Project 2722: J. M. Mirande. 2018. Morphology, molecules, and the phylogeny of the Characidae (Teleostei, Characiformes). Cladistics. (In Press)
Specimen: Exodon paradoxus
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Abstract

This is the most comprehensive phylogenetic analysis of the Characidae to date and the first large-scale hypothesis of the family, combining myriad morphological data with molecular information. A total of 520 morphological characters were analysed herein, of which 98 are newly defined. Among the analysed taxa, 259 species were coded by examining specimens, three fossil species were coded from the literature, one species was coded almost completely from published figures, 122 were partially coded from the literature, and 88 were analysed exclusively from molecular data. The total number of species in the analysed dataset is 473. Analyses were made by parsimony under equal and extended implied weighting with a broad range of parameters. The final hypothesis was selected using a stability criterion that chooses among the most parsimonious trees of all searches. It was found by weighting molecular characters with the average homoplasy of entire partitions (markers). The resulting hypothesis is congruent with previous molecular-based phylogenies of the family. The Characidae are monophyletic, with four main clades: the Spintherobolinae new subfamily; an expanded Stethaprioninae including the Grundulini, Gymnocharacini, Rhoadsiini and Stethaprionini; the Stevardiinae; and a clade composed of the Aphyocharacinae, Characinae, Cheirodontinae, Exodontinae and Tetragonopterinae. Also, a stem Characidae was found, as formed by the Eocene-Oligocene genera +Bryconetes and +Paleotetra as successive sister groups of extant members of the family. A subfamilial classification is proposed, but deep changes in the systematics that are beyond the scope of this study are still needed to classify the Characidae into monophyletic genera.


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Article DOI: https://doi.org/10.1111/cla.12345

Project DOI: 10.7934/P2722, http://dx.doi.org/10.7934/P2722
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 1 Document
  • 392 Taxa
  • 1 Specimen
  • 520 Characters
Total size of project's media files: 6 MB

Download Project SDD File
Total scored cells: 140392
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 520
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 1056
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:520/0
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MorphoBank Project 2722
  • Creation Date:
    03 July 2017
  • Publication Date:
    09 August 2018
  • Project views: 988


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    Juan Marcos Mirande
    Project Administrator
    Last logged in 08/09/18
    3921105200000140392
    (134262, 0, 6130)
    00000


    Taxonomic Overview for Matrix 'M24978' (385 Taxa)

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    [1] Abramites hypselonotus
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    23432425072001
    [2] Acestrocephalus sardina
    Last Modified in 01/22/18
    59454445016001
    [3] Acestrorhynchus lacustris
    Last Modified in 01/22/18
    26461454040001
    [4] Acestrorhynchus pantaneiro
    Last Modified in 01/22/18
    9476468043001
    [5] Acrobrycon ipanquianus
    Last Modified in 01/22/18
    551650609001
    [6] Agoniates anchovia
    Last Modified in 01/22/18
    39459457024001
    [7] Agoniates halecinus
    Last Modified in 01/22/18
    31474468021001
    [8] Alestes macrophthalmus
    Last Modified in 01/22/18
    179312311030001
    [9] Apareiodon affinis
    Last Modified in 01/22/18
    44408407069001
    [10] Aphyocharacidium bolivianum
    Last Modified in 01/22/18
    27487479014001
    [11] Aphyocharax anisitsi
    Last Modified in 01/22/18
    9497492019001
    [12] Aphyocharax dentatus
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    4503497019001
    [13] Aphyocharax nattereri
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    10480478032001
    [14] Aphyodite grammica
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    [15] Argopleura magdalenensis
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    7520497016001
    [16] Astyanacinus moorii
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    [17] Astyanax abramis
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    [18] Astyanax lacustris
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    [19] Astyanax eigenmanniorum
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    [20] Astyanax cf. rutilus
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    [21] Astyanax chico
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    1550149609001
    [22] Astyanax correntinus
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    1550149609001
    [23] Astyanax cremnobates
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    11512498011001
    [24] Astyanax endy
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    [25] Astyanax latens
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    [26] Astyanax lineatus
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    [27] Astyanax mexicanus
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    [28] Astyanax paris
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    [29] Astyanax pelegrini
