Project 2707: L. Ramsköld, B. D. Chatterton. 1991. Revision and subdivision of the polyphyletic ‘Leonaspis’ (Trilobita). Transactions of the Royal Society of Edinburgh: Earth Sciences. 82 (4):333-371.
Abstract
A phylogenetic analysis is performed on all species previously assigned to the odontopleurid trilobite genus Leonaspis. The PAUP analysis shows this genus, as conventionally defined, to be polyphyletic, composed of four groups of equal taxonomic rank. Leonaspis (s.s.)is here restricted to one of these groups only, a monophyletic set of species characterised by 'four-spined' pygidia (i.e. with four spines between the major border spines) and a thorax of nine segments. Kettneraspis is recognised for the largest of these four groups, and is composed of 'two-spined' species with nine thoracic segments. Leonaspis and Kettneraspis belong to the Odontopleurinae. A third group of previous 'Leonaspis' species constitute a new genus of the Acidaspidinae. Its species are characterised by being 'four-spined' and having ten thoracic segments. For these the new genus Exallaspis is erected, with type species E. bufo. A fourth group of 'Leonaspis' species, being 'four-spined' with nine thoracic segments, belongs to Eoleonaspis, an Ordovician odontopleurine genus. Leonaspis and Exallaspis are temporally and spatially non-overlapping, Leonaspis being exclusively Gondwanan post-Wenlock, whereas Exallaspis is restricted to areas north of the Rheic Ocean and ranges from basal Llandovery to Ludlow. Kettneraspis is pandemic. The condition of five epiborder spines and ten border spines on the free cheek is proposed as plesiomorphic for odontopleurids, and the border spines are shown to originate as two separate rows with alternating spines. A shift in position of the facial suture is shown to transfer the genal spine from the cranidium to the free cheek in Kettneraspis meraspides of degree 1 or 2, and the previously suggested co-occurrence of a fixigenal and librigenal spine in these stages is refuted. Earlier proposed dimorphism is rejected, and two mechanisms for reduction in the number of pygidial border spines are proposed. Kettneraspis reetae sp. nov. is described.
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Article DOI: 10.1017/S026359330000420X
Project DOI: 10.7934/P2707, http://dx.doi.org/10.7934/P2707
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 1734 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 35 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 91 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:35/0 |
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MorphoBank Project 2707
MorphoBank Project 2707
- Creation Date:
15 June 2017 - Publication Date:
30 August 2018 - Project views: 12554
- Matrix downloads: 6
Authors' Institutions
- Swedish Museum of Natural History (Naturhistoriska riksmuseet) Stockholm
- University of Alberta
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Michael Pankowski Project Administrator | 64 | 0 | 0 | 35 | 0 | 0 | 1734 (1734, 0, 0) | 0 | 0 | 0 | ||||
Thomas Hegna Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M24718' (64 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Acidaspis cincinnatiensis Last Modified in 06/15/17 | 1 | 34 | 34 | 0 | 0 | 0 | 0 | 2 |
