Project 2607: Rose, K.D., L.T. Holbrook, K. Kumar, R.S. Rana, H.E. Ahrens, R.H. Dunn, A. Folie, K.E. Jones, T. Smith. 2020. Anatomy, relationships, and paleobiology of Cambaytherium (Mammalia, Perissodactylamorpha, Anthracobunia) from the lower Eocene of western India. Journal of Vertebrate Paleontology. 39 (sup1):1-147.
Abstract
The anatomy of Cambaytherium, a primitive, perissodactyl-like mammal from the lower Eocene Cambay Shale Formation of Gujarat, India, is described in detail on the basis of more than 350 specimens that represent almost the entire dentition and the skeleton. Cambaytherium combines plesiomorphic traits typical of archaic ungulates such as phenacodontids with derived traits characteristic of early perissodactyls. Cambaytherium was a subcursorial animal better adapted for running than phenacodontids but less specialized than early perissodactyls. The cheek teeth are bunodont with large upper molar conules, not lophodont as in early perissodactyls; like perissodactyls, however, the lower molars have twinned metaconids and m3 has an extended hypoconulid lobe. A steep wear gradient with heavy wear in the middle of the tooth row suggests an abrasive herbivorous diet. Three species of Cambaytherium are recognized: C. thewissi (∼23 kg), C. gracilis (∼10 kg), and C. marinus (∼99 kg). Body masses were estimated from tooth size and long bone dimensions. Biostratigraphic and isotopic evidence indicates an age of ca. 54.5 Ma for the Cambay Shale vertebrate fauna, the oldest Cenozoic continental vertebrate assemblage from India, near or prior to the initial collision with Asia. Cambaytheriidae (also including Nakusia and Perissobune) and Anthracobunidae are sister taxa, constituting the clade Anthracobunia, which is sister to Perissodactyla. We unite them in a new higher taxon, Perissodactylamorpha. The antiquity and occurrence of Cambaytherium—the most primitive known perissodactylamorph—in India near or before its collision with Asia suggest that Perissodactyla evolved during the Paleocene on the Indian Plate or in peripheral areas of southern or southwestern Asia.Read the article »
Article DOI: 10.1080/02724634.2020.1761370
Project DOI: 10.7934/P2607, http://dx.doi.org/10.7934/P2607
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 33244 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 642 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 827 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:642/0 |
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MorphoBank Project 2607
MorphoBank Project 2607
- Creation Date:
02 February 2017 - Publication Date:
24 November 2020 - Project views: 12022
- Matrix downloads: 22
This research
supported by
