Project 2191: J. C. Lamsdell, D. E. Briggs, H. P. Liu, B. J. Witzke, R. M. McKay. 2015. A new Ordovician arthropod from the Winneshiek Lagerstätte of Iowa (USA) reveals the ground plan of eurypterids and chasmataspidids. The Science of Nature. 102 (9-10):null.
Abstract
Euchelicerates were a major component of Palaeozoic faunas but their basal relationships are uncertain: it has been suggested that Xiphosura – xiphosurids (horseshoe crabs) and similar Palaeozoic forms, the synziphosurines – may not represent a natural group. Basal euchelicerates are rare in the fossil record, however, particularly during the initial Ordovician radiation of the group. Here, we describe Winneshiekia youngae gen. et sp. nov., a euchelicerate from the Middle Ordovician (Darriwilian) Winneshiek Lagerstätte of Iowa. Winneshiekia shares features with both xiphosurans (a large, semicircular carapace and ophthalmic ridges) and dekatriatan euchelicerates such as chasmataspidids and eurypterids (an opisthosoma of 13 tergites). Phylogenetic analysis resolves Winneshiekia at the base of Dekatriata, sister taxon to a clade comprising chasmataspidids, eurypterids, arachnids, and Houia. Winneshiekia provides further support for the polyphyly of synziphosurines, traditionally considered the stem-lineage to xiphosurid horseshoe crabs, and by extension the paraphyly of Xiphosura. The new taxon reveals the ground pattern of Dekatriata and provides evidence of character polarity in chasmataspidids and eurypterids. The Winneshiek Lagerstätte represents an important window into early chelicerate evolution.Read the article »
Article DOI: 10.1007/s00114-015-1312-5
Project DOI: 10.7934/P2191, http://dx.doi.org/10.7934/P2191
This project contains | Matrices |
---|---|
Download Project SDD File | Total scored cells: 17335 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 216 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 531 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:216/0 |
Currently Viewing:
MorphoBank Project 2191
MorphoBank Project 2191
- Creation Date:
12 June 2015 - Publication Date:
03 December 2015 - Project views: 31209
- Media downloads: 1
- Matrix downloads: 8
This research
supported by
Authors' Institutions
- University of Iowa
- Yale University
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
James Lamsdell Project Administrator | 105 | 1 | 1 | 216 | 0 | 0 | 17335 (14409, 0, 2926) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M23373' (105 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Fuxianhuia protensa Last Modified in 06/12/15 | 32 | 135 | 135 | 0 | 49 | 0 | 0 | 1 |
[2] Leanchoilia illecebrosa Last Modified in 06/12/15 | 32 | 149 | 149 | 0 | 35 | 0 | 0 | 1 |
[3] Alalcomenaeus cambricus Last Modified in 06/12/15 | 25 | 156 | 156 | 0 | 35 | 0 | 0 | 1 |
[4] Yohoia tenuis Last Modified in 06/12/15 | 67 | 120 | 120 | 0 | 29 | 0 | 0 | 1 |
[5] Emeraldella brocki Last Modified in 06/12/15 | 18 | 152 | 152 | 0 | 46 | 0 | 0 | 1 |
[6] Sydneyia inexpectans Last Modified in 06/12/15 | 39 | 134 | 134 | 0 | 43 | 0 | 0 | 1 |
[7] Olenoides serratus Last Modified in 06/12/15 | 10 | 166 | 166 | 0 | 40 | 0 | 0 | 1 |
