aorun zhaoi
    Last Modified in 01/16/15
    287257254011001
    [46] Gallimimus bullatus
    Last Modified in 01/16/15
    189328328038001
    [47] Garudimimus brevipes
    Last Modified in 01/16/15
    191318318046001
    [48] Giganotosaurus carolinii
    Last Modified in 01/16/15
    36019319104001
    [49] Gigantoraptor erlianensis
    Last Modified in 01/16/15
    4588383014001
    [50] Gorgosaurus libratus
    Last Modified in 01/16/15
    30125025004001
    [51] Guanlong wucaii
    Last Modified in 01/16/15
    63480469013001
    [52] Hagryphus giganteus
    Last Modified in 01/16/15
    519363600001
    [53] Haplocheirus sollers
    Last Modified in 01/16/15
    106435428015001
    [54] Harpymimus okladnikovi
    Last Modified in 01/16/15
    218301300036001
    [55] Huaxiagnathus orientalis
    Last Modified in 01/16/15
    305238238012001
    [56] Incisivosaurus gauthieri
    Last Modified in 01/16/15
    337205205013001
    [57] Ingenia yanshinii
    Last Modified in 01/16/15
    41014214203001
    [58] Irritator challengerii
    Last Modified in 01/16/15
    471777707001
    [59] Juravenator starki
    Last Modified in 01/16/15
    37417317008001
    [60] Kileskus aristotocus
    Last Modified in 01/16/15
    484707001001
    [61] Limusaurus inextricabilis
    Last Modified in 01/16/15
    206296293055001
    [62] Linheraptor exquisitus
    Last Modified in 01/16/15
    315230230010001
    [63] Mahakala omnogovae
    Last Modified in 01/16/15
    470807905001
    [64] Majungasaurus crenatissimus
    Last Modified in 01/16/15
    155388387012001
    [65] Mapusaurus roseae
    Last Modified in 01/16/15
    36718618602001
    [66] Masiakasaurus knopfleri
    Last Modified in 01/16/15
    376169164010001
    [67] Megalosaurus bucklandii
    Last Modified in 01/16/15
    36318518408001
    [68] Mei long
    Last Modified in 01/16/15
    236295295024001
    [69] Microraptor gui
    Last Modified in 01/16/15
    323218218014001
    [70] Microvenator celer
    Last Modified in 01/16/15
    37117817507001
    [71] Monolophosaurus jiangi
    Last Modified in 01/16/15
    254289288012001
    [72] Mononykus olecranus
    Last Modified in 01/16/15
    312223223020001
    [73] Neovenator salerii
    Last Modified in 01/16/15
    25230029803001
    [74] Nomingia gobiensis
    Last Modified in 01/16/15
    38616116108001
    [75] Nothronychus graffami
    Last Modified in 01/16/15
    37018218203001
    [76] Ornitholestes hermanni
    Last Modified in 01/16/15
    218324320014001
    [77] Ornithomimus edmontonicus
    Last Modified in 01/16/15
    259275275021001
    [78] Oviraptor philoceratops
    Last Modified in 01/16/15
    421119119015001
    [79] Parvicursor remotus
    Last Modified in 01/16/15
    432107104018001
    [80] Patagonykus puertai
    Last Modified in 01/16/15
    39815515003001
    [81] Pelecanimimus polyodon
    Last Modified in 01/16/15
    305236228014001
    [82] Piatnitzkysaurus floresi
    Last Modified in 01/16/15
    38416716604001
    [83] Proceratosaurus bradleyi
    Last Modified in 01/16/15
    40214614407001
    [84] Rahonavis ostromi
    Last Modified in 01/16/15
    34320720705001
    [85] Raptorex kriegsteini
    Last Modified in 01/16/15
    30025025005001
    [86] Rinchenia mongoliensis
    Last Modified in 01/16/15
    440101101014001
    [87] Sapeornis chaoyangensis
    Last Modified in 01/16/15
    250275275030001
    [88] Saurornithoides mongoliensis
    Last Modified in 01/16/15
    39515315307001
    [89] Saurornitholestes langstoni
    Last Modified in 01/16/15
    35419819303001
    [90] Scipionyx samniticus
    Last Modified in 01/16/15
    332212212011001
    [91] Segnosaurus galboensis
    Last Modified in 01/16/15
    41513713703001
    [92] Shenzhousaurus orientalis
    Last Modified in 01/16/15
    301225225029001
    [93] Shuvuuia deserti
    Last Modified in 01/16/15
    77433430045001