Authors' Institutions
- Harvard University
- Royal Belgian Institute of Natural Sciences
- Des Moines University
- Johns Hopkins University
- Rowan University
- Wadia Institute of Himalayan Geology
- H.N.B. Garhwal University
- High Point University
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Luke Holbrook Project Administrator | 74 | 1 | 1 | 642 | 0 | 0 | 33244 (31982, 0, 1262) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M24453' (72 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Eotitanops Last Modified in 11/24/20 | 59 | 257 | 255 | 0 | 7 | 0 | 0 | 1 |
[2] † Palaeosyops Last Modified in 11/24/20 | 9 | 308 | 306 | 0 | 6 | 0 | 0 | 1 |
[3] † Orohippus Last Modified in 11/24/20 | 22 | 298 | 298 | 0 | 1 | 0 | 0 | 1 |
[4] † Epihippus Last Modified in 11/24/20 | 97 | 224 | 223 | 0 | 1 | 0 | 0 | 1 |
[5] † Mesohippus Last Modified in 11/24/20 | 6 | 313 | 313 | 0 | 2 | 0 | 0 | 1 |
[6] † Arenahippus Last Modified in 11/24/20 | 78 | 246 | 242 | 0 | 1 | 0 | 0 | 1 |
[7] † Eohippus Last Modified in 11/24/20 | 45 | 276 | 275 | 0 | 1 | 0 | 0 | 1 |
[8] † Protorohippus Last Modified in 11/24/20 | 22 | 298 | 298 | 0 | 1 | 0 | 0 | 1 |
[9] † Xenicohippus Last Modified in 11/24/20 | 55 | 268 | 265 | 0 | 1 | 0 | 0 | 1 |
[10] † Sifrippus Last Modified in 11/24/20 | 159 | 168 | 160 | 0 | 2 | 0 | 0 | 1 |
[11] † Pliolophus Last Modified in 11/24/20 | 140 | 181 | 180 | 0 | 1 | 0 | 0 | 1 |
[12] † Hyracotherium Last Modified in 11/24/20 | 187 | 133 | 133 | 0 | 1 | 0 | 0 | 1 |
[13] † Palaeotherium Last Modified in 11/24/20 | 12 | 307 | 305 | 0 | 4 | 0 | 0 | 1 |
[14] † Plagiolophus Last Modified in 11/24/20 | 29 | 287 | 285 | 0 | 7 | 0 | 0 | 1 |
[15] † Leptolophus Last Modified in 11/24/20 | 155 | 164 | 164 | 0 | 2 | 0 | 0 | 1 |
[16] † Ghazijhippus Last Modified in 11/24/20 | 190 | 130 | 130 | 0 | 1 | 0 | 0 | 1 |
[17] † Hallensia Last Modified in 11/24/20 | 50 | 269 | 269 | 0 | 2 | 0 | 0 | 1 |
[18] † Propachynolophus Last Modified in 11/24/20 | 187 | 137 | 134 | 0 | 0 | 0 | 0 | 1 |
[19] † Propalaeotherium Last Modified in 11/24/20 | 30 | 296 | 290 | 0 | 1 | 0 | 0 | 1 |
[20] † Lambdotherium Last Modified in 11/24/20 | 74 | 246 | 245 | 0 | 2 | 0 | 0 | 1 |
[21] † Pachynolophus Last Modified in 11/24/20 | 124 | 199 | 197 | 0 | 0 | 0 | 0 | 1 |
[22] † Anchilophus Last Modified in 11/24/20 | 127 | 195 | 193 | 0 | 1 | 0 | 0 | 1 |
[23] † Lophiotherium Last Modified in 11/24/20 | 171 | 155 | 150 | 0 | 0 | 0 | 0 | 1 |
[24] † Homogalax Last Modified in 11/24/20 | 46 | 275 | 274 | 0 | 1 | 0 | 0 | 1 |
[25] † Cardiolophus Last Modified in 11/24/20 | 94 | 229 | 227 | 0 | 0 | 0 | 0 | 1 |
[26] † Eomoropus Last Modified in 11/24/20 | 62 | 256 | 256 | 0 | 3 | 0 | 0 | 1 |
