Morphobank

Project 2191: J. C. Lamsdell, D. E. Briggs, H. P. Liu, B. J. Witzke, R. M. McKay. 2015. A new Ordovician arthropod from the Winneshiek Lagerstätte of Iowa (USA) reveals the ground plan of eurypterids and chasmataspidids. The Science of Nature. 102 (9-10):null.

Specimen: Winneshiekia youngae (SIU:140288)

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Abstract

Euchelicerates were a major component of Palaeozoic faunas but their basal relationships are uncertain: it has been suggested that Xiphosura – xiphosurids (horseshoe crabs) and similar Palaeozoic forms, the synziphosurines – may not represent a natural group. Basal euchelicerates are rare in the fossil record, however, particularly during the initial Ordovician radiation of the group. Here, we describe Winneshiekia youngae gen. et sp. nov., a euchelicerate from the Middle Ordovician (Darriwilian) Winneshiek Lagerstätte of Iowa. Winneshiekia shares features with both xiphosurans (a large, semicircular carapace and ophthalmic ridges) and dekatriatan euchelicerates such as chasmataspidids and eurypterids (an opisthosoma of 13 tergites). Phylogenetic analysis resolves Winneshiekia at the base of Dekatriata, sister taxon to a clade comprising chasmataspidids, eurypterids, arachnids, and Houia. Winneshiekia provides further support for the polyphyly of synziphosurines, traditionally considered the stem-lineage to xiphosurid horseshoe crabs, and by extension the paraphyly of Xiphosura. The new taxon reveals the ground pattern of Dekatriata and provides evidence of character polarity in chasmataspidids and eurypterids. The Winneshiek Lagerstätte represents an important window into early chelicerate evolution.

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Article DOI: 10.1007/s00114-015-1312-5

Project DOI: 10.7934/P2191, http://dx.doi.org/10.7934/P2191

This project contains	Matrices
1 Media 1 Matrix 105 Taxa 1 Specimen 216 Characters Total size of project's media files: 4.14M Download Project SDD File	Total scored cells: 17335 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 216 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 531 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:216/0

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MorphoBank Project 2191

Creation Date:
12 June 2015
Publication Date:
03 December 2015
Media downloads: 1
Matrix downloads: 8

This research
supported by

Authors' Institutions

University of Iowa

Yale University

Members

James Lamsdell
Project Administrator

105

216

17335
(14409, 0, 2926)