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    [30] Astyanax puka
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    [31] Astyanax troya
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    [32] Atopomesus pachyodus
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    [33] Attonitus ephimeros
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    [34] Aulixidens eugeniae
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    [35] Axelrodia lindeae
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    [36] Bario steindachneri
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    [37] Boulengerella lateristriga
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    [38] Brachychalcinus copei
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    [39] Astyanax bransfordii
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    [40] Brittanichthys axelrodi
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    [41] Brachyalestes carolinae
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    [42] Brycon falcatus
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    [43] Brycon gouldingi
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    [44] Brycon orbignyanus
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    [45] Brycon pesu
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    [46] Brycon polylepis
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    [47] Knodus tanaothoros
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    [48] Bryconaethiops macrops
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    [49] Bryconamericus agna
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    [50] Bryconamericus uporas
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    [51] Hemibrycon caucanus
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    [52] Bryconamericus cf. iheringii
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    [53] Bryconamericus iheringii
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    [54] Bryconamericus ikaa
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    [55] Eretmobrycon emperador
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    [56] Bryconamericus exodon
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    [57] Bryconamericus mennii
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    [58] Eretmobrycon peruanus
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    [59] Bryconamericus rubropictus
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    [60] Eretmobrycon scleroparius
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    [61] Bryconamericus stramineus
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    [62] Piabina thomasi
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    [63] Bryconexodon juruenae
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    [64] Bryconops affinis
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    [65] Bryconops melanurus
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    [66] Carlana eigenmanni
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    [67] Carlastyanax aurocaudatus
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    [68] Carnegiella strigata
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    [69] Chalceus macrolepidotus
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    [70] Characidium zebra gr.
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    [71] Characidium rachovii
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    [72] Charax leticiae
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    [73] Charax stenopterus
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    [74] Cheirodon interruptus
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    [75] Coptobrycon bilineatus
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    [76] Creagrutus anary
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    [77] Creagrutus atrisignum
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    [78] Creagrutus cracentis
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    [79] Creagrutus gephyrus
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    [80] Creagrutus meridionalis
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    2049849208001
    [81] Creagrutus muelleri
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    97397396027001
    [82] Creagrutus peruanus
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    97399397026001
    [83] Creagrutus taphorni
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    4946546209001
    [84] Ctenobrycon hauxwellianus
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    [85] Diapoma alburnus
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    1251049909001
    [86] Diapoma obi
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    15511494011001
    [87] Diapoma uruguayense
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    [88] Cyanogaster noctivaga
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    73431423024001
    [89] Cynopotamus argenteus
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    [90] Cyphocharax spilotus
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    [91] Deuterodon iguape
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    [92] Deuterodon langei
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    [93] Deuterodon sp.