[27] † Litolophus Last Modified in 11/24/20 | 68 | 250 | 250 | 0 | 3 | 0 | 0 | 1 |
[28] † Moropus Last Modified in 11/24/20 | 12 | 289 | 287 | 0 | 22 | 0 | 0 | 1 |
[29] † Lophiodon Last Modified in 11/24/20 | 31 | 276 | 276 | 0 | 14 | 0 | 0 | 1 |
[30] † Isectolophus Last Modified in 11/24/20 | 90 | 231 | 229 | 0 | 2 | 0 | 0 | 1 |
[31] † Heptodon Last Modified in 11/24/20 | 20 | 298 | 297 | 0 | 4 | 0 | 0 | 1 |
[32] † Helaletes Last Modified in 11/24/20 | 14 | 302 | 301 | 0 | 6 | 0 | 0 | 1 |
[33] † Lophialetes Last Modified in 11/24/20 | 40 | 277 | 277 | 0 | 4 | 0 | 0 | 1 |
[34] † Schlosseria Last Modified in 11/24/20 | 119 | 197 | 197 | 0 | 5 | 0 | 0 | 1 |
[35] † Deperetella Last Modified in 11/24/20 | 112 | 197 | 196 | 0 | 13 | 0 | 0 | 1 |
[36] † Teleolophus Last Modified in 11/24/20 | 181 | 131 | 129 | 0 | 11 | 0 | 0 | 1 |
[37] † Hyrachyus Last Modified in 11/24/20 | 2 | 312 | 311 | 0 | 8 | 0 | 0 | 1 |
[38] † Phenacodus Last Modified in 11/24/20 | 5 | 314 | 311 | 0 | 5 | 0 | 0 | 1 |
[39] † Ectocion Last Modified in 11/24/20 | 107 | 211 | 209 | 0 | 5 | 0 | 0 | 1 |
[40] † Tetraclaenodon Last Modified in 11/24/20 | 76 | 242 | 237 | 0 | 8 | 0 | 0 | 1 |
[41] † Meniscotherium Last Modified in 11/24/20 | 7 | 313 | 308 | 0 | 6 | 0 | 0 | 1 |
[42] † Cambaytherium thewissi Last Modified in 11/24/20 | 71 | 244 | 240 | 0 | 10 | 0 | 0 | 1 |
[43] † Cambaytherium gracilis Last Modified in 11/24/20 | 172 | 149 | 147 | 0 | 2 | 0 | 0 | 1 |
[44] † Nakusia Last Modified in 11/24/20 | 279 | 41 | 41 | 0 | 1 | 0 | 0 | 1 |
[45] † Perissobune Last Modified in 11/24/20 | 204 | 120 | 117 | 0 | 0 | 0 | 0 | 1 |
[46] † Anthracobune pinfoldi Last Modified in 11/24/20 | 186 | 132 | 132 | 0 | 3 | 0 | 0 | 1 |
[47] † Anthracobune wardi Last Modified in 11/24/20 | 194 | 128 | 127 | 0 | 0 | 0 | 0 | 1 |
[48] † Obergfellia Last Modified in 11/24/20 | 230 | 91 | 91 | 0 | 0 | 0 | 0 | 1 |
[49] † Radinskya Last Modified in 11/24/20 | 217 | 102 | 102 | 0 | 2 | 0 | 0 | 1 |
[50] † Phenacolophus Last Modified in 11/24/20 | 217 | 102 | 102 | 0 | 2 | 0 | 0 | 1 |
[51] † Olbitherium millenariusum Last Modified in 11/24/20 | 237 | 84 | 84 | 0 | 0 | 0 | 0 | 1 |
[52] † Eritherium Last Modified in 11/24/20 | 174 | 138 | 137 | 0 | 10 | 0 | 0 | 1 |
[53] † Phosphatherium Last Modified in 11/24/20 | 140 | 173 | 172 | 0 | 9 | 0 | 0 | 1 |
[54] † Moeritherium Last Modified in 11/24/20 | 57 | 248 | 245 | 0 | 19 | 0 | 0 | 1 |
[55] † Numidotherium Last Modified in 11/24/20 | 72 | 237 | 237 | 0 | 12 | 0 | 0 | 1 |
[56] † Seggeurius Last Modified in 11/24/20 | 194 | 122 | 122 | 0 | 5 | 0 | 0 | 1 |
[57] † Dimaitherium Last Modified in 11/24/20 | 114 | 205 | 201 | 0 | 6 | 0 | 0 | 1 |
[58] † Microhyrax Last Modified in 11/24/20 | 203 | 117 | 117 | 0 | 1 | 0 | 0 | 1 |
