Project 2627: E. Vlachos, M. Rabi. 2017. Total evidence analysis and body size evolution of extant and extinct tortoises (Testudines: Cryptodira: Pan-Testudinidae). Cladistics. online (online):1-35.
Specimen: Siebenrockiella crassicollis (unvouchered)
View: Dorsal

Abstract

Testudinidae (tortoises) is an extant clade of terrestrial turtles of worldwide distribution and with a rich fossil record that provides an exceptional context for studying their evolutionary history. Because of the lack of global phylogenetic analyses integrating extinct taxa, our current knowledge of the relationships of the total clade of Testudinidae is rather poor. To resolve this issue, we performed the first total evidence analysis of Pan-Testudinidae. The total evidence trees are congruent with the molecular topology and agree on the dichotomy of derived Testudinidae (=Testudininae; Converted Clade Name) into two previously recognized major clades, Testudona and Geochelona (New Clade Name). The integration of extinct taxa into the analysis allowed the stratigraphic fit of the total evidence trees, indicating that crown Testudininae, Testudona and Geochelona all originated by the Late Eocene, in agreement with recent molecular estimates. Ghost lineage analysis indicates high diversification in the Late Eocene and in the Miocene. The age of crown Testudo is Late Miocene, again in accordance with some molecular dates. Phylogenetic placement of fossils demonstrates that giant body size independently evolved in multiple continental mainland taxa and confirms recent results deduced from living taxa


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Article DOI: 10.1111/cla.12227

Project DOI: 10.7934/P2627, http://dx.doi.org/10.7934/P2627
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 13 Documents
  • 72 Taxa
  • 1 Specimen
  • 169 Characters
Total size of project's media files: 1 MB

Download Project SDD File
Total scored cells: 7855
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 169
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 395
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:144/25
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MorphoBank Project 2627


    Members

    member name taxa specimens media media
    notes
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    notes  media
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    Evangelos Vlachos
    Project Administrator
    Last logged in 11/17/17
    7211116901007855
    (7576, 0, 279)
    041000


    Taxonomic Overview for Matrix 'M24521' (72 Taxa)

