Project 2626: J. N. Choiniere, C. A. Forster, W. J. de Klerk. 2012. New information on Nqwebasaurus thwazi, a coelurosaurian theropod from the Early Cretaceous Kirkwood Formation in South Africa. Journal of African Earth Sciences. 71:1-17.
Abstract
We performed additional preparation on the holotype skeleton of Nqwebasaurus thwazi and discovered new skeletal material. We describe this material, which includes a maxilla with small, conical, unserrated teeth and bones of the braincase, as well as parts of the holotype postcranial anatomy that were previously poorly documented. We incorporate this new anatomical information into a broadly sampled matrix designed to test theropod relationships. Our phylogenetic results hypothesize that Nqwebasaurus is the basalmost ornithomimosaur, and recover numerous characters supporting this relationship, including features of the maxilla, frontal, dentition, axial skeleton, forelimb and hindlimb. Nqwebasaurus is the first African ornithomimosaur and the first Gondwanan member of this group known from articulated skeletal material, supporting the hypothesis that coelurosaurian groups were cosmopolitan during their early evolutionary history. The presence of reduced dentition and a gastric mill in Nqwebasaurus strongly suggest that this taxon was herbivorous.Read the article »
Article DOI: 10.1016/j.jafrearsci.2012.05.005
Project DOI: 10.7934/P2626, http://dx.doi.org/10.7934/P2626
| This project contains | Matrices |
|---|---|
Download Project SDD File  | Total scored cells: 25550 Total media associated with cells: 0 Total labels associated with cell media: 0 |
| Characters | |
| Total characters: 568 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 1290 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:510/58 |
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MorphoBank Project 2626
MorphoBank Project 2626
- Creation Date:
23 February 2017 - Publication Date:
23 February 2017 - Media downloads: 2
- Matrix downloads: 10
Authors' Institutions
- Albany Museum, Grahamstown
- The George Washington University
- University of the Witwatersrand
- American Museum of Natural History (AMNH)
- Rhodes University
Members
| member name | taxa |
specimens |
media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
| MorphoBank Curator Project Administrator | 97 | 11 | 10 | 568 | 0 | 0 | 25550 (24118, 0, 1432) | 0 | 0 | 0 | ||||
| Jonah Choiniere Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
| Catherine Forster Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Taxonomic Overview for Matrix 'M24524' (97 Taxa)
| taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
| [1] † Herrerasaurus ischigualastensis Taxon name last Modified on 02/23/17 | 189 | 353 | 352 | 0 | 26 | 0 | 0 | 3 |
| [2] † Nqwebasaurus thwazi Taxon name last Modified on 02/23/17 | 325 | 229 | 226 | 0 | 14 | 0 | 0 | 3 |
| [3] † Acrocanthosaurus atokensis Taxon name last Modified on 02/23/17 | 258 | 303 | 300 | 0 | 8 | 0 | 0 | 3 |
| [4] † Afrovenator abakensis Taxon name last Modified on 02/23/17 | 315 | 247 | 246 | 0 | 7 | 0 | 0 | 3 |
| [5] † Albertosaurus sarcophagus Taxon name last Modified on 02/23/17 | 160 | 401 | 401 | 0 | 7 | 0 | 0 | 3 |
| [6] † Allosaurus fragilis Taxon name last Modified on 02/23/17 | 45 | 510 | 501 | 0 | 15 | 0 | 0 | 3 |
| [7] † Alvarezsaurus calvoi Taxon name last Modified on 02/23/17 | 448 | 111 | 101 | 0 | 9 | 0 | 0 | 3 |
