Project 2662: E. Gorscak, P. M. O'Connor, E. M. Roberts, N. J. Stevens. 2017. The second titanosaurian (Dinosauria: Sauropoda) from the middle Cretaceous Galula Formation, southwestern Tanzania with remarks on African titanosaurian diversity. Journal of Vertebrate Paleontology. 30:e1343250–23.
Specimen: † Shingopana songwensis (RRBP/02100)
View: anterior


The paleobiogeographic significance of continental Africa during the middle and Late Cretaceous is not well understood, in part due to incomplete sampling from large portions of the landmass during these intervals. Intensified field efforts in the Galula Formation exposed in southwestern Tanzania have revealed a diverse vertebrate fauna, including the novel titanosaurian Shingopana songwensis, gen. et sp. nov., described herein. Based on a left angular, cervical vertebrae, cervical and dorsal ribs, a left humerus, and a partial left pubis, Shingopana exhibits morphology indicating affinities with the Late Cretaceous aeolosaurine titanosaurians of South America. The bulbous expansion of the cervical vertebral neural spine is similar to the condition in Bonitasaura salgadoi, Overosaurus paradasorum, and Trigonosaurus pricei. The dorsal ribs of Shingopana also present proximal anterior and posterior flanges that previously were proposed to be unique to Overosaurus. Furthermore, Shingopana is diagnosed by a divided spinoprezygapophyseal lamina in the middle-to-posterior cervical vertebrae. Parsimony and both uncalibrated and tip-dated Bayesian phylogenetic approaches support Shingopana as the first African titanosaurian that is closely related to aeolosaurines. Comparisons with other African titanosaurians, such as the co-occurring Rukwatitan bisepultus and geographically proximate Malawisaurus dixeyi, suggest that southern African forms represent diverse taxa rather than forming a monophyletic group. Moreover, southern African forms exhibit stronger affinities with South American clades than with representative northern African form, suggesting that tectonically driven separation of the two landmasses may have influenced the development of progressively isolated southern African faunas throughout the Cretaceous.

Read the article »

Project DOI: 10.7934/P2662,
This project contains
  • 28 Media
  • 5 Documents
  • 5 Folios
  • 1 Taxon
  • 1 Specimen
Total size of project's media files: 129 MB

Download Project SDD File
Currently Viewing:
MorphoBank Project 2662

    This research
    supported by


    member name taxa specimens media
    Eric Gorscak
    Project Administrator
    Last logged in 08/28/17

    Project has no matrices defined.

    Project views

    type number of views Individual items viewed (where applicable)
    Total project views6777
    Project overview797
    Media views3714Media search (744 views); M451712 (111 views); M451729 (103 views); M451730 (104 views); M451736 (122 views); M451721 (98 views); M451733 (98 views); M451734 (102 views); M451711 (115 views); M451715 (104 views); M451713 (79 views); M451727 (197 views); M451738 (101 views); M451735 (96 views); M451731 (100 views); M451728 (103 views); M451726 (96 views); M451718 (85 views); M451714 (94 views); M451737 (106 views); M451725 (116 views); M451720 (93 views); M451719 (108 views); M451717 (126 views); M451723 (91 views); M451722 (102 views); M451724 (95 views); M451732 (103 views); M451716 (122 views);
    Documents list245
    Taxon list371
    Folio views409Folio list (173 views); Humerus (54 views); Cervical Ribs (47 views); Cervical Vertebra A (43 views); Dorsal Ribs (48 views); Cervical Vertebra B (44 views);
    Specimen list339
    Views for media list738

    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project39
    Media downloads1M451711 (1 download);
    Project downloads34
    Document downloads4Nexus File for Parsimony/Uncalibrated Bayesian Analysis (4 downloads);