Project 2707: Lars Ramsk├Âld,Brian D. E. Chatterton. 1991. Revision and subdivision of the polyphyletic 'Leonaspis' (Trilobita). Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 82:333-371.
Specimen: † Leonaspis (unvouchered)
View: ventral

Abstract

A phylogenetic analysis is performed on all species previously assigned to the odontopleurid trilobite genus Leonaspis. The PAUP analysis shows this genus, as conventionally defined, to be polyphyletic, composed of four groups of equal taxonomic rank. Leonaspis (s.s.)
is here restricted to one of these groups only, a monophyletic set of species characterised by 'four-spined' pygidia (i.e. with four spines between the major border spines) and a thorax of nine segments. Kettneraspis is recognised for the largest of these four groups, and is composed of 'two-spined' species with nine thoracic segments. Leonaspis and Kettneraspis belong to the Odontopleurinae. A third group of previous 'Leonaspis' species constitute a new genus of the Acidaspidinae. Its species are characterised by being 'four-spined' and having ten thoracic segments. For these the new genus Exallaspis is erected, with type species E. bufo. A fourth group of 'Leonaspis' species, being 'four-spined' with nine thoracic segments, belongs to Eoleonaspis, an Ordovician odontopleurine genus. Leonaspis and Exallaspis are temporally and spatially non-overlapping, Leonaspis being exclusively Gondwanan post-Wenlock, whereas Exallaspis is restricted to areas north of the Rheic Ocean and ranges from basal Llandovery to Ludlow. Kettneraspis is pandemic. The condition of five epiborder spines and ten border spines on the free cheek is proposed as plesiomorphic for odontopleurids, and the border spines are shown to originate as two separate rows with alternating spines. A shift in position of the facial suture is shown to transfer the genal spine from the cranidium to the free cheek in Kettneraspis meraspides of degree 1 or 2, and the previously suggested co-occurrence of a fixigenal and librigenal spine in these stages is refuted. Earlier proposed dimorphism is rejected, and two mechanisms for reduction in the number of pygidial border spines are proposed. Kettneraspis reetae sp. nov. is described.


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Article DOI: https://doi.org/10.1017/S026359330000420X

Project DOI: 10.7934/P2707, http://dx.doi.org/10.7934/P2707
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 65 Taxa
  • 1 Specimen
  • 35 Characters
Total size of project's media files: 12 MB

Download Project SDD File
Total scored cells: 1734
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 35
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 91
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:35/0
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MorphoBank Project 2707


    Members

    member name taxa specimens media chars character
    media
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    (scored, NPA, "-")
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    Michael Pankowski
    Project Administrator
    Last logged in 11/24/17
    640035001734
    (1734, 0, 0)
    000
    Thomas Hegna
    Full membership
    Last logged in 07/09/18
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    Taxonomic Overview for Matrix 'M24718' (64 Taxa)