    [94] Sinornithoides youngi
    Last Modified in 01/16/15
    39415415407001
    [95] Sinornithosaurus millenii
    Last Modified in 01/16/15
    315228228012001
    [96] Sinosauropteryx prima
    Last Modified in 01/16/15
    254288288013001
    [97] Sinovenator changii
    Last Modified in 01/16/15
    30624524504001
    [98] Sinraptor dongi
    Last Modified in 01/16/15
    103439435013001
    [99] Stokesosaurus langhami
    Last Modified in 01/16/15
    43212211801001
    [100] Struthiomimus altus
    Last Modified in 01/16/15
    181330330044001
    [101] Suchomimus tenerensis
    Last Modified in 01/16/15
    36218718508001
    [102] Syntarsus rhodesiensis
    Last Modified in 01/16/15
    173366363017001
    [103] Tanycolagreus topwilsoni
    Last Modified in 01/16/15
    32123223002001
    [104] Tarbosaurus bataar
    Last Modified in 01/16/15
    35519319307001
    [105] Tawa hallae
    Last Modified in 01/16/15
    273262262020001
    [106] Torvosaurus tanneri
    Last Modified in 01/16/15
    33121521409001
    [107] Troodon formosus
    Last Modified in 01/16/15
    262279276014001
    [108] Tsaagan mangas
    Last Modified in 01/16/15
    342200200013001
    [109] Tugulusaurus faciles
    Last Modified in 01/16/15
    497585800001
    [110] Tyrannosaurus rex
    Last Modified in 01/16/15
    29509507017001
    [111] Unenlagia plus neuquenraptor
    Last Modified in 01/16/15
    43611611203001
    [112] Velociraptor mongoliensis
    Last Modified in 01/16/15
    116421421018001
    [113] Zanabazar junior
    Last Modified in 01/16/15
    366178178011001
    [114] Zuolong salleei
    Last Modified in 01/16/15
    31323523207001


    Taxonomic Overview for Matrix 'M23208' (114 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] "Herrerasaurus ischigualastensis"
    Last Modified in 01/16/15
    184346345025001
    [2] "Abelisaurus comahuensis"
    Last Modified in 01/16/15
    478767502001
    [3] "Acrocanthosaurus atokensis"
    Last Modified in 01/16/15
    25229729407001
    [4] "Aerosteon riocoloradensis"
    Last Modified in 01/16/15
    458949403001
    [5] "Afrovenator abakensis"
    Last Modified in 01/16/15
    30524424307001
    [6] "Albertonykus borealis"
    Last Modified in 01/16/15
    522323101001
    [7] "Albertosaurus sarcophagus"
    Last Modified in 01/16/15
    15539439406001
    [8] "Allosaurus fragilis"
    Last Modified in 01/16/15
    41502493014001
    [9] "Alvarezsaurus calvoi"
    Last Modified in 01/16/15
    467848304001
    [10] "Alxasaurus elesitaiensis"
    Last Modified in 01/16/15
    37417717604001
    [11] "Anserimimus planinychus"
    Last Modified in 01/16/15
    45010010005001
    [12] "Apsaravis ukhaana"
    Last Modified in 01/16/15
    337193193025001
    [13] "Archaeopteryx lithographica"
    Last Modified in 01/16/15
    193341340021001
    [14] "Avimimus portentosus"
    Last Modified in 01/16/15
    248295295012001
    [15] "Beipiaosaurus inexpectus"
    Last Modified in 01/16/15
    43511211208001
    [16] "Buitreraptor gonzalezorum"
    Last Modified in 01/16/15
    37917017006001
    [17] "Byronosaurus jaffeei"
    Last Modified in 01/16/15
    385157156013001
    [18] "Carcharodontosaurus saharicus"
    Last Modified in 01/16/15
    42811911808001
    [19] "Carnotaurus sastrei"
    Last Modified in 01/16/15
    210324323021001
    [20] "Caudipteryx zhoui"
    Last Modified in 01/16/15
    298246241012001
    [21] "Ceratosaurus nasicornis"
    Last Modified in 01/16/15
    159382381014001
    [22] "Chirostenotes pergracilis"
    Last Modified in 01/16/15
    370172172013001
    [23] "Citipati osmolskae"
    Last Modified in 01/16/15
    178329329048001
    [24] "Coelophysis bauri"
    Last Modified in 01/16/15
    222316314018001
    [25] "Coelurus fragilis"
    Last Modified in 01/16/15
    41314114101001
    [26] "Compsognathus longipes"
    Last Modified in 01/16/15
    224316311015001
