Project 2286: S. TARASOV. 2017. A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data. Zootaxa. 4329 (2):101.
Abstract
Two Oriental dung beetle genera: Parachorius Harold, 1873 and Cassolus Sharp, 1875 have long had an ambiguous tribal position in Scarabaeinae (Coleoptera: Scarabaeidae), but have never been considered as closely related. A recently discovered species representing the morphological link between the two genera gave a hint to their possible close affiliation. To assess phylogenetic and taxonomic placement of these genera, I conducted phylogenetic analyses of global dung beetle samples using morphological (134 taxa, 232 characters) and molecular (551 terminals, 8 gene regions) data. Both morphological and molecular analyses strongly support the monophyly of Parachorius + Cassolus. This leads to the synonymy of Parachorius with Cassolus new synonymy, and resulted in the new generic concept for Parachorius. The isolated phylogenetic position of Parachorius and its morphological distinctiveness from all other known Scarabaeinae tribes suggest recognition of a new tribe, Parachoriini new tribe, to maintain the stability of tribal classification in dung beetles. Investigation of old and recent material of Parachorius revealed a large number of undescribed species and the need for a taxonomic revision of this genus. The revision of Parachorius, powered by the 3i cybertaxonomic tool, is presented in this study. The revised Parachorius is comprised of 19 species from the Oriental and southeastern Palaearctic Regions, of which seven are newly described (P. asymmetricus new species, P. bolavensis new species, P. longipenis new species, P. newthayerae new species, P. pseudojavanus new species, P. schuelkei new species, and P. solodovnikovi new species). Three species names in Parachorius are synonymized, namely, P. fungorum Kryzhanovsky & Medvedev, 1966 = P. krali Utsunomiya & Masumoto, 2001 new synonymy; P. thomsoni Harold, 1873 = P. lannathai Hanboonsong & Masumoto, 2001 new synonymy; and P. peninsularis (Arrow, 1907) = C. pongchaii Masumoto, 2001 new synonymy. Two species originally described in Cassolus (C. sumatranus and C. minutus) are transferred to the genus Panelus Lewis, 1895. The rank of the genus Macropanelus is lowered to a subgenus within Panelus (i.e. Panelus (Macropanelus) new status).Read the article »
Article DOI: 10.11646/zootaxa.4329.2.1
Project DOI: 10.7934/P2286, http://dx.doi.org/10.7934/P2286
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 30640 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 232 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 468 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:232/0 |
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MorphoBank Project 2286
MorphoBank Project 2286
- Creation Date:
01 October 2015 - Publication Date:
13 September 2017 - Project views: 24444
- Matrix downloads: 12
Authors' Institutions
- University of Oslo
- University of Tennessee
Members
member name | taxa | specimens | media | media notes | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||||||||||
Sergei Tarasov Project Administrator | 134 | 1 | 1 | 0 | 232 | 0 | 0 | 0 | 0 | 30640 (30052, 0, 588) | 0 | 7 | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M24757' (134 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Aphodius erraticus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[2] Podotenus fulviventris Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[3] Aulonocnemis crassecostata Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[4] Manjarivolo sp Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[5] Coptorhina auspicata Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[6] Delopleurus sp Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[7] Frankenbergerius armatus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[8] Sarophorus costatus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[9] Odontoloma sp Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[10] Nebulasilvius insularis Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[11] Endroedyolus paradoxus Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[12] Dicranocara deschodti Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[13] Byrrhidium namaquensis Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[14] Paraphytus sp Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[15] Epirinus sp Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[16] Neosisyphus sp Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[17] Macroderes mutilans Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[18] Xinidium dentilabris Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[19] Pedaria sp Last Modified in 08/14/17 | 1 | 231 | 231 | 0 | 0 | 0 | 0 | 1 |
[20] Gyronotus carinatus Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[21] Anachalcos convexus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[22] Hammondantus psammophilus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[23] Pycnopanelus krikkeni Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[24] Janssensantus pauliani Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[25] Aphengoecus multiserratus Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[26] Bohepilissus sp Last Modified in 08/14/17 | 2 | 209 | 209 | 0 | 21 | 0 | 0 | 1 |
[27] Circellium bacchus Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[28] Dwesasilvasedis medinae Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[29] Coptodactyla nitida Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[30] Amphistomus calcaratus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[31] Aulacopris maximus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[32] Canthonosoma macleayi Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[33] Boletoscapter cornutus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[34] Gen. nov. sp. nov. Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[35] Cephalodesmius armiger Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[36] Coproecus hemisphaericus Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[37] Demarziella interrupta Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[38] Diorygopyx tibialis Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[39] Mentophilus hollandiae Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[40] Onthobium gutierrez Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[41] Pseudonthobium sinuatotibiale Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[42] Tesserodon erratum Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[43] Saphobius sp Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[44] Bdelyrus seminudus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[45] Tesserodoniella elguetai Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[46] Canthonella silphoides Last Modified in 08/14/17 | 7 | 225 | 225 | 0 | 0 | 0 | 0 | 1 |
