Project 2662: E. Gorscak, P. M. O'Connor, E. M. Roberts, N. J. Stevens. 2017. The second titanosaurian (Dinosauria: Sauropoda) from the middle Cretaceous Galula Formation, southwestern Tanzania, with remarks on African titanosaurian diversity. Journal of Vertebrate Paleontology. 37 (4):e1343250.
Specimen: † Shingopana songwensis (RRBP/02100)
View: anterior


The paleobiogeographic significance of continental Africa during the middle and Late Cretaceous is not well understood, in part due to incomplete sampling from large portions of the landmass during these intervals. Intensified field efforts in the Galula Formation exposed in southwestern Tanzania have revealed a diverse vertebrate fauna, including the novel titanosaurian Shingopana songwensis, gen. et sp. nov., described herein. Based on a left angular, cervical vertebrae, cervical and dorsal ribs, a left humerus, and a partial left pubis, Shingopana exhibits morphology indicating affinities with the Late Cretaceous aeolosaurine titanosaurians of South America. The bulbous expansion of the cervical vertebral neural spine is similar to the condition in Bonitasaura salgadoi, Overosaurus paradasorum, and Trigonosaurus pricei. The dorsal ribs of Shingopana also present proximal anterior and posterior flanges that previously were proposed to be unique to Overosaurus. Furthermore, Shingopana is diagnosed by a divided spinoprezygapophyseal lamina in the middle-to-posterior cervical vertebrae. Parsimony and both uncalibrated and tip-dated Bayesian phylogenetic approaches support Shingopana as the first African titanosaurian that is closely related to aeolosaurines. Comparisons with other African titanosaurians, such as the co-occurring Rukwatitan bisepultus and geographically proximate Malawisaurus dixeyi, suggest that southern African forms represent diverse taxa rather than forming a monophyletic group. Moreover, southern African forms exhibit stronger affinities with South American clades than with representative northern African form, suggesting that tectonically driven separation of the two landmasses may have influenced the development of progressively isolated southern African faunas throughout the Cretaceous.

Read the article »

Article DOI: 10.1080/02724634.2017.1343250

Project DOI: 10.7934/P2662,
This project contains
  • 28 Media
  • 5 Documents
  • 5 Folios
  • 1 Taxon
  • 1 Specimen
Total size of project's media files: 116.69M

Download Project SDD File
Currently Viewing:
MorphoBank Project 2662

    This research
    supported by

    Authors' Institutions

    • Ohio University

    • James Cook University


    member name taxa specimens media
    Eric Gorscak
    Project Administrator

    Project has no matrices defined.

    Project views

    type number of views Individual items viewed (where applicable)
    Total project views19767
    Project overview1768
    Media views10386Media search (2029 views); M451712 (296 views); M451729 (286 views); M451730 (311 views); M451736 (307 views); M451721 (315 views); M451733 (299 views); M451734 (280 views); M451711 (305 views); M451715 (299 views); M451713 (273 views); M451727 (400 views); M451738 (299 views); M451735 (283 views); M451731 (312 views); M451728 (302 views); M451726 (278 views); M451718 (268 views); M451714 (269 views); M451737 (311 views); M451725 (295 views); M451720 (280 views); M451719 (284 views); M451717 (342 views); M451723 (282 views); M451722 (307 views); M451724 (279 views); M451732 (297 views); M451716 (298 views);
    Documents list618
    Taxon list995
    Folio views1469Folio list (486 views); Humerus (200 views); Cervical Ribs (218 views); Cervical Vertebra A (188 views); Dorsal Ribs (191 views); Cervical Vertebra B (186 views);
    Specimen list1024
    Views for media list2684

    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project176
    Media downloads1M451711 (1 download);
    Project downloads171
    Document downloads4Nexus File for Parsimony/Uncalibrated Bayesian Analysis (4 downloads);