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    [94] Diapoma speculiferum
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    [95] Diapoma terofali
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    10501492018001
    [96] Distichodus maculatus
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    23420416081001
    [97] Ectrepopterus uruguayensis
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    29495481010001
    [98] Engraulisoma taeniatum
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    [99] Exodon paradoxus
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    [100] Galeocharax humeralis
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    [101] Galeocharax gulo
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    [102] Gnathocharax steindachneri
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    [103] Grundulus bogotensis
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    [104] Gymnocharacinus bergii
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    [105] Gymnocorymbus ternetzi
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    19499491010001
    [106] Hasemania nana
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    10486480030001
    [107] Hasemania crenuchoides
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    22491482016001
    [108] Hemibrycon dariensis
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    [109] Hemibrycon surinamensis
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    [110] Hemigrammus bleheri
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    [111] Hemigrammus erythrozonus
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    [112] Hemigrammus marginatus
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    [113] Hemigrammus ulreyi
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    [114] Hemigrammus unilineatus
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    [115] Hemiodus thayeria
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    [116] Heterocharax macrolepis
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    [117] Heterocheirodon yatai
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    [118] Hollandichthys multifasciatus
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    [119] Hoplias cf. malabaricus
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    [120] Hoplocharax goethei
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    31469458031001
    [121] Hyphessobrycon anisitsi
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    [122] Hyphessobrycon bifasciatus
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    [123] Hyphessobrycon compressus
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    [124] Hyphessobrycon elachys
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    [125] Hyphessobrycon eques
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    [126] Hyphessobrycon herbertaxelrodi
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    [127] Hyphessobrycon megalopterus
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    [128] Hyphessobrycon meridionalis
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    [129] Hyphessobrycon pulchripinnis
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    [130] Hyphessobrycon socolofi
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    [131] Hyphessobrycon wajat
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    [132] Iguanodectes geisleri
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    [133] Inpaichthys kerri
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    [134] Jupiaba mucronata
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    [135] Jupiaba scologaster
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    [136] Jupiaba acanthogaster
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    [137] Jupiaba anteroides
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    [138] Jupiaba polylepis
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    [139] Knodus breviceps
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    [140] Knodus gamma
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    [141] Knodus heteresthes
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    [142] Knodus meridae
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    [143] Bryconacidnus pectinatus
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    [144] Leporinus striatus
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    [145] Lignobrycon myersi
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    [146] Lonchogenys ilisha
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    [147] Markiana nigripinnis
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    [148] Metynnis mola
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    [149] Micralestes stormsi
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    [150] Microgenys minuta