[47] Cryptocanthon paradoxus Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[48] Paracanthon sp sp n Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[49] Streblopus opatroides Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[50] Zonocopris gibbicollis Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[51] Genieridium margareteae Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[52] Onoreidium howdeni Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[53] Trichillum pauliani Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[54] Scatimus strandi Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[55] Aphengium cupreum Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[56] Agamopus lampros Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[57] Agamopus viridis Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[58] Canthidium bokermanni Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[59] Chalcocopris hesperus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[60] Dichotomius sp Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[61] Dichotomius sericeus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[62] Homocopris torulosus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[63] Ontherus appendiculatus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[64] Uroxys latesulcatus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[65] Uroxys epipleuralis Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[66] Uroxys pauliani Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[67] Bdelyropsis bowditchi Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[68] Canthon virens Last Modified in 08/14/17 | 0 | 232 | 231 | 0 | 0 | 0 | 0 | 1 |
[69] Malagoniella yucateca Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[70] Megathoposoma candezei Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[71] Scybalophagus rugosus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[72] Deltochilum sp Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[73] Scybalocanthon nigriceps Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[74] Hansreia sp Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[75] Silvicanthon bridarollii Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[76] Tetraechma sanguineomaculata Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[77] Anomiopus edmondsi Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[78] Scatonomus fasciculatus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[79] Sinapisoma sp Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[80] Heliocopris sp Last Modified in 08/14/17 | 2 | 230 | 229 | 0 | 0 | 0 | 0 | 1 |
[81] Catharsius sp Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[82] Metacatharsius sp Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[83] Copris sp Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[84] Pseudopedaria grossa Last Modified in 08/14/17 | 0 | 232 | 231 | 0 | 0 | 0 | 0 | 1 |
[85] Microcopris sp Last Modified in 08/14/17 | 0 | 232 | 231 | 0 | 0 | 0 | 0 | 1 |
[86] Gromphas aeruginosa Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[87] Coprophanaeus telamon Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[88] Oruscatus davus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[89] Phanaeus splendidulus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[90] Ennearabdus lobocephalus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[91] Eucranium arachnoides Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[92] Glyphoderus monticola Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[93] Eurysternus hamaticollis Last Modified in 08/14/17 | 0 | 232 | 231 | 0 | 0 | 0 | 0 | 1 |
[94] Pachysoma aesculapius Last Modified in 08/14/17 | 0 | 211 | 211 | 0 | 21 | 0 | 0 | 1 |
[95] Scarabaeus aegyptiorum Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[96] Paragymnopleurus sp Last Modified in 08/14/17 | 1 | 231 | 231 | 0 | 0 | 0 | 0 | 1 |
[97] Gymnopleurus leei Last Modified in 08/14/17 | 1 | 231 | 231 | 0 | 0 | 0 | 0 | 1 |
[98] Epactoides sp1 Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[99] Epactoides sp2 Last Modified in 08/14/17 | 2 | 230 | 230 | 0 | 0 | 0 | 0 | 1 |
[100] Ochicanthon sp Last Modified in 08/14/17 | 0 | 232 | 231 | 0 | 0 | 0 | 0 | 1 |
[101] Apotolamprus quadrimaculatus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[102] Arachnodes sp Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[103] Tanzanolus sp Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[104] Onthophagus avocetta Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[105] Helictopleurus quadripunctatus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[106] Bubas bison Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[107] Onitis sp Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[108] Ateuchus squalidus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[109] Ateuchus histeroides Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[110] Lobateuchus parisii Last Modified in 08/14/17 | 152 | 80 | 80 | 0 | 0 | 0 | 0 | 1 |
[111] Canthochilum tureyra Last Modified in 08/14/17 | 6 | 226 | 226 | 0 | 0 | 0 | 0 | 1 |
[112] Canthochilum oakleyi Last Modified in 08/14/17 | 6 | 226 | 226 | 0 | 0 | 0 | 0 | 1 |
[113] Canthochilum alleni Last Modified in 08/14/17 | 129 | 103 | 103 | 0 | 0 | 0 | 0 | 1 |
[114] Canthochilum philaivii Last Modified in 08/14/17 | 130 | 102 | 102 | 0 | 0 | 0 | 0 | 1 |
[115] Panelus sp Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[116] Parachorius longipenis Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[117] Parachorius solodovnikovi Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[118] Parachorius nudus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[119] Parachorius peninsularis Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[120] Parachorius bolavensis Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[121] Parachorius fukiensis Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[122] Parachorius assymetricus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[123] Parachorius humeralis Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[124] Parachorius gotoi Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[125] Parachorius semsanganus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[126] Parachorius globosus Last Modified in 08/14/17 | 1 | 231 | 231 | 0 | 0 | 0 | 0 | 1 |
[127] Parachorius fungorum Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[128] Parachorius thomsoni Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[129] Parachorius hookeri Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[130] Parachorius maruyamai Last Modified in 08/14/17 | 8 | 224 | 224 | 0 | 0 | 0 | 0 | 1 |
[131] Parachorius newthayerae Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[132] Parachorius pseudojavanus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[133] Parachorius javanus Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
[134] Parachorius schuelkei Last Modified in 08/14/17 | 0 | 232 | 232 | 0 | 0 | 0 | 0 | 1 |
Project views
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Total project views | 24444 | |
Project overview | 3026 | |
Media views | 3527 | Media search (2979 views); M402182 (548 views); |
Views for media list | 1149 | |
Bibliography | 802 | |
Matrix views | 1375 | Matrix landing page (1133 views); Scarabaeinae character matrix (242 views); |
Specimen list | 1835 | |
Taxon list | 12724 | |
Documents list | 6 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 214 | |
Matrix downloads | 12 | Scarabaeinae character matrix (12 downloads); |
Project downloads | 202 |