Project 2789: A. Bignon, B. G. Waisfeld, N. E. Vaccari, B. D. Chatterton. 2020. Reassessment of the Order Trinucleida (Trilobita). Journal of Systematic Palaeontology. 18 (13):1061-1077.
Specimen: † Trinucleida (unvouchered)
View: 2D image

Abstract

Trinucleoids have been treated as a superfamily of the Order Asaphida, based on: a pre-occipital glabellar tubercle, a cephalic median suture in most ancestral forms and the presence of broadly similar globular protaspid larvae in the life cycle. Recent discoveries cast doubts on the monophyly of the two last characters. This study provides a phylogenetic analysis whose main purpose is to test the affiliation of trinucleoids. The topology of the strict consensus tree obtained indicates that trinucleoids share a more recent common ancestor with members of the Order Ptychopariida than they do with any asaphid. The stratigraphically higher and most derived trinucleoids are well defined and easy to recognize. Their basal representatives can also be identified with confidence, in spite of substantial variation in the diagnostic characters of the group. These early trinucleoids are similar enough to the typical basal “ptychopariid” bauplan to suggest ancestry within that diffuse and poorly defined trilobite order. Indeed, Ptychopariida is presently regarded as being paraphyletic. We consider that raising trinucleids to ordinal status, by proposing the Order Trinucleida, will make it easier to decipher higher level relationships among trilobites. This action, sustained by improved morphologic and ontogenetic information of Cambro-Ordovician trilobites, reveals convergences of the three characters that have been used to support the inclusion of trinucleids within Asaphida. Based on this new information, we propose six distinct morphotypes of commutavi protaspis and, as a probable sensorial organ, that the cephalic axial tubercle migration might be caused by an increase in the height of the glabella and thus may not bear an important systematic signal. Moreover, the tree topology provided herein offers a preliminary overview of evolution within the order. Alsataspididae and Raphiophoridae are paraphyletic, suggesting that more investigation is needed to define them clearly. Dionidae appears to be more closely related to Raphiophoridae than to Trinucleidae, suggesting that the character state of ‘a perforated fringe’ evolved more than once.


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Article DOI: 10.1080/14772019.2020.1720324

Project DOI: 10.7934/P2789, http://dx.doi.org/10.7934/P2789
This project containsMatrices
  • 151 Media
  • 1 Matrix
  • 2 Documents
  • 91 Taxa
  • 3 Specimens
  • 75 Characters
Total size of project's media files: 21.82M

Download Project SDD File
Total scored cells: 4661
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 75
Total characters with associated media: 14
Total characters with media with labels: 0
Total character states: 185
Total character states with associated media: 130
Total character states with media with labels:0
Total unordered/ordered characters:75/0
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MorphoBank Project 2789

    Authors' Institutions

    • Universidad Nacional de Cordoba (UNC)

    • University of Alberta

    • Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET)

    • Universidad Nacional de la Rioja



    Members

    member name taxa specimens media chars character
    media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
    medialabels
    rules
    Arnaud Bignon
    Project Administrator
    9131517514404661
    (4276, 0, 385)
    000
    Brian Chatterton
    Full membership
    0000000
    (0, 0, 0)
    000
    Emilio Vaccari
    Full membership
    0000000
    (0, 0, 0)
    000
    Beatriz Waisfeld
    Full membership
    0000000
    (0, 0, 0)
    000