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    [151] Microschemobrycon casiquiare
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    [152] Mimagoniates rheocharis
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    [153] Moenkhausia ceros
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    [154] Moenkhausia cotinho
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    [155] Moenkhausia bonita
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    [156] Moenkhausia dichroura
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    [157] Moenkhausia heikoi
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    [158] Moenkhausia jamesi
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    [159] Moenkhausia lata
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    [160] Moenkhausia sanctaefilomenae
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    [161] Moenkhausia xinguensis
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    [162] Myxiops aphos
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    [163] Nantis indefessus
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    [164] Nematobrycon palmeri
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    [165] Nematocharax venustus
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    [166] Odontostilbe microcephala
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    [167] Odontostilbe paraguayensis
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    [168] Holoshesthes pequira
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    [169] Odontostoechus lethostigmus
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    [170] Oligosarcus bolivianus
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    [171] Oligosarcus itau
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    [172] Oligosarcus jenynsii
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    [173] Oligosarcus longirostris
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    [174] Oligosarcus menezesi
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    [175] Oligosarcus pintoi
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    [176] Orthospinus franciscensis
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    [177] Paleotetra entrecorregos
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    [178] Paleotetra aiuruoca
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    [179] Paracheirodon axelrodi
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    [180] Paragoniates alburnus
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    [181] Parecbasis cyclolepis
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    [182] Parodon nasus
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    [183] Petitella georgiae
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    [184] Phenacogaster franciscoensis
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    [185] Phenacogaster tegatus
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    [186] Phenagoniates macrolepis
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    [187] Piabarchus analis
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    [188] Piabina argentea
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    [189] Piabucus melanostoma
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    [190] Piaractus mesopotamicus
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    [191] Poptella paraguayensis
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    [192] Prionobrama filigera
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    [193] Prionobrama paraguayensis
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    [194] Pristella maxillaris
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    [195] Probolodus heterostomus
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    [196] Prochilodus lineatus
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    [197] Prodontocharax melanotus
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    [198] Psellogrammus kennedyi
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    [199] Pseudochalceus kyburzi
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    [200] Pseudocorynopoma doriae
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    [201] Pseudocorynopoma heterandria
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    [202] Pyrrhulina australis
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    [203] Rhaphiodon vulpinus
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    [204] Rhoadsia altipinna
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    [205] Roeboexodon guyanensis
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    [206] Roeboides descalvadensis
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    [207] Roeboides microlepis
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    [208] Salminus brasiliensis