[2] † Diacanthaspis cooperi Last Modified in 06/16/17 | 4 | 31 | 31 | 0 | 0 | 0 | 0 | 2 |
[3] † Diacanthaspis scitula Last Modified in 06/29/17 | 6 | 29 | 29 | 0 | 0 | 0 | 0 | 2 |
[4] † Exallaspis bufo Last Modified in 06/29/17 | 1 | 34 | 34 | 0 | 0 | 0 | 0 | 2 |
[5] † Exallaspis acarescola Last Modified in 06/16/17 | 5 | 30 | 30 | 0 | 0 | 0 | 0 | 2 |
[6] † Exallaspis coronata Last Modified in 06/16/17 | 15 | 20 | 20 | 0 | 0 | 0 | 0 | 2 |
[7] † Exallaspis deflexa Last Modified in 06/16/17 | 6 | 29 | 29 | 0 | 0 | 0 | 0 | 2 |
[8] † Exallaspis cf. illinoiensis Last Modified in 06/20/17 | 18 | 17 | 17 | 0 | 0 | 0 | 0 | 2 |
[9] † Exallaspis lothiana Last Modified in 06/16/17 | 7 | 28 | 28 | 0 | 0 | 0 | 0 | 2 |
[10] † Exallaspis muldensis Last Modified in 06/16/17 | 5 | 30 | 30 | 0 | 0 | 0 | 0 | 2 |
[11] † Exallaspis mutica Last Modified in 06/16/17 | 3 | 32 | 32 | 0 | 0 | 0 | 0 | 2 |
[12] † Exallaspis quadrimucronata 1 Last Modified in 06/16/17 | 0 | 35 | 35 | 0 | 0 | 0 | 0 | 2 |
[13] † Exallaspis quadrimucronata 2 Last Modified in 06/16/17 | 4 | 31 | 31 | 0 | 0 | 0 | 0 | 2 |
[14] † Exallaspis varbolensis Last Modified in 06/16/17 | 13 | 22 | 22 | 0 | 0 | 0 | 0 | 2 |
[15] † Kettneraspis belli Last Modified in 06/16/17 | 7 | 29 | 28 | 0 | 0 | 0 | 0 | 2 |
[16] † Kettneraspis beni Last Modified in 06/16/17 | 9 | 27 | 26 | 0 | 0 | 0 | 0 | 2 |
[17] † Kettneraspis besti Last Modified in 06/19/17 | 11 | 25 | 24 | 0 | 0 | 0 | 0 | 2 |
[18] † Kettneraspis bispinosa Last Modified in 06/19/17 | 10 | 25 | 25 | 0 | 0 | 0 | 0 | 2 |
[19] † Kettneraspis boltoni Last Modified in 06/19/17 | 8 | 28 | 27 | 0 | 0 | 0 | 0 | 2 |
[20] † Kettneraspis centrina Last Modified in 06/19/17 | 10 | 25 | 25 | 0 | 0 | 0 | 0 | 2 |
[21] † Kettneraspis chacaltayana Last Modified in 06/19/17 | 17 | 18 | 18 | 0 | 0 | 0 | 0 | 2 |
[22] † Kettneraspis clavata Last Modified in 06/19/17 | 4 | 31 | 31 | 0 | 0 | 0 | 0 | 2 |
[23] † Kettneraspis crenata crenata Last Modified in 06/19/17 | 1 | 34 | 34 | 0 | 0 | 0 | 0 | 2 |
[24] † Kettneraspis crenata angelini Last Modified in 06/19/17 | 1 | 34 | 34 | 0 | 0 | 0 | 0 | 2 |
[25] † Kettneraspis crenata brutoni Last Modified in 06/19/17 | 4 | 31 | 31 | 0 | 0 | 0 | 0 | 2 |
[26] † Kettneraspis dormitzeri Last Modified in 06/19/17 | 7 | 28 | 28 | 0 | 0 | 0 | 0 | 2 |
[27] † Kettneraspis 'elliptica' Last Modified in 06/19/17 | 3 | 32 | 32 | 0 | 0 | 0 | 0 | 2 |
[28] † Kettneraspis franconica Last Modified in 06/20/17 | 16 | 19 | 19 | 0 | 0 | 0 | 0 | 2 |
[29] † Kettneraspis geinitziana Last Modified in 06/20/17 | 19 | 16 | 16 | 0 | 0 | 0 | 0 | 2 |
[30] † Kettneraspis hirta Last Modified in 06/20/17 | 10 | 25 | 25 | 0 | 0 | 0 | 0 | 2 |
[31] † Kettneraspis jaanussoni Last Modified in 06/20/17 | 4 | 32 | 31 | 0 | 0 | 0 | 0 | 2 |
[32] † Kettneraspis lenzi Last Modified in 06/20/17 | 2 | 34 | 33 | 0 | 0 | 0 | 0 | 2 |
[33] † Kettneraspis leridae Last Modified in 06/20/17 | 15 | 20 | 20 | 0 | 0 | 0 | 0 | 2 |
[34] † Kettneraspis leucothea Last Modified in 06/20/17 | 8 | 27 | 27 | 0 | 0 | 0 | 0 | 2 |
[35] † Kettneraspis longstaffei Last Modified in 06/20/17 | 2 | 34 | 33 | 0 | 0 | 0 | 0 | 2 |
[36] † Kettneraspis myops Last Modified in 06/20/17 | 11 | 24 | 24 | 0 | 0 | 0 | 0 | 2 |