    [27] "Conchoraptor gracilis"
    Last Modified in 01/16/15
    363175175017001
    [28] "Condorraptor currumili"
    Last Modified in 01/16/15
    41713613602001
    [29] "Confuciusornis sanctus"
    Last Modified in 01/16/15
    267240240048001
    [30] "Cryolophosaurus ellioti"
    Last Modified in 01/16/15
    40115014706001
    [31] "Daspletosaurus torosus"
    Last Modified in 01/16/15
    35119719607001
    [32] "Deinonychus antirrhopus"
    Last Modified in 01/16/15
    127415415013001
    [33] "Dilong paradoxus"
    Last Modified in 01/16/15
    222318315015001
    [34] "Dilophosaurus wetherilli"
    Last Modified in 01/16/15
    199343340015001
    [35] "Dromaeosaurus albertensis"
    Last Modified in 01/16/15
    39315315309001
    [36] "Dubreuillosaurus valesdunensis"
    Last Modified in 01/16/15
    41413413209001
    [37] "Duriavenator hesperis"
    Last Modified in 01/16/15
    489636303001
    [38] "EK troodontid"
    Last Modified in 01/16/15
    505494901001
    [39] "Eocarcharia dinops"
    Last Modified in 01/16/15
    497575701001
    [40] "Eoraptor lunensis"
    Last Modified in 01/16/15
    311221221023001
    [41] "Epidexipteryx hui"
    Last Modified in 01/16/15
    37816916908001
    [42] "Erlikosaurus andrewsi"
    Last Modified in 01/16/15
    359174173022001
    [43] "Eustreptospondylus oxiensis"
    Last Modified in 01/16/15
    190350348016001
    [44] "Falcarius utahensis"
    Last Modified in 01/16/15
    18336836605001
    [45] "Aorun zhaoi"
    Last Modified in 01/16/15
    309235232011001
    [46] "Gallimimus bullatus"
    Last Modified in 01/16/15
    189328328038001
    [47] "Garudimimus brevipes"
    Last Modified in 01/16/15
    192317317046001
    [48] "Giganotosaurus carolinii"
    Last Modified in 01/16/15
    36019319104001
    [49] "Gigantoraptor erlianensis"
    Last Modified in 01/16/15
    4588383014001
    [50] "Gorgosaurus libratus"
    Last Modified in 01/16/15
    30125025004001
    [51] "Guanlong wucaii"
    Last Modified in 01/16/15
    63480469013001
    [52] "Hagryphus giganteus"
    Last Modified in 01/16/15
    519363600001
    [53] "Haplocheirus sollers"
    Last Modified in 01/16/15
    106435428015001
    [54] "Harpymimus okladnikovi"
    Last Modified in 01/16/15
    218301300036001
    [55] "Huaxiagnathus orientalis"
    Last Modified in 01/16/15
    305238238012001
    [56] "Incisivosaurus gauthieri"
    Last Modified in 01/16/15
    337205205013001
    [57] "Ingenia yanshinii"
    Last Modified in 01/16/15
    41014214203001
    [58] "Irritator challengerii"
    Last Modified in 01/16/15
    471777707001
    [59] "Juravenator starki"
    Last Modified in 01/16/15
    39115615308001
    [60] "Kileskus aristotocus"
    Last Modified in 01/16/15
    485696901001
    [61] "Limusaurus inextricabilis"
    Last Modified in 01/16/15
    206296293055001
    [62] "Linheraptor exquisitus"
    Last Modified in 01/16/15
    315230230010001
    [63] "Mahakala omnogovae"
    Last Modified in 01/16/15
    470807905001
    [64] "Majungasaurus crenatissimus"
    Last Modified in 01/16/15
    155388387012001
    [65] "Mapusaurus roseae"
    Last Modified in 01/16/15
    36718618602001
    [66] "Masiakasaurus knopfleri"
    Last Modified in 01/16/15
    376169164010001
    [67] "Megalosaurus bucklandii"
    Last Modified in 01/16/15
    36318518408001
    [68] "Mei long"
    Last Modified in 01/16/15
    236295295024001
    [69] "Microraptor gui"
    Last Modified in 01/16/15
    323218218014001
    [70] "Microvenator celer"
    Last Modified in 01/16/15
    37117817507001
    [71] "Monolophosaurus jiangi"
    Last Modified in 01/16/15
    254289288012001
    [72] "Mononykus olecranus"
    Last Modified in 01/16/15
    312223223020001
    [73] "Neovenator salerii"
    Last Modified in 01/16/15
    25230029803001
    [74] "Nomingia gobiensis"
    Last Modified in 01/16/15
    38616116108001