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    [209] Salminus hilarii
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    [210] Serrapinnus calliurus
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    [211] Serrapinnus notomelas
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    [212] Serrasalmus maculatus
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    [213] Serrasalmus rhombeus
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    [214] Spintherobolus ankoseion
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    [215] Spintherobolus papilliferus
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    [216] Stethaprion erythrops
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    [217] Stichonodon insignis
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    [218] Tetragonopterus argenteus
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    [219] Tetragonopterus chalceus
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    [220] Thayeria boehlkei
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    [221] Thayeria obliqua
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    [222] Thoracocharax stellatus
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    [223] Triportheus angulatus
    Last Modified in 01/22/18
    29469465026001
    [224] Triportheus nematurus
    Last Modified in 01/22/18
    13484480027001
    [225] Triportheus pantanensis
    Last Modified in 01/22/18
    18478475027001
    [226] Xenagoniates bondi
    Last Modified in 01/22/18
    54439436030001
    [227] Deuterodon longirostris
    Last Modified in 01/22/18
    35480466019001
    [228] Astyanax intermedius
    Last Modified in 01/22/18
    14500492014001
    [229] Deuterodon potaroensis
    Last Modified in 01/22/18
    12500495013001
    [230] Deuterodon rosae
    Last Modified in 01/22/18
    26479474020001
    [231] Deuterodon stigmaturus
    Last Modified in 01/22/18
    21488479020001
    [232] Diapoma guarani
    Last Modified in 01/22/18
    40480469011001
    [233] Macropsobrycon uruguayanae
    Last Modified in 01/22/18
    32472466022001
    [234] Knodus moenkhausii
    Last Modified in 01/22/18
    2750148706001
    [235] Hyphessobrycon erythrostigma
    Last Modified in 01/22/18
    3648447806001
    [236] Hemigrammus rodwayi
    Last Modified in 01/22/18
    4847146606001
    [237] Protocheirodon pi
    Last Modified in 01/22/18
    71433424025001
    [238] Odontostilbe fugitiva
    Last Modified in 01/22/18
    48462460012001
    [239] Odontostilbe pulchra
    Last Modified in 01/22/18
    27493481012001
    [240] Pseudocheirodon terrabae
    Last Modified in 01/22/18
    23490480017001
    [241] Compsura heterura
    Last Modified in 01/22/18
    23476474023001
    [242] Amazonspinther dalmata
    Last Modified in 01/22/18
    68401397055001
    [243] Oligosarcus paranensis
    Last Modified in 01/22/18
    6498490024001
    [244] Hyphessobrycon boulengeri
    Last Modified in 01/22/18
    18501490012001
    [245] Hyphessobrycon poecilioides
    Last Modified in 01/22/18
    10500493017001
    [246] Hemigrammus pulcher
    Last Modified in 01/22/18
    6507500014001
    [247] Hemigrammus aguaruna
    Last Modified in 01/22/18
    21492485014001
    [248] Bryconella pallidifrons
    Last Modified in 01/22/18
    6501491023001
    [249] Phycocharax rasbora
    Last Modified in 01/22/18
    9500485026001
    [250] Hyphessobrycon montagi
    Last Modified in 01/22/18
    11496489020001
    [251] Characidae n.sp. r
    Last Modified in 01/22/18
    14492488018001
    [252] Hyphessobrycon vanzolinii
    Last Modified in 01/22/18
    17492482021001
    [253] Hemigrammus haraldi
    Last Modified in 01/22/18
    11496494015001
    [254] Erythrocharax altipinnis
    Last Modified in 01/22/18
    22463453045001
    [255] Hyphessobrycon loweae
    Last Modified in 01/22/18
    17487481022001
    [256] Hemigrammus cf. lunatus
    Last Modified in 01/22/18
    11500492017001
    [257] Hyphessobrycon moniliger
    Last Modified in 01/22/18
    15488481024001
    [258] Bryconetes enigmaticus
    Last Modified in 01/22/18
    36015815802001
    [259] Moenkhausia pirauba
    Last Modified in 01/22/18
    14503494012001
    [260] Moenkhausia lepidura
    Last Modified in 01/22/18
    12491485023001
    [261] Moenkhausia collettii
    Last Modified in 01/22/18
    9496491020001
    [262] Hyphessobrycon santae
    Last Modified in 01/22/18
    16495489015001
    [263] Serrapinnus microdon
    Last Modified in 01/22/18
    4516504012001
    [264] Acinocheirodon melanogramma
    Last Modified in 01/22/18
    433908403001
    [265] Aphyocheirodon hemigrammus
    Last Modified in 01/22/18
    454636303001
    [266] Argopleura chocoensis
    Last Modified in 01/22/18
    462565503001
    [267] Astyanax aeneus
    Last Modified in 01/22/18
    487333003001
