Project 2234: S. Fujimoto, A. Jørgensen, J. G. Hansen. 2017. A molecular approach to arthrotardigrade phylogeny (Heterotardigrada, Tardigrada). Zoologica Scripta. 46 (4):496-505.
Specimen: Coronarctus sp. (unvouchered)
View: Micrographs taken at 400x magnification


The marine order Arthrotardigrada (class Heterotardigrada, phylum Tardigrada) is known for its conspicuously high morphological diversity and has been traditionally recognized as the most ancestral group within the phylum. Despite its potential importance in understanding the evolution of the phylum, the phylogenetic relationships of Arthrotardigrada have not been clarified. This study conducted molecular phylogenetic analyses of the order encompassing all families except Neoarctidae using nuclear 18S and 28S rRNA fragments. Data from two rare families, Coronarctidae and Renaudarctidae, were included for the first time. The analyses confirmed the monophyly of Heterotardigrada and inferred Coronarctidae as the sister group to all other heterotardigrade taxa. Furthermore, the results support a monophyletic Renaudarctidae + Stygarctidae clade, which has been previously suggested on morphology. Our data indicated that two subfamilies currently placed in Halechiniscidae are only distantly related to this family. We propose that these taxa are each elevated to family level (Styraconyxidae (new rank) and Tanarctidae (new rank)). The morphology of tardigrades is discussed in the context of the inferred phylogeny. (from Fujimoto, S., Jørgensen, A. & Hansen, J.G. (2017). A molecular approach to arthrotardigrade phylogeny (Heterotardigrada, Tardigrada). Zoologica Scripta, 46, 496–505.)

Read the article »

Article DOI: 10.1111/zsc.12221

Project DOI: 10.7934/P2234,
This project contains
  • 42 Media
  • 22 Taxa
  • 22 Specimens
Total size of project's media files: 344.83M

Download Project SDD File
Currently Viewing:
MorphoBank Project 2234
  • Creation Date:
    13 August 2015
  • Publication Date:
    06 February 2019
  • Project views: 20017

    Authors' Institutions

    • Copenhagen University

    • Kyoto University

    • Tohoku University

    • Natural History Museum of Denmark


    member name taxa specimens media
    Shinta Fujimoto
    Project Administrator
    Jesper G. Hansen
    Full membership
    Aslak Jørgensen
    Full membership

    Project has no matrices defined.

    Project views

    type number of views Individual items viewed (where applicable)
    Total project views20017
    Project overview987
    Media views8955Media search (1370 views); M390613 (176 views); M390614 (169 views); M390615 (181 views); M390616 (182 views); M390617 (178 views); M390618 (179 views); M390619 (180 views); M390620 (175 views); M390621 (193 views); M390622 (181 views); M390623 (173 views); M390624 (170 views); M390625 (177 views); M390626 (166 views); M390627 (171 views); M390628 (169 views); M390629 (170 views); M390630 (166 views); M390631 (223 views); M390632 (186 views); M390633 (180 views); M390634 (176 views); M390635 (166 views); M390636 (201 views); M390637 (174 views); M390638 (185 views); M390639 (205 views); M390640 (188 views); M390641 (180 views); M390642 (176 views); M390643 (161 views); M390644 (188 views); M390645 (177 views); M390646 (180 views); M390647 (171 views); M390648 (181 views); M390649 (234 views); M390650 (178 views); M390651 (172 views); M390652 (179 views); M390653 (172 views); M390654 (196 views);
    Views for media list437
    Specimen list6179
    Taxon list3148
    Documents list1

    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project144
    Project downloads144