    [75] "Nothronychus graffami"
    Last Modified in 01/16/15
    37018218203001
    [76] "Ornitholestes hermanni"
    Last Modified in 01/16/15
    218324320014001
    [77] "Ornithomimus edmontonicus"
    Last Modified in 01/16/15
    259275275021001
    [78] "Oviraptor philoceratops"
    Last Modified in 01/16/15
    421119119015001
    [79] "Parvicursor remotus"
    Last Modified in 01/16/15
    432107104018001
    [80] "Patagonykus puertai"
    Last Modified in 01/16/15
    39815515003001
    [81] "Pelecanimimus polyodon"
    Last Modified in 01/16/15
    305236228014001
    [82] "Piatnitzkysaurus floresi"
    Last Modified in 01/16/15
    38416716604001
    [83] "Proceratosaurus bradleyi"
    Last Modified in 01/16/15
    40214614407001
    [84] "Rahonavis ostromi"
    Last Modified in 01/16/15
    34320720705001
    [85] "Raptorex kriegsteini"
    Last Modified in 01/16/15
    30025025005001
    [86] "Rinchenia mongoliensis"
    Last Modified in 01/16/15
    440101101014001
    [87] "Sapeornis chaoyangensis"
    Last Modified in 01/16/15
    250275275030001
    [88] "Saurornithoides mongoliensis"
    Last Modified in 01/16/15
    39515315307001
    [89] "Saurornitholestes langstoni"
    Last Modified in 01/16/15
    35419819303001
    [90] "Scipionyx samniticus"
    Last Modified in 01/16/15
    352192192011001
    [91] "Segnosaurus galboensis"
    Last Modified in 01/16/15
    41513713703001
    [92] "Shenzhousaurus orientalis"
    Last Modified in 01/16/15
    301225225029001
    [93] "Shuvuuia deserti"
    Last Modified in 01/16/15
    77433430045001
    [94] "Sinornithoides youngi"
    Last Modified in 01/16/15
    39415415407001
    [95] "Sinornithosaurus millenii"
    Last Modified in 01/16/15
    315228228012001
    [96] "Sinosauropteryx prima"
    Last Modified in 01/16/15
    254288288013001
    [97] "Sinovenator changii"
    Last Modified in 01/16/15
    30624524504001
    [98] "Sinraptor dongi"
    Last Modified in 01/16/15
    103439435013001
    [99] "Stokesosaurus langhami"
    Last Modified in 01/16/15
    43212211801001
    [100] "Struthiomimus altus"
    Last Modified in 01/16/15
    181330330044001
    [101] "Suchomimus tenerensis"
    Last Modified in 01/16/15
    36218718508001
    [102] "Syntarsus rhodesiensis"
    Last Modified in 01/16/15
    173366363017001
    [103] "Tanycolagreus topwilsoni"
    Last Modified in 01/16/15
    32123223002001
    [104] "Tarbosaurus bataar"
    Last Modified in 01/16/15
    35519319307001
    [105] "Tawa hallae"
    Last Modified in 01/16/15
    273262262020001
    [106] "Torvosaurus tanneri"
    Last Modified in 01/16/15
    33121521409001
    [107] "Troodon formosus"
    Last Modified in 01/16/15
    262279276014001
    [108] "Tsaagan mangas"
    Last Modified in 01/16/15
    342200200013001
    [109] "Tugulusaurus faciles"
    Last Modified in 01/16/15
    497585800001
    [110] "Tyrannosaurus rex"
    Last Modified in 01/16/15
    29509507017001
    [111] "Unenlagia plus neuquenraptor"
    Last Modified in 01/16/15
    43611611203001
    [112] "Velociraptor mongoliensis"
    Last Modified in 01/16/15
    116421421018001
    [113] "Zanabazar junior"
    Last Modified in 01/16/15
    366178178011001
    [114] "Zuolong salleei"
    Last Modified in 01/16/15
    31323523207001


    Project views

    type number of views Individual items viewed (where applicable)
    Total project views20607
    Project overview2792
    Matrix views2251Matrix landing page (1843 views); Choiniere et al 2014 Full matrix (221 views); Choiniere et al 2014 Ontogenetic matrix (187 views);
    Media views4022Media search (3511 views); M365941 (511 views);
    Taxon list10158
    Specimen list1384




    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project376
    Matrix downloads16Choiniere et al 2014 Full matrix (15 downloads); Choiniere et al 2014 Ontogenetic matrix (1 download);
    Project downloads360