| [8] † Alxasaurus elesitaiensis Taxon name last Modified on 02/23/17 | 386 | 178 | 177 | 0 | 4 | 0 | 0 | 3 |
| [9] † Anserimimus planinychus Taxon name last Modified on 02/23/17 | 460 | 103 | 103 | 0 | 5 | 0 | 0 | 3 |
| [10] † Apsaravis ukhaana Taxon name last Modified on 02/23/17 | 345 | 197 | 197 | 0 | 26 | 0 | 0 | 3 |
| [11] † Archaeopteryx lithographica Taxon name last Modified on 02/23/17 | 197 | 349 | 348 | 0 | 22 | 0 | 0 | 3 |
| [12] † Avimimus portentosus Taxon name last Modified on 02/23/17 | 254 | 301 | 301 | 0 | 13 | 0 | 0 | 3 |
| [13] † Beipiaosaurus inexpectus Taxon name last Modified on 02/23/17 | 443 | 116 | 116 | 0 | 9 | 0 | 0 | 3 |
| [14] † Beishanlong grandis Taxon name last Modified on 02/23/17 | 468 | 100 | 100 | 0 | 0 | 0 | 0 | 3 |
| [15] † Buitreraptor gonzalezorum Taxon name last Modified on 02/23/17 | 383 | 179 | 179 | 0 | 6 | 0 | 0 | 3 |
| [16] † Byronosaurus jaffeei Taxon name last Modified on 02/23/17 | 395 | 160 | 159 | 0 | 13 | 0 | 0 | 3 |
| [17] † Carnotaurus sastrei Taxon name last Modified on 02/23/17 | 221 | 326 | 325 | 0 | 21 | 0 | 0 | 3 |
| [18] † Caudipteryx zhoui Taxon name last Modified on 02/23/17 | 304 | 252 | 247 | 0 | 13 | 0 | 0 | 3 |
| [19] † Ceratosaurus nasicornis Taxon name last Modified on 02/23/17 | 168 | 386 | 385 | 0 | 14 | 0 | 0 | 3 |
| [20] † Chirostenotes pergracilis Taxon name last Modified on 02/23/17 | 380 | 175 | 175 | 0 | 13 | 0 | 0 | 3 |
| [21] † Citipati osmolskae Taxon name last Modified on 02/23/17 | 182 | 337 | 336 | 0 | 49 | 0 | 0 | 3 |
| [22] † Coelophysis bauri Taxon name last Modified on 02/23/17 | 227 | 323 | 321 | 0 | 19 | 0 | 0 | 3 |
| [23] † Compsognathus longipes Taxon name last Modified on 02/23/17 | 230 | 322 | 316 | 0 | 16 | 0 | 0 | 3 |
| [24] † Conchoraptor gracilis Taxon name last Modified on 02/23/17 | 372 | 179 | 179 | 0 | 17 | 0 | 0 | 3 |
| [25] † Confuciusornis sanctus Taxon name last Modified on 02/23/17 | 273 | 246 | 246 | 0 | 49 | 0 | 0 | 3 |
| [26] † Cryolophosaurus ellioti Taxon name last Modified on 02/23/17 | 414 | 150 | 147 | 0 | 6 | 0 | 0 | 3 |
| [27] † Daspletosaurus torosus Taxon name last Modified on 02/23/17 | 358 | 202 | 201 | 0 | 8 | 0 | 0 | 3 |
| [28] † Deinonychus antirrhopus Taxon name last Modified on 02/23/17 | 134 | 420 | 420 | 0 | 14 | 0 | 0 | 3 |
| [29] † Dilong paradoxus Taxon name last Modified on 02/23/17 | 228 | 324 | 321 | 0 | 16 | 0 | 0 | 3 |
| [30] † Dilophosaurus wetherilli Taxon name last Modified on 02/23/17 | 206 | 348 | 345 | 0 | 16 | 0 | 0 | 3 |
| [31] † Dromaeosaurus albertensis Taxon name last Modified on 02/23/17 | 399 | 159 | 159 | 0 | 10 | 0 | 0 | 3 |
| [32] † EK troodontid Taxon name last Modified on 02/23/17 | 516 | 50 | 50 | 0 | 2 | 0 | 0 | 3 |
| [33] † Eoraptor lunensis Taxon name last Modified on 02/23/17 | 323 | 222 | 222 | 0 | 23 | 0 | 0 | 3 |
| [34] † Epidexipteryx hui Taxon name last Modified on 02/23/17 | 389 | 171 | 171 | 0 | 8 | 0 | 0 | 3 |
| [35] † Erlikosaurus andrewsi Taxon name last Modified on 02/23/17 | 367 | 179 | 178 | 0 | 22 | 0 | 0 | 3 |
| [36] † Eustreptospondylus oxiensis Taxon name last Modified on 02/23/17 | 195 | 357 | 355 | 0 | 17 | 0 | 0 | 3 |
| [37] † Falcarius utahensis Taxon name last Modified on 02/23/17 | 185 | 378 | 376 | 0 | 6 | 0 | 0 | 3 |
| [38] † Gallimimus bullatus Taxon name last Modified on 02/23/17 | 192 | 338 | 338 | 0 | 38 | 0 | 0 | 3 |