[59] † Saghatherium Last Modified in 11/24/20 | 71 | 245 | 244 | 0 | 6 | 0 | 0 | 1 |
[60] Orycteropus Last Modified in 02/02/17 | 3 | 192 | 191 | 0 | 127 | 0 | 0 | 1 |
[61] Elephantulus Last Modified in 02/02/17 | 13 | 286 | 283 | 0 | 25 | 0 | 0 | 1 |
[62] Petrodromus Last Modified in 02/02/17 | 8 | 294 | 288 | 0 | 25 | 0 | 0 | 1 |
[63] Rhynchocyon Last Modified in 02/02/17 | 14 | 291 | 283 | 0 | 24 | 0 | 0 | 1 |
[64] † Diacodexis Last Modified in 11/24/20 | 19 | 290 | 286 | 0 | 16 | 0 | 0 | 1 |
[65] † Homacodon Last Modified in 11/24/20 | 111 | 203 | 200 | 0 | 10 | 0 | 0 | 1 |
[66] † Vulpavus Last Modified in 11/24/20 | 34 | 268 | 268 | 0 | 19 | 0 | 0 | 1 |
[67] † Paramys Last Modified in 11/24/20 | 10 | 268 | 268 | 0 | 43 | 0 | 0 | 1 |
[68] † Rhombomylus Last Modified in 11/24/20 | 25 | 262 | 259 | 0 | 37 | 0 | 0 | 1 |
[69] † Notharctus Last Modified in 11/24/20 | 20 | 289 | 289 | 0 | 12 | 0 | 0 | 1 |
[70] † Asioryctes Last Modified in 11/24/20 | 56 | 245 | 245 | 0 | 20 | 0 | 0 | 1 |
[71] † Behemotops proteus Last Modified in 11/24/20 | 200 | 119 | 119 | 0 | 2 | 0 | 0 | 1 |
[72] Didelphis Last Modified in 02/02/17 | 23 | 282 | 281 | 0 | 17 | 0 | 0 | 1 |
Taxonomic Overview for Matrix 'M24454' (74 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Eotitanops Last Modified in 11/24/20 | 59 | 257 | 255 | 0 | 7 | 0 | 0 | 1 |
[2] † Palaeosyops Last Modified in 11/24/20 | 9 | 308 | 306 | 0 | 6 | 0 | 0 | 1 |
[3] † Orohippus Last Modified in 11/24/20 | 22 | 298 | 298 | 0 | 1 | 0 | 0 | 1 |
[4] † Epihippus Last Modified in 11/24/20 | 97 | 224 | 223 | 0 | 1 | 0 | 0 | 1 |
[5] † Mesohippus Last Modified in 11/24/20 | 6 | 313 | 313 | 0 | 2 | 0 | 0 | 1 |
[6] † Arenahippus Last Modified in 11/24/20 | 78 | 246 | 242 | 0 | 1 | 0 | 0 | 1 |
[7] † Eohippus Last Modified in 11/24/20 | 45 | 276 | 275 | 0 | 1 | 0 | 0 | 1 |
[8] † Protorohippus Last Modified in 11/24/20 | 22 | 298 | 298 | 0 | 1 | 0 | 0 | 1 |
[9] † Xenicohippus Last Modified in 11/24/20 | 55 | 268 | 265 | 0 | 1 | 0 | 0 | 1 |
[10] † Sifrippus Last Modified in 11/24/20 | 159 | 168 | 160 | 0 | 2 | 0 | 0 | 1 |
[11] † Pliolophus Last Modified in 11/24/20 | 140 | 181 | 180 | 0 | 1 | 0 | 0 | 1 |
[12] † Hyracotherium Last Modified in 11/24/20 | 187 | 133 | 133 | 0 | 1 | 0 | 0 | 1 |
[13] † Palaeotherium Last Modified in 11/24/20 | 12 | 307 | 305 | 0 | 4 | 0 | 0 | 1 |
[14] † Plagiolophus Last Modified in 11/24/20 | 29 | 287 | 285 | 0 | 7 | 0 | 0 | 1 |
[15] † Leptolophus Last Modified in 11/24/20 | 155 | 164 | 164 | 0 | 2 | 0 | 0 | 1 |
[16] † Ghazijhippus Last Modified in 11/24/20 | 190 | 130 | 130 | 0 | 1 | 0 | 0 | 1 |
[17] † Hallensia Last Modified in 11/24/20 | 50 | 269 | 269 | 0 | 2 | 0 | 0 | 1 |