    [268] Attonitus bounites
    Last Modified in 01/22/18
    488313002001
    [269] Attonitus irisae
    Last Modified in 01/22/18
    488313002001
    [270] Astyanax baileyi
    Last Modified in 01/22/18
    499201902001
    [271] Astyanax caballeroi
    Last Modified in 01/22/18
    33218718404001
    [272] Eretmobrycon brevirostris
    Last Modified in 01/22/18
    478403804001
    [273] Hemibrycon cristiani
    Last Modified in 01/22/18
    474444303001
    [274] Eretmobrycon dahli
    Last Modified in 01/22/18
    466494905001
    [275] Hemibrycon loisae
    Last Modified in 01/22/18
    467515003001
    [276] Knodus orteguasae
    Last Modified in 01/22/18
    483343403001
    [277] Bryconamericus pachacuti
    Last Modified in 01/22/18
    481373603001
    [278] Bryconamericus patriciae
    Last Modified in 01/22/18
    465565203001
    [279] Hemibrycon plutarcoi
    Last Modified in 01/22/18
    470484604001
    [280] Eretmobrycon terrabensis
    Last Modified in 01/22/18
    472474503001
    [281] Ceratobranchia obtusirostris
    Last Modified in 01/22/18
    423939304001
    [282] Cheirodon australe
    Last Modified in 01/22/18
    460535208001
    [283] Cheirodon kiliani
    Last Modified in 01/22/18
    460525208001
    [284] Chrysobrycon myersi
    Last Modified in 01/22/18
    457646003001
    [285] Clupeacharax anchoveoides
    Last Modified in 01/22/18
    39910199022001
    [286] Compsura gorgonae
    Last Modified in 01/22/18
    441767207001
    [287] Corynopoma riisei
    Last Modified in 01/22/18
    452646404001
    [288] Creagrutus affinis
    Last Modified in 01/22/18
    441757504001
    [289] Creagrutus beni
    Last Modified in 01/22/18
    423969403001
    [290] Creagrutus brevipinnis
    Last Modified in 01/22/18
    441767603001
    [291] Creagrutus britskii
    Last Modified in 01/22/18
    423969403001
    [292] Creagrutus ignotus
    Last Modified in 01/22/18
    423969403001
    [293] Creagrutus occidaneus
    Last Modified in 01/22/18
    423969403001
    [294] Creagrutus seductus
    Last Modified in 01/22/18
    423949403001
    [295] Creagrutus phasma
    Last Modified in 01/22/18
    423949403001
    [296] Creagrutus maxillaris
    Last Modified in 01/22/18
    4188685017001
    [297] Creagrutus ungulus
    Last Modified in 01/22/18
    423979403001
    [298] Creagrutus pila
    Last Modified in 01/22/18
    423969403001
    [299] Creagrutus changae
    Last Modified in 01/22/18
    423969304001
    [300] Creagrutus melanzonus
    Last Modified in 01/22/18
    423979403001
    [301] Creagrutus barrigai
    Last Modified in 01/22/18
    423959403001
    [302] Diapoma dicropotamicus
    Last Modified in 01/22/18
    455636203001
    [303] Diapoma lepiclastus
    Last Modified in 01/22/18
    441807702001
    [304] Diapoma tipiaia
    Last Modified in 01/22/18
    450686802001
    [305] Eretmobrycon bayano
    Last Modified in 01/22/18
    461575603001
    [306] Gephyrocharax atracaudatus
    Last Modified in 01/22/18
    37814514002001
    [307] Gephyrocharax chocoensis
    Last Modified in 01/22/18
    424979402001
    [308] Gephyrocharax valencia
    Last Modified in 01/22/18
    4181059903001
    [309] Gilbertolus maracaiboensis
    Last Modified in 01/22/18
    457626102001
    [310] Glandulocauda melanopleura
    Last Modified in 01/22/18
    440797802001
    [311] Hemibrycon beni
    Last Modified in 01/22/18
    447727003001
    [312] Hemibrycon boquiae
    Last Modified in 01/22/18
    494252402001
    [313] Hemibrycon brevispini
    Last Modified in 01/22/18
    457636102001
    [314] Hemibrycon huambonicus
    Last Modified in 01/22/18
    445757203001
    [315] Hemibrycon helleri
    Last Modified in 01/22/18
    445757203001
    [316] Hemibrycon inambari
    Last Modified in 01/22/18
    445767203001
    [317] Hemibrycon jabonero
    Last Modified in 01/22/18
    475474302001
    [318] Hemibrycon jelskii
    Last Modified in 01/22/18
    446757202001
    [319] Hemibrycon quindos
    Last Modified in 01/22/18
    464545303001
    [320] Hemibrycon taeniurus
    Last Modified in 01/22/18
    447767102001
    [321] Hypobrycon leptorhynchus
    Last Modified in 01/22/18
    4495855016001
    [322] Hypobrycon maromba
    Last Modified in 01/22/18
    4515453016001
    [323] Knodus megalops
    Last Modified in 01/22/18
    482353503001
    [324] Knodus smithi
    Last Modified in 01/22/18
    462504909001
    [325] Kolpotocheirodon figueiredoi
    Last Modified in 01/22/18
    433898502001
    [326] Kolpotocheirodon theloura
    Last Modified in 01/22/18
    433898502001
    [327] Lepidocharax burnsi
    Last Modified in 01/22/18
    40611511103001
    [328] Leptagoniates steindachneri
    Last Modified in 01/22/18
    467454508001
    [329] Lophiobrycon weitzmani
    Last Modified in 01/22/18
    451706603001
    [330] Macropsobrycon xinguensis
    Last Modified in 01/22/18