| [39] † Garudimimus brevipes Taxon name last Modified on 02/23/17 | 196 | 326 | 326 | 0 | 46 | 0 | 0 | 3 |
| [40] † Giganotosaurus carolinii Taxon name last Modified on 02/23/17 | 373 | 193 | 191 | 0 | 4 | 0 | 0 | 3 |
| [41] † Gigantoraptor erlianensis Taxon name last Modified on 02/23/17 | 467 | 86 | 86 | 0 | 15 | 0 | 0 | 3 |
| [42] † Gorgosaurus libratus Taxon name last Modified on 02/23/17 | 308 | 255 | 255 | 0 | 5 | 0 | 0 | 3 |
| [43] † Guanlong wucaii Taxon name last Modified on 02/23/17 | 68 | 487 | 476 | 0 | 14 | 0 | 0 | 3 |
| [44] † Haplocheirus sollers Taxon name last Modified on 02/23/17 | 109 | 444 | 437 | 0 | 16 | 0 | 0 | 3 |
| [45] † Harpymimus okladnikovi Taxon name last Modified on 02/23/17 | 226 | 306 | 305 | 0 | 36 | 0 | 0 | 3 |
| [46] † Huaxiagnathus orientalis Taxon name last Modified on 02/23/17 | 313 | 242 | 242 | 0 | 13 | 0 | 0 | 3 |
| [47] † Incisivosaurus gauthieri Taxon name last Modified on 02/23/17 | 348 | 207 | 207 | 0 | 13 | 0 | 0 | 3 |
| [48] † Limusaurus inextricabilis Taxon name last Modified on 02/23/17 | 213 | 301 | 298 | 0 | 56 | 0 | 0 | 3 |
| [49] † Linheraptor exquisitus Taxon name last Modified on 02/23/17 | 321 | 236 | 236 | 0 | 11 | 0 | 0 | 3 |
| [50] † Mahakala omnogovae Taxon name last Modified on 02/23/17 | 478 | 84 | 83 | 0 | 6 | 0 | 0 | 3 |
| [51] † Majungasaurus crenatissimus Taxon name last Modified on 02/23/17 | 164 | 392 | 391 | 0 | 12 | 0 | 0 | 3 |
| [52] † Mapusaurus roseae Taxon name last Modified on 02/23/17 | 380 | 186 | 186 | 0 | 2 | 0 | 0 | 3 |
| [53] † Masiakasaurus knopfleri Taxon name last Modified on 02/23/17 | 385 | 172 | 167 | 0 | 11 | 0 | 0 | 3 |
| [54] † Megalosaurus bucklandii Taxon name last Modified on 02/23/17 | 376 | 185 | 184 | 0 | 8 | 0 | 0 | 3 |
| [55] † Mei long Taxon name last Modified on 02/23/17 | 242 | 301 | 301 | 0 | 25 | 0 | 0 | 3 |
| [56] † Microraptor gui Taxon name last Modified on 02/23/17 | 330 | 223 | 223 | 0 | 15 | 0 | 0 | 3 |
| [57] † Microvenator celer Taxon name last Modified on 02/23/17 | 382 | 179 | 176 | 0 | 8 | 0 | 0 | 3 |
| [58] † Monolophosaurus jiangi Taxon name last Modified on 02/23/17 | 265 | 291 | 290 | 0 | 12 | 0 | 0 | 3 |
| [59] † Mononykus olecranus Taxon name last Modified on 02/23/17 | 319 | 228 | 228 | 0 | 21 | 0 | 0 | 3 |
| [60] † Neovenator salerii Taxon name last Modified on 02/23/17 | 261 | 303 | 301 | 0 | 4 | 0 | 0 | 3 |
| [61] † Nothronychus graffami Taxon name last Modified on 02/23/17 | 384 | 180 | 180 | 0 | 4 | 0 | 0 | 3 |
| [62] † Ornitholestes hermanni Taxon name last Modified on 02/23/17 | 224 | 330 | 325 | 0 | 15 | 0 | 0 | 3 |
| [63] † Ornithomimus edmontonicus Taxon name last Modified on 02/23/17 | 264 | 283 | 283 | 0 | 21 | 0 | 0 | 3 |
| [64] † Oviraptor philoceratops Taxon name last Modified on 02/23/17 | 433 | 120 | 120 | 0 | 15 | 0 | 0 | 3 |
| [65] † Parvicursor remotus Taxon name last Modified on 02/23/17 | 439 | 112 | 109 | 0 | 19 | 0 | 0 | 3 |
| [66] † Patagonykus puertai Taxon name last Modified on 02/23/17 | 408 | 155 | 142 | 0 | 6 | 0 | 0 | 3 |
| [67] † Pelecanimimus polyodon Taxon name last Modified on 02/23/17 | 317 | 237 | 230 | 0 | 14 | 0 | 0 | 3 |
| [68] † Piatnitzkysaurus floresi Taxon name last Modified on 02/23/17 | 395 | 169 | 168 | 0 | 4 | 0 | 0 | 3 |
| [69] † Proceratosaurus bradleyi Taxon name last Modified on 02/23/17 | 415 | 147 | 145 | 0 | 6 | 0 | 0 | 3 |