    Taxonomic Overview for Matrix 'M24906' (72 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] † Myinda uriconii
    Last Modified in 08/28/19
    37363602004
    [2] † Myindella crux
    Last Modified in 08/28/19
    6676702004
    [3] † Hanchungolithus richthofeni
    Last Modified in 11/29/18
    5686802004
    [4] † Ningkianolithus welleri
    Last Modified in 08/28/19
    32414102004
    [5] † Reedolithus quebecensis
    Last Modified in 12/04/18
    3707002004
    [6] † Protoincaia ancestor
    Last Modified in 12/04/18
    10565609004
    [7] † Tretaspis ceryx
    Last Modified in 12/04/18
    6606009004
    [8] † Stapeleyella inconstans
    Last Modified in 12/04/18
    11555509004
    [9] † Bancroftolithus hughesi
    Last Modified in 12/04/18
    0737302004
    [10] † Pragolithus praecedens
    Last Modified in 11/29/18
    15515109004
    [11] † Trinucleus fimbriatus
    Last Modified in 12/04/18
    6606009004
    [12] † Cryptolithus tesselatus
    Last Modified in 11/29/18
    06363012004
    [13] † Deanaspis goldfusii
    Last Modified in 11/29/18
    55858012004
    [14] † Onnia s. superba
    Last Modified in 11/29/18
    26161012004
    [15] † Marrolithus bureaui
    Last Modified in 11/29/18
    26161012004
    [16] † Dionide mareki
    Last Modified in 11/29/18
    16262012004
    [17] † Trinucleoides reussi
    Last Modified in 08/28/19
    55858012004
    [18] † Ampyxella edgelli
    Last Modified in 11/29/18
    95555011004
    [19] † Globampyx trinucleoides
    Last Modified in 08/28/19
    36161011004
    [20] † Damghanampyx ginteri
    Last Modified in 11/29/18
    65959010004
    [21] † Raphioampyx sinankylosus
    Last Modified in 12/04/18
    194545011004
    [22] † Rhombampyx glaber
    Last Modified in 12/04/18
    154949011004
    [23] † Raphiophorus parvulus
    Last Modified in 12/04/19
    26262011004
    [24] † Cnemidopyge costatus
    Last Modified in 12/08/18
    145050011004
    [25] † Lonchodomas tenuis
    Last Modified in 11/29/18
    154949011004
    [26] † Ampyx nasutus
    Last Modified in 08/28/19
    184646011004
    [27] † Mendolaspis sanjuaninus
    Last Modified in 11/29/18
    204444011004
    [28] † Nambeetella fitzroyensis
    Last Modified in 11/29/18
    135151011004
    [29] † Pseudampyxina trisegmentata
    Last Modified in 12/04/18
    184646011004
    [30] † Lehnertia nawisapa
    Last Modified in 11/29/18
    2676706004
    [31] † Pyrimetopus pyrifrons
    Last Modified in 08/29/19
    10636302004
    [32] † Orometopus elatifrons
    Last Modified in 11/29/18
    13606002004
    [33] † Araiopleura reticulata
    Last Modified in 11/29/18
    7666602004
    [34] † Hapalopleura clavata
    Last Modified in 11/25/17
    7666602004
    [35] † Seleneceme acuticaudata
    Last Modified in 12/04/18
    25414109004
    [36] † Skljarella cracens
    Last Modified in 12/04/18
    5686802004
    [37] † Joshuaspis parvus
    Last Modified in 11/29/18
    28454502004
    [38] † Doremataspis ornata
    Last Modified in 08/28/19
    14595902004
    [39] † Gibbura lepida
    Last Modified in 08/28/19
    30434302004
    [40] † Liostracina simesi
    Last Modified in 11/29/18
    10636302004
    [41] † Liostracina volens
    Last Modified in 11/29/18
    27464602004
    [42] † Liostracina tangwangzhaiensis
    Last Modified in 11/29/18
    11626202004
    [43] † Nepea narinosa
    Last Modified in 08/28/19
    17565602004
    [44] † Norwoodia boninoi
    Last Modified in 08/28/19
    6676702004
    [45] † Glaphyraspis parva
    Last Modified in 08/28/19
    14595902004
    [46] † Aphelaspis brachyphasis
    Last Modified in 11/29/18
    10636302004
    [47] † Ptychoparia striata
    Last Modified in 12/04/18
    3707002004
    [48] † Elrathia kingi
    Last Modified in 11/29/18
    5686802004
    [49] † Altikolia kurshabica
    Last Modified in 08/28/19
    12616102004
    [50] † Bolaspidella housensis
    Last Modified in 08/28/19
    0737302004
    [51] † Kochaspis liliana
    Last Modified in 08/28/19
    11626202004
    [52] † Gunnia fava
    Last Modified in 08/28/19
    10636302004
    [53] † Modocia whiteleyi
    Last Modified in 08/28/19
    4696902004
    [54] † Hardyoides minor
    Last Modified in 11/29/18
    7666602004
    [55] † Asaphus (A.) expansus
    Last Modified in 11/29/18
    5666604004
    [56] † Megistaspis (m.) limbata
    Last Modified in 11/29/18
    14585803004
    [57] † Asaphellus homfrayi
    Last Modified in 08/28/19
    7656503004
    [58] † Tsinania canens
    Last Modified in 08/28/19
    10626203004
    [59] † Liomegalaspides hupeiensis
    Last Modified in 11/29/18
    4656506004
    [60] † Taihungshania shui
    Last Modified in 12/04/18
    9606006004
    [61] † Nileus armadillo
    Last Modified in 08/28/19
    2686805004
    [62] † Remopleurides xiazhenensis
    Last Modified in 12/04/18
    12575706004
    [63] † Pterocephalia norfordi
    Last Modified in 12/04/18
    2717102004
    [64] † Maladioides coreanicus
    Last Modified in 11/29/18
    10626203004
    [65] † Ivshinaspis reticulata
    Last Modified in 11/29/18
    19535303004
    [66] † Haniwa quadrata
    Last Modified in 11/29/18
    10626203004
    [67] † Abharella sp.
    Last Modified in 11/29/18
    12606003004
    [68] † Ceratopyge forficuloides
    Last Modified in 11/29/18
    16565603004
    [69] † Proceratopyge cf. p. lata
    Last Modified in 11/29/18
    11616103004
    [70] † Parabolina acanthura
    Last Modified in 11/29/18
    4696902004
    [71] † Olenellus gilberti
    Last Modified in 08/28/19
    2707003004
    [72] † Redlichia takooensis
    Last Modified in 12/04/18
    4686803004


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    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project165
    Document downloads6All Trees (4 downloads); Consensus Strict (2 downloads);
    Project downloads138
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