    taxon unscored
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    scored
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    [1] † Acidaspis cincinnatiensis
    Last Modified in 06/15/17
    1343400002
    [2] † Diacanthaspis cooperi
    Last Modified in 06/16/17
    4313100002
    [3] † Diacanthaspis scitula
    Last Modified in 06/29/17
    6292900002
    [4] † Exallaspis bufo
    Last Modified in 06/29/17
    1343400002
    [5] † Exallaspis acarescola
    Last Modified in 06/16/17
    5303000002
    [6] † Exallaspis coronata
    Last Modified in 06/16/17
    15202000002
    [7] † Exallaspis deflexa
    Last Modified in 06/16/17
    6292900002
    [8] † Exallaspis cf. illinoiensis
    Last Modified in 06/20/17
    18171700002
    [9] † Exallaspis lothiana
    Last Modified in 06/16/17
    7282800002
    [10] † Exallaspis muldensis
    Last Modified in 06/16/17
    5303000002
    [11] † Exallaspis mutica
    Last Modified in 06/16/17
    3323200002
    [12] † Exallaspis quadrimucronata 1
    Last Modified in 06/16/17
    0353500002
    [13] † Exallaspis quadrimucronata 2
    Last Modified in 06/16/17
    4313100002
    [14] † Exallaspis varbolensis
    Last Modified in 06/16/17
    13222200002
    [15] † Kettneraspis belli
    Last Modified in 06/16/17
    7292800002
    [16] † Kettneraspis beni
    Last Modified in 06/16/17
    9272600002
    [17] † Kettneraspis besti
    Last Modified in 06/19/17
    11252400002
    [18] † Kettneraspis bispinosa
    Last Modified in 06/19/17
    10252500002
    [19] † Kettneraspis boltoni
    Last Modified in 06/19/17
    8282700002
    [20] † Kettneraspis centrina
    Last Modified in 06/19/17
    10252500002
    [21] † Kettneraspis chacaltayana
    Last Modified in 06/19/17
    17181800002
    [22] † Kettneraspis clavata
    Last Modified in 06/19/17
    4313100002
    [23] † Kettneraspis crenata crenata
    Last Modified in 06/19/17
    1343400002
    [24] † Kettneraspis crenata angelini
    Last Modified in 06/19/17
    1343400002
    [25] † Kettneraspis crenata brutoni
    Last Modified in 06/19/17
    4313100002
    [26] † Kettneraspis dormitzeri
    Last Modified in 06/19/17
    7282800002
    [27] † Kettneraspis 'elliptica'
    Last Modified in 06/19/17
    3323200002
    [28] † Kettneraspis franconica
    Last Modified in 06/20/17
    16191900002
    [29] † Kettneraspis geinitziana
    Last Modified in 06/20/17
    19161600002
    [30] † Kettneraspis hirta
    Last Modified in 06/20/17
    10252500002
    [31] † Kettneraspis jaanussoni
    Last Modified in 06/20/17
    4323100002
    [32] † Kettneraspis lenzi
    Last Modified in 06/20/17
    2343300002
    [33] † Kettneraspis leridae
    Last Modified in 06/20/17
    15202000002
    [34] † Kettneraspis leucothea
    Last Modified in 06/20/17
    8272700002
    [35] † Kettneraspis longstaffei
    Last Modified in 06/20/17
    2343300002
    [36] † Kettneraspis myops
    Last Modified in 06/20/17
    11242400002
    [37] † Kettneraspis orphana
    Last Modified in 06/20/17
    15202000002
    [38] † Kettneraspis parkini
    Last Modified in 06/20/17
    7282800002
    [39] † Kettneraspis pigra pigra
    Last Modified in 06/20/17
    6292900002
    [40] † Kettneraspis pigra ornata
    Last Modified in 06/20/17
    17181800002
    [41] † Kettneraspis rattei
    Last Modified in 06/20/17
    7282800002
    [42] † Kettneraspis risbeyi
    Last Modified in 06/20/17
    16191900002
    [43] † Kettneraspis tuberculata tuberculata
    Last Modified in 06/20/17
    2333300002
    [44] † Kettneraspis tuberculata favonia
    Last Modified in 06/20/17
    7282800002
    [45] † Kettneraspis wellingtonensis
    Last Modified in 06/20/17
    6292900002
    [46] † Kettneraspis williamsi a
    Last Modified in 06/29/17
    2353300002
    [47] † Kettneraspis williamsi b
    Last Modified in 06/29/17
    4313100002
    [48] † Kettneraspis cf. williamsi
    Last Modified in 06/20/17
    4313100002
    [49] † Kettneraspis sp. a
    Last Modified in 06/20/17
    4313100002
    [50] † Kettneraspis sp. nov. nsw
    Last Modified in 06/20/17
    4313100002
    [51] † Leonaspis argentea
    Last Modified in 06/20/17
    17181800002
    [52] † Leonaspis belisarius
    Last Modified in 06/20/17
    9262600002
    [53] † Leonaspis britannica
    Last Modified in 06/20/17
    4313100002
    [54] † Leonaspis confluens
    Last Modified in 06/20/17
    17181800002
    [55] † Leonaspis ezellina
    Last Modified in 06/20/17
    17181800002
    [56] † Leonaspis glabrata
    Last Modified in 06/20/17
    18171700002
    [57] † Leonaspis hastata
    Last Modified in 06/20/17
    16191900002
    [58] † Leonaspis hoernesi
    Last Modified in 06/20/17
    7282800002
    [59] † Leonaspis jenkinsi
    Last Modified in 06/20/17
    7282800002
    [60] † Leonaspis lochkovensis
    Last Modified in 06/20/17
    4323100002
    [61] † Leonaspis leonhardi
    Last Modified in 06/20/17
    2333300002
    [62] † Leonaspis maura
    Last Modified in 06/20/17
    13222200002
    [63] † Leonaspis truncata
    Last Modified in 06/20/17
    15202000002
    [64] † Meadowtownella ascita
    Last Modified in 06/20/17
    2333300002


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