| [70] † Rahonavis ostromi Taxon name last Modified on 02/23/17 | 350 | 212 | 212 | 0 | 6 | 0 | 0 | 3 |
| [71] † Rinchenia mongoliensis Taxon name last Modified on 02/23/17 | 453 | 101 | 101 | 0 | 14 | 0 | 0 | 3 |
| [72] † Sapeornis chaoyangensis Taxon name last Modified on 02/23/17 | 259 | 278 | 278 | 0 | 31 | 0 | 0 | 3 |
| [73] † Saurornithoides mongoliensis Taxon name last Modified on 02/23/17 | 406 | 155 | 155 | 0 | 7 | 0 | 0 | 3 |
| [74] † Saurornitholestes langstoni Taxon name last Modified on 02/23/17 | 363 | 201 | 196 | 0 | 4 | 0 | 0 | 3 |
| [75] † Segnosaurus galboensis Taxon name last Modified on 02/23/17 | 424 | 141 | 141 | 0 | 3 | 0 | 0 | 3 |
| [76] † Shenzhousaurus orientalis Taxon name last Modified on 02/23/17 | 313 | 225 | 225 | 0 | 30 | 0 | 0 | 3 |
| [77] † Shuvuuia deserti Taxon name last Modified on 02/23/17 | 78 | 442 | 438 | 0 | 48 | 0 | 0 | 3 |
| [78] † Sinornithoides youngi Taxon name last Modified on 02/23/17 | 403 | 157 | 157 | 0 | 8 | 0 | 0 | 3 |
| [79] † Sinornithosaurus millenii Taxon name last Modified on 02/23/17 | 323 | 232 | 232 | 0 | 13 | 0 | 0 | 3 |
| [80] † Sinosauropteryx prima Taxon name last Modified on 02/23/17 | 262 | 292 | 292 | 0 | 14 | 0 | 0 | 3 |
| [81] † Sinovenator changii Taxon name last Modified on 02/23/17 | 316 | 248 | 248 | 0 | 4 | 0 | 0 | 3 |
| [82] † Sinraptor dongi Taxon name last Modified on 02/23/17 | 109 | 445 | 441 | 0 | 14 | 0 | 0 | 3 |
| [83] † Stokesosaurus langhami Taxon name last Modified on 02/23/17 | 445 | 122 | 118 | 0 | 1 | 0 | 0 | 3 |
| [84] † Struthiomimus altus Taxon name last Modified on 02/23/17 | 146 | 376 | 376 | 0 | 46 | 0 | 0 | 3 |
| [85] † Suchomimus tenerensis Taxon name last Modified on 02/23/17 | 375 | 187 | 185 | 0 | 8 | 0 | 0 | 3 |
| [86] † Syntarsus rhodesiensis Taxon name last Modified on 02/23/17 | 180 | 371 | 368 | 0 | 18 | 0 | 0 | 3 |
| [87] † Tanycolagreus topwilsoni Taxon name last Modified on 02/23/17 | 328 | 237 | 235 | 0 | 3 | 0 | 0 | 3 |
| [88] † Tarbosaurus bataar Taxon name last Modified on 02/23/17 | 361 | 199 | 199 | 0 | 8 | 0 | 0 | 3 |
| [89] † Tawa hallae Taxon name last Modified on 02/23/17 | 284 | 264 | 264 | 0 | 20 | 0 | 0 | 3 |
| [90] † Torvosaurus tanneri Taxon name last Modified on 02/23/17 | 344 | 215 | 214 | 0 | 9 | 0 | 0 | 3 |
| [91] † Troodon formosus Taxon name last Modified on 02/23/17 | 269 | 284 | 281 | 0 | 15 | 0 | 0 | 3 |
| [92] † Tsaagan mangas Taxon name last Modified on 02/23/17 | 350 | 205 | 205 | 0 | 13 | 0 | 0 | 3 |
| [93] † Tyrannosaurus rex Taxon name last Modified on 02/23/17 | 32 | 518 | 516 | 0 | 18 | 0 | 0 | 3 |
| [94] † Unenlagia comahuensis Taxon name last Modified on 02/23/17 | 446 | 120 | 112 | 0 | 2 | 0 | 0 | 3 |
| [95] † Velociraptor mongoliensis Taxon name last Modified on 02/23/17 | 118 | 431 | 431 | 0 | 19 | 0 | 0 | 3 |
| [96] † Zanabazar junior Taxon name last Modified on 02/23/17 | 378 | 179 | 179 | 0 | 11 | 0 | 0 | 3 |
| [97] † Zuolong salleei Taxon name last Modified on 02/23/17 | 321 | 240 | 237 | 0 | 7 | 0 | 0 | 3 |
Project downloads
| type | number of downloads | Individual items downloaded (where applicable) |
| Total downloads from project | 378 | |
| Project downloads | 359 | |
| Document downloads | 7 | Matrix (3 downloads); character names and states (4 downloads); |
| Matrix downloads | 10 | matrix (10 downloads); |
| Media downloads | 2 | M437349 (2 downloads); |