[18] † Propachynolophus Last Modified in 11/24/20 | 187 | 137 | 134 | 0 | 0 | 0 | 0 | 1 |
[19] † Propalaeotherium Last Modified in 11/24/20 | 30 | 296 | 290 | 0 | 1 | 0 | 0 | 1 |
[20] † Lambdotherium Last Modified in 11/24/20 | 74 | 246 | 245 | 0 | 2 | 0 | 0 | 1 |
[21] † Pachynolophus Last Modified in 11/24/20 | 124 | 199 | 197 | 0 | 0 | 0 | 0 | 1 |
[22] † Anchilophus Last Modified in 11/24/20 | 127 | 195 | 193 | 0 | 1 | 0 | 0 | 1 |
[23] † Lophiotherium Last Modified in 11/24/20 | 171 | 155 | 150 | 0 | 0 | 0 | 0 | 1 |
[24] † Homogalax Last Modified in 11/24/20 | 46 | 275 | 274 | 0 | 1 | 0 | 0 | 1 |
[25] † Cardiolophus Last Modified in 11/24/20 | 94 | 229 | 227 | 0 | 0 | 0 | 0 | 1 |
[26] † Eomoropus Last Modified in 11/24/20 | 62 | 256 | 256 | 0 | 3 | 0 | 0 | 1 |
[27] † Litolophus Last Modified in 11/24/20 | 68 | 250 | 250 | 0 | 3 | 0 | 0 | 1 |
[28] † Moropus Last Modified in 11/24/20 | 12 | 289 | 287 | 0 | 22 | 0 | 0 | 1 |
[29] † Lophiodon Last Modified in 11/24/20 | 31 | 276 | 276 | 0 | 14 | 0 | 0 | 1 |
[30] † Isectolophus Last Modified in 11/24/20 | 90 | 231 | 229 | 0 | 2 | 0 | 0 | 1 |
[31] † Heptodon Last Modified in 11/24/20 | 20 | 298 | 297 | 0 | 4 | 0 | 0 | 1 |
[32] † Helaletes Last Modified in 11/24/20 | 14 | 302 | 301 | 0 | 6 | 0 | 0 | 1 |
[33] † Lophialetes Last Modified in 11/24/20 | 40 | 277 | 277 | 0 | 4 | 0 | 0 | 1 |
[34] † Schlosseria Last Modified in 11/24/20 | 119 | 197 | 197 | 0 | 5 | 0 | 0 | 1 |
[35] † Deperetella Last Modified in 11/24/20 | 112 | 197 | 196 | 0 | 13 | 0 | 0 | 1 |
[36] † Teleolophus Last Modified in 11/24/20 | 181 | 131 | 129 | 0 | 11 | 0 | 0 | 1 |
[37] † Hyrachyus Last Modified in 11/24/20 | 2 | 312 | 311 | 0 | 8 | 0 | 0 | 1 |
[38] † Phenacodus Last Modified in 11/24/20 | 5 | 314 | 311 | 0 | 5 | 0 | 0 | 1 |
[39] † Ectocion Last Modified in 11/24/20 | 107 | 211 | 209 | 0 | 5 | 0 | 0 | 1 |
[40] † Tetraclaenodon Last Modified in 11/24/20 | 76 | 242 | 237 | 0 | 8 | 0 | 0 | 1 |
[41] † Meniscotherium Last Modified in 11/24/20 | 7 | 313 | 308 | 0 | 6 | 0 | 0 | 1 |
[42] † Cambaytherium thewissi Last Modified in 11/24/20 | 71 | 244 | 240 | 0 | 10 | 0 | 0 | 1 |
[43] † Cambaytherium gracilis Last Modified in 11/24/20 | 172 | 149 | 147 | 0 | 2 | 0 | 0 | 1 |
[44] † Nakusia Last Modified in 11/24/20 | 279 | 41 | 41 | 0 | 1 | 0 | 0 | 1 |
[45] † Perissobune Last Modified in 11/24/20 | 204 | 120 | 117 | 0 | 0 | 0 | 0 | 1 |
[46] † Anthracobune pinfoldi Last Modified in 11/24/20 | 186 | 132 | 132 | 0 | 3 | 0 | 0 | 1 |
[47] † Anthracobune wardi Last Modified in 11/24/20 | 194 | 128 | 127 | 0 | 0 | 0 | 0 | 1 |
[48] † Obergfellia Last Modified in 11/24/20 | 230 | 91 | 91 | 0 | 0 | 0 | 0 | 1 |
[49] † Radinskya Last Modified in 11/24/20 | 217 | 102 | 102 | 0 | 2 | 0 | 0 | 1 |