[8] Pycnogonum litorale Last Modified in 06/12/15 | 8 | 97 | 97 | 0 | 111 | 0 | 0 | 1 |
[9] Haliestes dasos Last Modified in 06/12/15 | 37 | 82 | 82 | 0 | 97 | 0 | 0 | 1 |
[10] Palaeoisopus problematicus Last Modified in 06/12/15 | 20 | 110 | 110 | 0 | 86 | 0 | 0 | 1 |
[11] Offacolus kingi Last Modified in 06/12/15 | 77 | 115 | 115 | 0 | 24 | 0 | 0 | 1 |
[12] Dibasterium durgae Last Modified in 06/12/15 | 53 | 132 | 132 | 0 | 31 | 0 | 0 | 1 |
[13] Anderella parva Last Modified in 06/12/15 | 149 | 63 | 63 | 0 | 4 | 0 | 0 | 1 |
[14] Weinbergina opitzi Last Modified in 06/12/15 | 62 | 136 | 136 | 0 | 18 | 0 | 0 | 1 |
[15] Camanchia grovensis Last Modified in 06/12/15 | 140 | 72 | 72 | 0 | 4 | 0 | 0 | 1 |
[16] Venustulus waukeshaensis Last Modified in 06/12/15 | 119 | 90 | 90 | 0 | 7 | 0 | 0 | 1 |
[17] Willwerathia laticeps Last Modified in 06/12/15 | 140 | 74 | 74 | 0 | 2 | 0 | 0 | 1 |
[18] Legrandella lombardi Last Modified in 06/12/15 | 134 | 80 | 80 | 0 | 2 | 0 | 0 | 1 |
[19] Bembicosoma pomphicus Last Modified in 06/12/15 | 150 | 64 | 64 | 0 | 2 | 0 | 0 | 1 |
[20] Bunodes lunula Last Modified in 06/12/15 | 147 | 65 | 65 | 0 | 4 | 0 | 0 | 1 |
[21] Limuloides limuloides Last Modified in 06/12/15 | 143 | 71 | 71 | 0 | 2 | 0 | 0 | 1 |
[22] Pasternakevia podolica Last Modified in 06/12/15 | 148 | 66 | 66 | 0 | 2 | 0 | 0 | 1 |
[23] Cyamocephalus loganensis Last Modified in 06/12/15 | 144 | 70 | 70 | 0 | 2 | 0 | 0 | 1 |
[24] Pseudoniscus roosevelti Last Modified in 06/12/15 | 149 | 65 | 65 | 0 | 2 | 0 | 0 | 1 |
[25] Euproops anthrax Last Modified in 06/12/15 | 94 | 101 | 101 | 0 | 21 | 0 | 0 | 1 |
[26] Kasibelinurus amoricum Last Modified in 06/12/15 | 145 | 67 | 67 | 0 | 4 | 0 | 0 | 1 |
[27] Winneshiekia youngae Last Modified in 06/12/15 | 147 | 67 | 67 | 0 | 2 | 0 | 0 | 1 |
[28] Houia yueya Last Modified in 06/12/15 | 138 | 71 | 71 | 0 | 7 | 0 | 0 | 1 |
[29] Lunataspis aurora Last Modified in 06/12/15 | 140 | 66 | 66 | 0 | 10 | 0 | 0 | 1 |
[30] Limulus polyphemus Last Modified in 06/12/15 | 0 | 179 | 179 | 0 | 37 | 0 | 0 | 1 |
[31] Paleolimulus signatus Last Modified in 06/12/15 | 42 | 146 | 146 | 0 | 28 | 0 | 0 | 1 |
[32] Stoermeropterus conicus Last Modified in 06/12/15 | 25 | 179 | 179 | 0 | 12 | 0 | 0 | 1 |
[33] Moselopterus ancylotelson Last Modified in 06/12/15 | 24 | 176 | 176 | 0 | 16 | 0 | 0 | 1 |
[34] Onychopterella kokomoensis Last Modified in 06/12/15 | 30 | 174 | 174 | 0 | 12 | 0 | 0 | 1 |
[35] Eurypterus tetragonophthalmus Last Modified in 06/12/15 | 8 | 194 | 194 | 0 | 14 | 0 | 0 | 1 |
[36] Erieopterus macrophthalmus Last Modified in 06/12/15 | 30 | 172 | 172 | 0 | 14 | 0 | 0 | 1 |
[37] Strobilopterus proteus Last Modified in 06/12/15 | 43 | 165 | 165 | 0 | 8 | 0 | 0 | 1 |
[38] Dolichopterus macrocheirus Last Modified in 06/12/15 | 22 | 183 | 183 | 0 | 11 | 0 | 0 | 1 |
[39] Carcinosoma newlini Last Modified in 06/12/15 | 27 | 174 | 174 | 0 | 15 | 0 | 0 | 1 |
[40] Megalograptus ohioensis Last Modified in 06/12/15 | 22 | 180 | 180 | 0 | 14 | 0 | 0 | 1 |
[41] Mixopterus kiaeri Last Modified in 06/12/15 | 28 | 173 | 173 | 0 | 15 | 0 | 0 | 1 |
[42] Orcanopterus manitoulinensis Last Modified in 06/12/15 | 69 | 131 | 131 | 0 | 16 | 0 | 0 | 1 |