    taxon unscored
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    "-" cellscell images labels on
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    [1] Pleurodira
    Last Modified in 11/17/17
    23147125021001
    [2] Chelydra serpentina
    Last Modified in 11/17/17
    5150148016001
    [3] Kinosternon odoratus
    Last Modified in 11/17/17
    3154154012001
    [4] Caretta caretta
    Last Modified in 11/17/17
    72888809001
    [5] Apalone spinifera
    Last Modified in 11/17/17
    279494048001
    [6] Platysternon megacephalum
    Last Modified in 11/17/17
    1514714707001
    [7] Batagur baska
    Last Modified in 11/17/17
    416316104001
    [8] Mauremys caspica
    Last Modified in 11/17/17
    716015605001
    [9] Siebenrockiella crassicollis
    Last Modified in 11/17/17
    1614914706001
    [10] Mauremys sinensis
    Last Modified in 11/17/17
    4012712504001
    [11] Chrysemys picta
    Last Modified in 11/17/17
    116416107001
    [12] Emys orbicularis
    Last Modified in 11/17/17
    3113313008001
    [13] Trachemys scripta
    Last Modified in 11/17/17
    815915704001
    [14] Gopherus agassizii
    Last Modified in 02/23/17
    816315803001
    [15] Chelonoidis carbonaria
    Last Modified in 11/17/17
    018615403001
    [16] Chelonoidis chilensis
    Last Modified in 11/17/17
    418216203001
    [17] Chelonoidis denticulata
    Last Modified in 11/17/17
    019416503001
    [18] Geochelone elegans
    Last Modified in 02/23/17
    5311611303001
    [19] Indotestudo elongata
    Last Modified in 02/23/17
    2614414102001
    [20] Manouria emys
    Last Modified in 11/17/17
    701009701001
    [21] Kinixys erosa
    Last Modified in 11/17/17
    12147139011001
    [22] Indotestudo forstenii
    Last Modified in 11/17/17
    316416203001
    [23] Testudo graeca
    Last Modified in 11/17/17
    217816204001
    [24] Chersine hermanni
    Last Modified in 02/23/17
    616716002001
    [25] Kinixys homeana
    Last Modified in 11/17/17
    2913313208001
    [26] Agrionemys horsfieldii
    Last Modified in 02/23/17
    5811510902001
    [27] Manouria impressa
    Last Modified in 11/17/17
    2913713703001
    [28] Testudo kleinmanni
    Last Modified in 11/17/17
    3113813205001
    [29] Testudo marginata
    Last Modified in 11/17/17
    79968802001
    [30] Chelonoidis nigra
    Last Modified in 11/17/17
    217216403001
    [31] Psammobates oculifera
    Last Modified in 02/23/17
    73969501001
    [32] Stigmochelys pardalis
    Last Modified in 11/17/17
    1315615303001
    [33] Gopherus polyphemus
    Last Modified in 11/17/17
    616416003001
    [34] Astrochelys radiata
    Last Modified in 11/17/17
    915915802001
    [35] Centrochelys sulcata
    Last Modified in 02/23/17
    217016402001
    [36] Malacochersus tornieri
    Last Modified in 11/17/17
    2151149018001
    [37] † Gigantochersina ammon
    Last Modified in 02/23/17
    113625600001
    [38] † Testudo antiqua
    Last Modified in 02/23/17
    106656101001
    [39] † Megalochelys atlas
    Last Modified in 02/23/17
    133403500001
    [40] † Titanochelon bacharidisi
    Last Modified in 02/23/17
    701019602001
    [41] † Hesperotestudo bermudae
    Last Modified in 02/23/17
    99737000001
    [42] † Titanochelon bolivari
    Last Modified in 02/23/17
    3513613103001
    [43] † Ergilemys bruneti
    Last Modified in 02/23/17
    111595700001
    [44] † Paleotestudo canetotiana
    Last Modified in 02/23/17
    112575601001
    [45] † Achilemys cassouleti
    Last Modified in 02/23/17
    103666400001
    [46] † Hadrianus castrensis
    Last Modified in 02/23/17
    113555301001
    [47] † Hadrianus corsoni
    Last Modified in 11/17/17
    104656201001
    [48] † Geochelone costaricensis
    Last Modified in 02/23/17
    112575700001
    [49] † Hesperotestudo crassiscutata
    Last Modified in 11/17/17
    107626000001
    [50] † Testudo eocaenica
    Last Modified in 02/23/17
    114625500001
    [51] † Taraschelon gigas
    Last Modified in 02/23/17
    125414103001
    [52] † Chelonoidis gringorum
    Last Modified in 02/23/17
    130373702001
    [53] † Chelonoidis hesterna
    Last Modified in 02/23/17
    108595902001
    [54] † Ergilemys insolitus
    Last Modified in 02/23/17
    123514600001
    [55] † Testudo kaiseni
    Last Modified in 02/23/17
    127414101001
    [56] † Oligopherus laticuneus
    Last Modified in 02/23/17
    91777701001
    [57] † Testudo lunellensis
    Last Modified in 02/23/17
    111575701001
    [58] † Testudo marmorum
    Last Modified in 02/23/17
    111575701001
    [59] † Cheirogaster maurini
    Last Modified in 02/23/17
    116505003001
    [60] † Namibchersus namaquensis
    Last Modified in 02/23/17
    108616100001
    [61] † Stylemys nebrascensis
    Last Modified in 11/17/17
    5511511301001
    [62] † Mesochersus orangeus
    Last Modified in 02/23/17
    135363301001
    [63] † Hesperotestudo osborniana
    Last Modified in 02/23/17
    6011110900001
    [64] † Testudo oughlamensis
    Last Modified in 02/23/17
    119524901001
    [65] † Impregnochelys pachytestis
    Last Modified in 02/23/17
    130393900001
    [66] † Titanochelon perpiniana
    Last Modified in 02/23/17
    87797704001
    [67] † Testudo promarginata
    Last Modified in 02/23/17
    111585800001
    [68] † Testudo sharanensis
    Last Modified in 02/23/17
    115535201001
    [69] † Testudo shensiensis
    Last Modified in 02/23/17
    102676601001
    [70] † Anhuichelys siaoshihensis
    Last Modified in 02/23/17
    105606004001
    [71] † Pelorochelon soriana
    Last Modified in 02/23/17
    106656300001
    [72] † Titanochelon vitodurana
    Last Modified in 02/23/17
    81878602001


    Project views

    type number of views Individual items viewed (where applicable)
    Total project views3088
    Project overview488
    Taxon list1153
    Documents list206
    Media views493Media search (410 views); M464981 (83 views);
    Matrix views208Matrix landing page (167 views); Vlachos and Rabi, Cladistics; Nexus file of morphological taxon-character matrix (41 views);
    Specimen list271
    Views for media list152
    Bibliography117




    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project65
    Document downloads16Suppl. File S8 (2 downloads); Suppl. File S5 (4 downloads); Suppl. File S6 (3 downloads); Body Size matrix (3 downloads); Suppl. File S4 (1 download); Suppl. File S7 (1 download); Trees of morphological analysis, with constrains (1 download); Total evidence MP trees (1 download);
    Project downloads46
    Matrix downloads3Vlachos and Rabi, Cladistics; Nexus file of morphological taxon-character matrix (3 downloads);