[37] † Kettneraspis orphana Last Modified in 06/20/17 | 15 | 20 | 20 | 0 | 0 | 0 | 0 | 2 |
[38] † Kettneraspis parkini Last Modified in 06/20/17 | 7 | 28 | 28 | 0 | 0 | 0 | 0 | 2 |
[39] † Kettneraspis pigra pigra Last Modified in 06/20/17 | 6 | 29 | 29 | 0 | 0 | 0 | 0 | 2 |
[40] † Kettneraspis pigra ornata Last Modified in 06/20/17 | 17 | 18 | 18 | 0 | 0 | 0 | 0 | 2 |
[41] † Kettneraspis rattei Last Modified in 06/20/17 | 7 | 28 | 28 | 0 | 0 | 0 | 0 | 2 |
[42] † Kettneraspis risbeyi Last Modified in 06/20/17 | 16 | 19 | 19 | 0 | 0 | 0 | 0 | 2 |
[43] † Kettneraspis tuberculata tuberculata Last Modified in 06/20/17 | 2 | 33 | 33 | 0 | 0 | 0 | 0 | 2 |
[44] † Kettneraspis tuberculata favonia Last Modified in 06/20/17 | 7 | 28 | 28 | 0 | 0 | 0 | 0 | 2 |
[45] † Kettneraspis wellingtonensis Last Modified in 06/20/17 | 6 | 29 | 29 | 0 | 0 | 0 | 0 | 2 |
[46] † Kettneraspis williamsi a Last Modified in 06/29/17 | 2 | 35 | 33 | 0 | 0 | 0 | 0 | 2 |
[47] † Kettneraspis williamsi b Last Modified in 06/29/17 | 4 | 31 | 31 | 0 | 0 | 0 | 0 | 2 |
[48] † Kettneraspis cf. williamsi Last Modified in 06/20/17 | 4 | 31 | 31 | 0 | 0 | 0 | 0 | 2 |
[49] † Kettneraspis sp. a Last Modified in 06/20/17 | 4 | 31 | 31 | 0 | 0 | 0 | 0 | 2 |
[50] † Kettneraspis sp. nov. nsw Last Modified in 06/20/17 | 4 | 31 | 31 | 0 | 0 | 0 | 0 | 2 |
[51] † Leonaspis argentea Last Modified in 06/20/17 | 17 | 18 | 18 | 0 | 0 | 0 | 0 | 2 |
[52] † Leonaspis belisarius Last Modified in 06/20/17 | 9 | 26 | 26 | 0 | 0 | 0 | 0 | 2 |
[53] † Leonaspis britannica Last Modified in 06/20/17 | 4 | 31 | 31 | 0 | 0 | 0 | 0 | 2 |
[54] † Leonaspis confluens Last Modified in 06/20/17 | 17 | 18 | 18 | 0 | 0 | 0 | 0 | 2 |
[55] † Leonaspis ezellina Last Modified in 06/20/17 | 17 | 18 | 18 | 0 | 0 | 0 | 0 | 2 |
[56] † Leonaspis glabrata Last Modified in 06/20/17 | 18 | 17 | 17 | 0 | 0 | 0 | 0 | 2 |
[57] † Leonaspis hastata Last Modified in 06/20/17 | 16 | 19 | 19 | 0 | 0 | 0 | 0 | 2 |
[58] † Leonaspis hoernesi Last Modified in 06/20/17 | 7 | 28 | 28 | 0 | 0 | 0 | 0 | 2 |
[59] † Leonaspis jenkinsi Last Modified in 06/20/17 | 7 | 28 | 28 | 0 | 0 | 0 | 0 | 2 |
[60] † Leonaspis lochkovensis Last Modified in 06/20/17 | 4 | 32 | 31 | 0 | 0 | 0 | 0 | 2 |
[61] † Leonaspis leonhardi Last Modified in 06/20/17 | 2 | 33 | 33 | 0 | 0 | 0 | 0 | 2 |
[62] † Leonaspis maura Last Modified in 06/20/17 | 13 | 22 | 22 | 0 | 0 | 0 | 0 | 2 |
[63] † Leonaspis truncata Last Modified in 06/20/17 | 15 | 20 | 20 | 0 | 0 | 0 | 0 | 2 |
[64] † Meadowtownella ascita Last Modified in 06/20/17 | 2 | 33 | 33 | 0 | 0 | 0 | 0 | 2 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 12554 | |
Project overview | 1561 | |
Bibliography | 628 | |
Matrix views | 1025 | Matrix landing page (850 views); Revision and subdivision of the polyphyletic 'Leonaspis' (Trilobita) (175 views); |
Media views | 2956 | Media search (2536 views); M585175 (420 views); |
Views for media list | 949 | |
Specimen list | 1515 | |
Taxon list | 3913 | |
Documents list | 7 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 169 | |
Project downloads | 163 | |
Matrix downloads | 6 | Revision and subdivision of the polyphyletic 'Leonaspis' (Trilobita) (6 downloads); |