[43] Adelophthalmus sievertsi Last Modified in 06/12/15 | 46 | 152 | 152 | 0 | 18 | 0 | 0 | 1 |
[44] Nanahughmilleria norvegica Last Modified in 06/12/15 | 23 | 181 | 181 | 0 | 12 | 0 | 0 | 1 |
[45] Hughmilleria socialis Last Modified in 06/12/15 | 23 | 177 | 177 | 0 | 16 | 0 | 0 | 1 |
[46] Slimonia acuminata Last Modified in 06/12/15 | 24 | 176 | 176 | 0 | 16 | 0 | 0 | 1 |
[47] Erettopterus bilobus Last Modified in 06/12/15 | 24 | 173 | 173 | 0 | 19 | 0 | 0 | 1 |
[48] Pterygotus anglicus Last Modified in 06/12/15 | 31 | 166 | 166 | 0 | 19 | 0 | 0 | 1 |
[49] Brachyopterus stubblefieldi Last Modified in 06/12/15 | 66 | 136 | 136 | 0 | 14 | 0 | 0 | 1 |
[50] Rhenopterus diensti Last Modified in 06/12/15 | 22 | 181 | 181 | 0 | 13 | 0 | 0 | 1 |
[51] Parastylonurus ornatus Last Modified in 06/12/15 | 28 | 175 | 175 | 0 | 13 | 0 | 0 | 1 |
[52] Laurieipterus elegans Last Modified in 06/12/15 | 88 | 116 | 116 | 0 | 12 | 0 | 0 | 1 |
[53] Kokomopterus longicaudatus Last Modified in 06/12/15 | 40 | 163 | 163 | 0 | 13 | 0 | 0 | 1 |
[54] Hardieopterus macrophthalmus Last Modified in 06/12/15 | 71 | 135 | 135 | 0 | 10 | 0 | 0 | 1 |
[55] Drepanopterus pentlandicus Last Modified in 06/12/15 | 26 | 179 | 178 | 0 | 12 | 0 | 0 | 1 |
[56] Hibbertopterus scouleri Last Modified in 06/12/15 | 27 | 177 | 177 | 0 | 12 | 0 | 0 | 1 |
[57] Megarachne servinei Last Modified in 06/12/15 | 157 | 54 | 53 | 0 | 6 | 0 | 0 | 1 |
[58] Chasmataspis laurencii Last Modified in 06/12/15 | 105 | 102 | 102 | 0 | 9 | 0 | 0 | 1 |
[59] Loganamaraspis dunlopi Last Modified in 06/12/15 | 83 | 114 | 114 | 0 | 19 | 0 | 0 | 1 |
[60] Achanarraspis reedi Last Modified in 06/12/15 | 145 | 67 | 67 | 0 | 4 | 0 | 0 | 1 |
[61] Heteroaspis stoermeri Last Modified in 06/12/15 | 140 | 68 | 68 | 0 | 8 | 0 | 0 | 1 |
[62] Diploaspis casteri Last Modified in 06/12/15 | 99 | 106 | 106 | 0 | 11 | 0 | 0 | 1 |
[63] Diploaspis muelleri Last Modified in 06/12/15 | 141 | 67 | 67 | 0 | 8 | 0 | 0 | 1 |
[64] Dvulikiaspis menneri Last Modified in 06/12/15 | 108 | 91 | 91 | 0 | 17 | 0 | 0 | 1 |
[65] Octoberaspis ushakovi Last Modified in 06/12/15 | 61 | 136 | 136 | 0 | 19 | 0 | 0 | 1 |
[66] Centruroides vittatus Last Modified in 06/12/15 | 0 | 186 | 186 | 0 | 30 | 0 | 0 | 1 |
[67] Androctonus australis Last Modified in 06/12/15 | 0 | 186 | 186 | 0 | 30 | 0 | 0 | 1 |
[68] Proscorpius osborni Last Modified in 06/12/15 | 26 | 159 | 159 | 0 | 31 | 0 | 0 | 1 |
[69] Palaeophonus nuncius Last Modified in 06/12/15 | 52 | 136 | 136 | 0 | 28 | 0 | 0 | 1 |
[70] Caddo agilis Last Modified in 06/12/15 | 7 | 166 | 166 | 0 | 43 | 0 | 0 | 1 |
[71] Chileogovea oedipus Last Modified in 06/12/15 | 14 | 162 | 162 | 0 | 40 | 0 | 0 | 1 |
[72] Cyphophthalmus duricorius Last Modified in 06/12/15 | 14 | 162 | 162 | 0 | 40 | 0 | 0 | 1 |
[73] Gonyleptes horridus Last Modified in 06/12/15 | 7 | 167 | 167 | 0 | 42 | 0 | 0 | 1 |
[74] Leiobunum rotundum Last Modified in 06/12/15 | 7 | 166 | 166 | 0 | 43 | 0 | 0 | 1 |
[75] Sclerobunus robustus Last Modified in 06/12/15 | 7 | 167 | 167 | 0 | 42 | 0 | 0 | 1 |
[76] Allothrombium simoni Last Modified in 06/12/15 | 14 | 151 | 150 | 0 | 52 | 0 | 0 | 1 |