[50] † Phenacolophus Last Modified in 11/24/20 | 217 | 102 | 102 | 0 | 2 | 0 | 0 | 1 |
[51] † Minchenella Last Modified in 11/24/20 | 275 | 46 | 46 | 0 | 0 | 0 | 0 | 1 |
[52] † Quettacyonidae Last Modified in 11/24/20 | 228 | 92 | 91 | 0 | 2 | 0 | 0 | 1 |
[53] † Olbitherium millenariusum Last Modified in 11/24/20 | 237 | 84 | 84 | 0 | 0 | 0 | 0 | 1 |
[54] † Eritherium Last Modified in 11/24/20 | 174 | 138 | 137 | 0 | 10 | 0 | 0 | 1 |
[55] † Phosphatherium Last Modified in 11/24/20 | 140 | 173 | 172 | 0 | 9 | 0 | 0 | 1 |
[56] † Moeritherium Last Modified in 11/24/20 | 57 | 248 | 245 | 0 | 19 | 0 | 0 | 1 |
[57] † Numidotherium Last Modified in 11/24/20 | 72 | 237 | 237 | 0 | 12 | 0 | 0 | 1 |
[58] † Seggeurius Last Modified in 11/24/20 | 194 | 122 | 122 | 0 | 5 | 0 | 0 | 1 |
[59] † Dimaitherium Last Modified in 11/24/20 | 114 | 205 | 201 | 0 | 6 | 0 | 0 | 1 |
[60] † Microhyrax Last Modified in 11/24/20 | 203 | 117 | 117 | 0 | 1 | 0 | 0 | 1 |
[61] † Saghatherium Last Modified in 11/24/20 | 71 | 245 | 244 | 0 | 6 | 0 | 0 | 1 |
[62] Orycteropus Last Modified in 02/02/17 | 3 | 192 | 191 | 0 | 127 | 0 | 0 | 1 |
[63] Elephantulus Last Modified in 02/02/17 | 13 | 286 | 283 | 0 | 25 | 0 | 0 | 1 |
[64] Petrodromus Last Modified in 02/02/17 | 8 | 294 | 288 | 0 | 25 | 0 | 0 | 1 |
[65] Rhynchocyon Last Modified in 02/02/17 | 14 | 291 | 283 | 0 | 24 | 0 | 0 | 1 |
[66] † Diacodexis Last Modified in 11/24/20 | 19 | 290 | 286 | 0 | 16 | 0 | 0 | 1 |
[67] † Homacodon Last Modified in 11/24/20 | 111 | 203 | 200 | 0 | 10 | 0 | 0 | 1 |
[68] † Vulpavus Last Modified in 11/24/20 | 34 | 268 | 268 | 0 | 19 | 0 | 0 | 1 |
[69] † Paramys Last Modified in 11/24/20 | 10 | 268 | 268 | 0 | 43 | 0 | 0 | 1 |
[70] † Rhombomylus Last Modified in 11/24/20 | 25 | 262 | 259 | 0 | 37 | 0 | 0 | 1 |
[71] † Notharctus Last Modified in 11/24/20 | 20 | 289 | 289 | 0 | 12 | 0 | 0 | 1 |
[72] † Asioryctes Last Modified in 11/24/20 | 56 | 245 | 245 | 0 | 20 | 0 | 0 | 1 |
[73] † Behemotops proteus Last Modified in 11/24/20 | 200 | 119 | 119 | 0 | 2 | 0 | 0 | 1 |
[74] Didelphis Last Modified in 02/02/17 | 23 | 282 | 281 | 0 | 17 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 12022 | |
Project overview | 1049 | |
Matrix views | 878 | Matrix landing page (601 views); Cambaythere matrix (117 views); Cambaythere matrix (160 views); |
Media views | 2144 | Media search (1811 views); M737892 (333 views); |
Views for media list | 629 | |
Specimen list | 1092 | |
Taxon list | 5796 | |
Bibliography | 429 | |
Documents list | 5 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 161 | |
Matrix downloads | 22 | Cambaythere matrix (11 downloads); Cambaythere matrix (11 downloads); |
Project downloads | 139 |