    4308377013001
    [331] Mimagoniates inequalis
    Last Modified in 01/22/18
    423989601001
    [332] Mimagoniates microlepis
    Last Modified in 01/22/18
    36914814605001
    [333] Moenkhausia agnesae
    Last Modified in 01/22/18
    468504903001
    [334] Moenkhausia australe
    Last Modified in 01/22/18
    495232302001
    [335] Moenkhausia celibela
    Last Modified in 01/22/18
    4238481016001
    [336] Moenkhausia comma
    Last Modified in 01/22/18
    477424102001
    [337] Moenkhausia copei
    Last Modified in 01/22/18
    475434203001
    [338] Moenkhausia cosmops
    Last Modified in 01/22/18
    4268279015001
    [339] Moenkhausia costae
    Last Modified in 01/22/18
    474444105001
    [340] Moenkhausia forestii
    Last Modified in 01/22/18
    442757107001
    [341] Moenkhausia gracilima
    Last Modified in 01/22/18
    492282503001
    [342] Moenkhausia hemigrammoides
    Last Modified in 01/22/18
    4545350016001
    [343] Moenkhausia inrai
    Last Modified in 01/22/18
    435848203001
    [344] Moenkhausia intermedia
    Last Modified in 01/22/18
    485333302001
    [345] Moenkhausia nigromarginata
    Last Modified in 01/22/18
    4297675016001
    [346] Moenkhausia oligolepis
    Last Modified in 01/22/18
    471504603001
    [347] Moenkhausia phaeonota
    Last Modified in 01/22/18
    4367268016001
    [348] Moenkhausia pyrophthalma
    Last Modified in 01/22/18
    444747204001
    [349] Nanocheirodon insignis
    Last Modified in 01/22/18
    38113313207001
    [350] Odontostilbe ecuadorensis
    Last Modified in 01/22/18
    40810710408001
    [351] Odontostilbe dialeptura
    Last Modified in 01/22/18
    4191039407001
    [352] Odontostilbe mitoptera
    Last Modified in 01/22/18
    419969308001
    [353] Odontostilbe splendida
    Last Modified in 01/22/18
    40610910608001
    [354] Cheirodon stenodon
    Last Modified in 01/22/18
    449726902001
    [355] Planaltina britskii
    Last Modified in 01/22/18
    38813112903001
    [356] Pseudocheirodon arnoldi
    Last Modified in 01/22/18
    446686707001
    [357] Rachoviscus crassiceps
    Last Modified in 01/22/18
    490302901001
    [358] Rhinobrycon negrensis
    Last Modified in 01/22/18
    41710010003001
    [359] Roestes ogilviei
    Last Modified in 01/22/18
    456606004001
    [360] Saccoderma melanostigma
    Last Modified in 01/22/18
    444706808001
    [361] Stygichthys typhlops
    Last Modified in 01/22/18
    4267978016001
    [362] Tetragonopterus anostomus
    Last Modified in 01/22/18
    4189186016001
    [363] Tetragonopterus araguaiensis
    Last Modified in 01/22/18
    433837908001
    [364] Tetragonopterus carvalhoi
    Last Modified in 01/22/18
    454656402001
    [365] Tetragonopterus denticulatus
    Last Modified in 01/22/18
    443707007001
    [366] Tetragonopterus rarus
    Last Modified in 01/22/18
    445747203001
    [367] Tyttocharax tambopatensis
    Last Modified in 01/22/18
    421989801001
    [368] Xenurobrycon heterodon
    Last Modified in 01/22/18
    433868601001
    [369] Xenurobrycon pteropus
    Last Modified in 01/22/18
    455646401001
    [370] Xenurobrycon polyancistrus
    Last Modified in 01/22/18
    461615801001
    [371] Paracheirodon innesi
    Last Modified in 01/22/18
    395116109016001
    [372] Phenacogaster calverti
    Last Modified in 01/22/18
    40111611306001
    [373] Knodus cinarucoense
    Last Modified in 01/22/18
    460615703001
    [374] Gymnocorymbus bondi
    Last Modified in 01/22/18
    4376765018001
    [375] Gymnocorymbus thayeri
    Last Modified in 01/22/18
    448686408001
    [376] Microschemobrycon melanotus
    Last Modified in 01/22/18
    461525207001
    [377] Hemibrycon galvisi
    Last Modified in 01/22/18
    462575503001
    [378] Eretmobrycon miraensis
    Last Modified in 01/22/18
    467505003001
    [379] Eretmobrycon gonzalezi
    Last Modified in 01/22/18
    450676604001
    [380] Bryconamericus diaphanus
    Last Modified in 01/22/18
    466525103001
    [381] Knodus deuterodonoides
    Last Modified in 01/22/18
    471514702001
    [382] Knodus alpha
    Last Modified in 01/22/18
    480393703001
    [383] Astyanax orthodus
    Last Modified in 01/22/18
    460595703001
    [384] Astyanax nasutus
    Last Modified in 01/22/18
    457636102001
    [385] Astyanax nicaraguensis
    Last Modified in 01/22/18
    466545202001


    Project views

    type number of views Individual items viewed (where applicable)
    Total project views988
    Project overview148
    Media views134Media search (116 views); M451802 (18 views);
    Taxon list362
    Matrix views126Matrix landing page (70 views); Dataset (56 views);
    Specimen list64
    Views for media list50
    Documents list69
    Bibliography35




    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project11
    Project downloads10
    Document downloads1Dataset (1 download);