Taxonomic Overview for Matrix 'M23373' (105 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] Fuxianhuia protensa Taxon name last Modified on 06/12/15	32	135	135	0	49	0	0	1
[2] Leanchoilia illecebrosa Taxon name last Modified on 06/12/15	32	149	149	0	35	0	0	1
[3] Alalcomenaeus cambricus Taxon name last Modified on 06/12/15	25	156	156	0	35	0	0	1
[4] Yohoia tenuis Taxon name last Modified on 06/12/15	67	120	120	0	29	0	0	1
[5] Emeraldella brocki Taxon name last Modified on 06/12/15	18	152	152	0	46	0	0	1
[6] Sydneyia inexpectans Taxon name last Modified on 06/12/15	39	134	134	0	43	0	0	1
[7] Olenoides serratus Taxon name last Modified on 06/12/15	10	166	166	0	40	0	0	1
[8] Pycnogonum litorale Taxon name last Modified on 06/12/15	8	97	97	0	111	0	0	1
[9] Haliestes dasos Taxon name last Modified on 06/12/15	37	82	82	0	97	0	0	1
[10] Palaeoisopus problematicus Taxon name last Modified on 06/12/15	20	110	110	0	86	0	0	1
[11] Offacolus kingi Taxon name last Modified on 06/12/15	77	115	115	0	24	0	0	1
[12] Dibasterium durgae Taxon name last Modified on 06/12/15	53	132	132	0	31	0	0	1
[13] Anderella parva Taxon name last Modified on 06/12/15	149	63	63	0	4	0	0	1
[14] Weinbergina opitzi Taxon name last Modified on 06/12/15	62	136	136	0	18	0	0	1
[15] Camanchia grovensis Taxon name last Modified on 06/12/15	140	72	72	0	4	0	0	1
[16] Venustulus waukeshaensis Taxon name last Modified on 06/12/15	119	90	90	0	7	0	0	1
[17] Willwerathia laticeps Taxon name last Modified on 06/12/15	140	74	74	0	2	0	0	1
[18] Legrandella lombardi Taxon name last Modified on 06/12/15	134	80	80	0	2	0	0	1
[19] Bembicosoma pomphicus Taxon name last Modified on 06/12/15	150	64	64	0	2	0	0	1
[20] Bunodes lunula Taxon name last Modified on 06/12/15	147	65	65	0	4	0	0	1
[21] Limuloides limuloides Taxon name last Modified on 06/12/15	143	71	71	0	2	0	0	1
[22] Pasternakevia podolica Taxon name last Modified on 06/12/15	148	66	66	0	2	0	0	1
[23] Cyamocephalus loganensis Taxon name last Modified on 06/12/15	144	70	70	0	2	0	0	1
[24] Pseudoniscus roosevelti Taxon name last Modified on 06/12/15	149	65	65	0	2	0	0	1
[25] Euproops anthrax Taxon name last Modified on 06/12/15	94	101	101	0	21	0	0	1
[26] Kasibelinurus amoricum Taxon name last Modified on 06/12/15	145	67	67	0	4	0	0	1
[27] Winneshiekia youngae Taxon name last Modified on 06/12/15	147	67	67	0	2	0	0	1
[28] Houia yueya Taxon name last Modified on 06/12/15	138	71	71	0	7	0	0	1
[29] Lunataspis aurora Taxon name last Modified on 06/12/15	140	66	66	0	10	0	0	1
[30] Limulus polyphemus Taxon name last Modified on 06/12/15	0	179	179	0	37	0	0	1
[31] Paleolimulus signatus Taxon name last Modified on 06/12/15	42	146	146	0	28	0	0	1
[32] Stoermeropterus conicus Taxon name last Modified on 06/12/15	25	179	179	0	12	0	0	1
[33] Moselopterus ancylotelson Taxon name last Modified on 06/12/15	24	176	176	0	16	0	0	1
[34] Onychopterella kokomoensis Taxon name last Modified on 06/12/15	30	174	174	0	12	0	0	1
[35] Eurypterus tetragonophthalmus Taxon name last Modified on 06/12/15	8	194	194	0	14	0	0	1
[36] Erieopterus macrophthalmus Taxon name last Modified on 06/12/15	30	172	172	0	14	0	0	1
[37] Strobilopterus proteus Taxon name last Modified on 06/12/15	43	165	165	0	8	0	0	1
[38] Dolichopterus macrocheirus Taxon name last Modified on 06/12/15	22	183	183	0	11	0	0	1
[39] Carcinosoma newlini Taxon name last Modified on 06/12/15	27	174	174	0	15	0	0	1
[40] Megalograptus ohioensis Taxon name last Modified on 06/12/15	22	180	180	0	14	0	0	1
[41] Mixopterus kiaeri Taxon name last Modified on 06/12/15	28	173	173	0	15	0	0	1
[42] Orcanopterus manitoulinensis Taxon name last Modified on 06/12/15	69	131	131	0	16	0	0	1
[43] Adelophthalmus sievertsi Taxon name last Modified on 06/12/15	46	152	152	0	18	0	0	1
[44] Nanahughmilleria norvegica Taxon name last Modified on 06/12/15	23	181	181	0	12	0	0	1
[45] Hughmilleria socialis Taxon name last Modified on 06/12/15	23	177	177	0	16	0	0	1
[46] Slimonia acuminata Taxon name last Modified on 06/12/15	24	176	176	0	16	0	0	1
[47] Erettopterus bilobus Taxon name last Modified on 06/12/15	24	173	173	0	19	0	0	1
[48] Pterygotus anglicus Taxon name last Modified on 06/12/15	31	166	166	0	19	0	0	1
[49] Brachyopterus stubblefieldi Taxon name last Modified on 06/12/15	66	136	136	0	14	0	0	1
[50] Rhenopterus diensti Taxon name last Modified on 06/12/15	22	181	181	0	13	0	0	1
[51] Parastylonurus ornatus Taxon name last Modified on 06/12/15	28	175	175	0	13	0	0	1
[52] Laurieipterus elegans Taxon name last Modified on 06/12/15	88	116	116	0	12	0	0	1
[53] Kokomopterus longicaudatus Taxon name last Modified on 06/12/15	40	163	163	0	13	0	0	1