[77] Amblyomma americanum Last Modified in 06/12/15 | 12 | 149 | 148 | 0 | 56 | 0 | 0 | 1 |
[78] Neocarus texanus Last Modified in 06/12/15 | 8 | 156 | 156 | 0 | 52 | 0 | 0 | 1 |
[79] Cryptocellus goodnighti Last Modified in 06/12/15 | 4 | 169 | 169 | 0 | 43 | 0 | 0 | 1 |
[80] Poliochera punctulata Last Modified in 06/12/15 | 59 | 115 | 115 | 0 | 42 | 0 | 0 | 1 |
[81] Ricinoides atewa Last Modified in 06/12/15 | 4 | 169 | 169 | 0 | 43 | 0 | 0 | 1 |
[82] Anthracomartus hindi Last Modified in 06/12/15 | 28 | 149 | 149 | 0 | 39 | 0 | 0 | 1 |
[83] Eophrynus prestvicii Last Modified in 06/12/15 | 28 | 152 | 152 | 0 | 36 | 0 | 0 | 1 |
[84] Gilboarachne griersoni Last Modified in 06/12/15 | 17 | 159 | 159 | 0 | 40 | 0 | 0 | 1 |
[85] Palaeocharinus sp. Last Modified in 06/12/15 | 5 | 171 | 171 | 0 | 40 | 0 | 0 | 1 |
[86] Mesotarbus peteri Last Modified in 06/12/15 | 46 | 128 | 128 | 0 | 42 | 0 | 0 | 1 |
[87] Plesiosiro madeleyi Last Modified in 06/12/15 | 40 | 136 | 136 | 0 | 40 | 0 | 0 | 1 |
[88] Eukoenenia mirabilis Last Modified in 06/12/15 | 3 | 170 | 169 | 0 | 44 | 0 | 0 | 1 |
[89] Prokoenenia wheeleri Last Modified in 06/12/15 | 3 | 170 | 169 | 0 | 44 | 0 | 0 | 1 |
[90] Mastigoproctus giganteus Last Modified in 06/12/15 | 3 | 176 | 176 | 0 | 37 | 0 | 0 | 1 |
[91] Thelyphonus caudatus Last Modified in 06/12/15 | 3 | 175 | 175 | 0 | 38 | 0 | 0 | 1 |
[92] Hubbardia pentapeltis Last Modified in 06/12/15 | 3 | 175 | 174 | 0 | 39 | 0 | 0 | 1 |
[93] Schizomus crassicaudatus Last Modified in 06/12/15 | 3 | 175 | 174 | 0 | 39 | 0 | 0 | 1 |
[94] Permarachne novokshonovi Last Modified in 06/12/15 | 87 | 100 | 100 | 0 | 29 | 0 | 0 | 1 |
[95] Attercopus filmbriunguis Last Modified in 06/12/15 | 138 | 66 | 66 | 0 | 12 | 0 | 0 | 1 |
[96] Liphistius malayanus Last Modified in 06/12/15 | 3 | 164 | 164 | 0 | 49 | 0 | 0 | 1 |
[97] Aphonopelma hentzi Last Modified in 06/12/15 | 3 | 157 | 157 | 0 | 56 | 0 | 0 | 1 |
[98] Hypochilus pococki Last Modified in 06/12/15 | 4 | 157 | 157 | 0 | 55 | 0 | 0 | 1 |
[99] Cupienius getazi Last Modified in 06/12/15 | 4 | 156 | 156 | 0 | 56 | 0 | 0 | 1 |
[100] Neobisium maritimum Last Modified in 06/12/15 | 10 | 155 | 154 | 0 | 52 | 0 | 0 | 1 |
[101] Chelifer cancroides Last Modified in 06/12/15 | 10 | 154 | 153 | 0 | 53 | 0 | 0 | 1 |
[102] Eremocosta striata Last Modified in 06/12/15 | 10 | 151 | 150 | 0 | 56 | 0 | 0 | 1 |
[103] Galeodes armeniacus Last Modified in 06/12/15 | 10 | 151 | 150 | 0 | 56 | 0 | 0 | 1 |
[104] Phrynus marginemaculatus Last Modified in 06/12/15 | 3 | 171 | 171 | 0 | 42 | 0 | 0 | 1 |
[105] Charinus victori Last Modified in 06/12/15 | 3 | 171 | 171 | 0 | 42 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 31209 | |
Matrix views | 2323 | Matrix landing page (2007 views); Phylogenetic analysis of Winneshiekia (316 views); |
Project overview | 3039 | |
Media views | 4893 | Media search (4177 views); M378310 (716 views); |
Specimen list | 2310 | |
Bibliography | 1125 | |
Taxon list | 17499 | |
Documents list | 20 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 258 | |
Project downloads | 249 | |
Matrix downloads | 8 | Phylogenetic analysis of Winneshiekia (8 downloads); |
Media downloads | 1 | M378310 (1 download); |