Project 2662: E. Gorscak, P. M. O'Connor, E. M. Roberts, N. J. Stevens. 2017. The second titanosaurian (Dinosauria: Sauropoda) from the middle Cretaceous Galula Formation, southwestern Tanzania, with remarks on African titanosaurian diversity. Journal of Vertebrate Paleontology. 37 (4):e1343250.
Specimen: † Shingopana songwensis (RRBP/02100)
View: anterior


The paleobiogeographic significance of continental Africa during the middle and Late Cretaceous is not well understood, in part due to incomplete sampling from large portions of the landmass during these intervals. Intensified field efforts in the Galula Formation exposed in southwestern Tanzania have revealed a diverse vertebrate fauna, including the novel titanosaurian Shingopana songwensis, gen. et sp. nov., described herein. Based on a left angular, cervical vertebrae, cervical and dorsal ribs, a left humerus, and a partial left pubis, Shingopana exhibits morphology indicating affinities with the Late Cretaceous aeolosaurine titanosaurians of South America. The bulbous expansion of the cervical vertebral neural spine is similar to the condition in Bonitasaura salgadoi, Overosaurus paradasorum, and Trigonosaurus pricei. The dorsal ribs of Shingopana also present proximal anterior and posterior flanges that previously were proposed to be unique to Overosaurus. Furthermore, Shingopana is diagnosed by a divided spinoprezygapophyseal lamina in the middle-to-posterior cervical vertebrae. Parsimony and both uncalibrated and tip-dated Bayesian phylogenetic approaches support Shingopana as the first African titanosaurian that is closely related to aeolosaurines. Comparisons with other African titanosaurians, such as the co-occurring Rukwatitan bisepultus and geographically proximate Malawisaurus dixeyi, suggest that southern African forms represent diverse taxa rather than forming a monophyletic group. Moreover, southern African forms exhibit stronger affinities with South American clades than with representative northern African form, suggesting that tectonically driven separation of the two landmasses may have influenced the development of progressively isolated southern African faunas throughout the Cretaceous.

Read the article »

Article DOI: 10.1080/02724634.2017.1343250

Project DOI: 10.7934/P2662,
This project contains
  • 28 Media
  • 5 Documents
  • 5 Folios
  • 1 Taxon
  • 1 Specimen
Total size of project's media files: 116.69M

Download Project SDD File
Currently Viewing:
MorphoBank Project 2662

    This research
    supported by

    Authors' Institutions

    • Ohio University

    • James Cook University


    member name taxa specimens media
    Eric Gorscak
    Project Administrator

    Project has no matrices defined.

    Project views

    type number of views Individual items viewed (where applicable)
    Total project views15948
    Project overview1520
    Media views8392Media search (1692 views); M451712 (238 views); M451729 (229 views); M451730 (253 views); M451736 (248 views); M451721 (251 views); M451733 (240 views); M451734 (223 views); M451711 (247 views); M451715 (243 views); M451713 (210 views); M451727 (340 views); M451738 (239 views); M451735 (224 views); M451731 (252 views); M451728 (245 views); M451726 (215 views); M451718 (214 views); M451714 (219 views); M451737 (254 views); M451725 (236 views); M451720 (221 views); M451719 (229 views); M451717 (280 views); M451723 (224 views); M451722 (226 views); M451724 (217 views); M451732 (241 views); M451716 (242 views);
    Documents list516
    Taxon list854
    Folio views1175Folio list (407 views); Humerus (158 views); Cervical Ribs (170 views); Cervical Vertebra A (143 views); Dorsal Ribs (151 views); Cervical Vertebra B (146 views);
    Specimen list813
    Views for media list2073

    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project131
    Media downloads1M451711 (1 download);
    Project downloads126
    Document downloads4Nexus File for Parsimony/Uncalibrated Bayesian Analysis (4 downloads);