[54] Hardieopterus macrophthalmus Taxon name last Modified on 06/12/15	71	135	135	0	10	0	0	1
[55] Drepanopterus pentlandicus Taxon name last Modified on 06/12/15	26	179	178	0	12	0	0	1
[56] Hibbertopterus scouleri Taxon name last Modified on 06/12/15	27	177	177	0	12	0	0	1
[57] Megarachne servinei Taxon name last Modified on 06/12/15	157	54	53	0	6	0	0	1
[58] Chasmataspis laurencii Taxon name last Modified on 06/12/15	105	102	102	0	9	0	0	1
[59] Loganamaraspis dunlopi Taxon name last Modified on 06/12/15	83	114	114	0	19	0	0	1
[60] Achanarraspis reedi Taxon name last Modified on 06/12/15	145	67	67	0	4	0	0	1
[61] Heteroaspis stoermeri Taxon name last Modified on 06/12/15	140	68	68	0	8	0	0	1
[62] Diploaspis casteri Taxon name last Modified on 06/12/15	99	106	106	0	11	0	0	1
[63] Diploaspis muelleri Taxon name last Modified on 06/12/15	141	67	67	0	8	0	0	1
[64] Dvulikiaspis menneri Taxon name last Modified on 06/12/15	108	91	91	0	17	0	0	1
[65] Octoberaspis ushakovi Taxon name last Modified on 06/12/15	61	136	136	0	19	0	0	1
[66] Centruroides vittatus Taxon name last Modified on 06/12/15	0	186	186	0	30	0	0	1
[67] Androctonus australis Taxon name last Modified on 06/12/15	0	186	186	0	30	0	0	1
[68] Proscorpius osborni Taxon name last Modified on 06/12/15	26	159	159	0	31	0	0	1
[69] Palaeophonus nuncius Taxon name last Modified on 06/12/15	52	136	136	0	28	0	0	1
[70] Caddo agilis Taxon name last Modified on 06/12/15	7	166	166	0	43	0	0	1
[71] Chileogovea oedipus Taxon name last Modified on 06/12/15	14	162	162	0	40	0	0	1
[72] Cyphophthalmus duricorius Taxon name last Modified on 06/12/15	14	162	162	0	40	0	0	1
[73] Gonyleptes horridus Taxon name last Modified on 06/12/15	7	167	167	0	42	0	0	1
[74] Leiobunum rotundum Taxon name last Modified on 06/12/15	7	166	166	0	43	0	0	1
[75] Sclerobunus robustus Taxon name last Modified on 06/12/15	7	167	167	0	42	0	0	1
[76] Allothrombium simoni Taxon name last Modified on 06/12/15	14	151	150	0	52	0	0	1
[77] Amblyomma americanum Taxon name last Modified on 06/12/15	12	149	148	0	56	0	0	1
[78] Neocarus texanus Taxon name last Modified on 06/12/15	8	156	156	0	52	0	0	1
[79] Cryptocellus goodnighti Taxon name last Modified on 06/12/15	4	169	169	0	43	0	0	1
[80] Poliochera punctulata Taxon name last Modified on 06/12/15	59	115	115	0	42	0	0	1
[81] Ricinoides atewa Taxon name last Modified on 06/12/15	4	169	169	0	43	0	0	1
[82] Anthracomartus hindi Taxon name last Modified on 06/12/15	28	149	149	0	39	0	0	1
[83] Eophrynus prestvicii Taxon name last Modified on 06/12/15	28	152	152	0	36	0	0	1
[84] Gilboarachne griersoni Taxon name last Modified on 06/12/15	17	159	159	0	40	0	0	1
[85] Palaeocharinus sp. Taxon name last Modified on 06/12/15	5	171	171	0	40	0	0	1
[86] Mesotarbus peteri Taxon name last Modified on 06/12/15	46	128	128	0	42	0	0	1
[87] Plesiosiro madeleyi Taxon name last Modified on 06/12/15	40	136	136	0	40	0	0	1
[88] Eukoenenia mirabilis Taxon name last Modified on 06/12/15	3	170	169	0	44	0	0	1
[89] Prokoenenia wheeleri Taxon name last Modified on 06/12/15	3	170	169	0	44	0	0	1
[90] Mastigoproctus giganteus Taxon name last Modified on 06/12/15	3	176	176	0	37	0	0	1
[91] Thelyphonus caudatus Taxon name last Modified on 06/12/15	3	175	175	0	38	0	0	1
[92] Hubbardia pentapeltis Taxon name last Modified on 06/12/15	3	175	174	0	39	0	0	1
[93] Schizomus crassicaudatus Taxon name last Modified on 06/12/15	3	175	174	0	39	0	0	1
[94] Permarachne novokshonovi Taxon name last Modified on 06/12/15	87	100	100	0	29	0	0	1
[95] Attercopus filmbriunguis Taxon name last Modified on 06/12/15	138	66	66	0	12	0	0	1
[96] Liphistius malayanus Taxon name last Modified on 06/12/15	3	164	164	0	49	0	0	1
[97] Aphonopelma hentzi Taxon name last Modified on 06/12/15	3	157	157	0	56	0	0	1
[98] Hypochilus pococki Taxon name last Modified on 06/12/15	4	157	157	0	55	0	0	1
[99] Cupienius getazi Taxon name last Modified on 06/12/15	4	156	156	0	56	0	0	1
[100] Neobisium maritimum Taxon name last Modified on 06/12/15	10	155	154	0	52	0	0	1
[101] Chelifer cancroides Taxon name last Modified on 06/12/15	10	154	153	0	53	0	0	1
[102] Eremocosta striata Taxon name last Modified on 06/12/15	10	151	150	0	56	0	0	1
[103] Galeodes armeniacus Taxon name last Modified on 06/12/15	10	151	150	0	56	0	0	1
[104] Phrynus marginemaculatus Taxon name last Modified on 06/12/15	3	171	171	0	42	0	0	1
[105] Charinus victori Taxon name last Modified on 06/12/15	3	171	171	0	42	0	0	1

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	256
Project downloads	247
Matrix downloads	8	Phylogenetic analysis of Winneshiekia (8 downloads);
Media downloads	1	M378310 (1 download);