Project 3168: S. P. Zack, T. A. Penkrot. 2019. Tarsals of the miniscule insectivorous mammal Batodonoides from the Eocene of San Diego County, California. Journal of Vertebrate Paleontology. 39:e1672076.
Specimen: † Batodonoides powayensis (SDSNH:76901)
View: Dorsal

Abstract

Isolated astragali and calcanei of one of the smallest known living or extinct mammals, the geolabidid lipotyphlan Batodonoides powayensis, are identified and described from early and late Uintan (middle Eocene) strata in San Diego County, California. The morphology of the tarsus of Batodonoides appears to be most consistent with arboreal or scansorial locomotion. Prior assessments of geolabidid relationships have allied the group with lipotyphlans, particularly soricids and Solenodon, but the tarsus of Batodonoides shows no particular similarity to any extant lipotyphlan family. Instead, there are unexpected tarsal similarities to the extinct Nyctitheriidae, suggesting that geolabidids may have their origins among Paleocene and Eocene nyctitheres. Results of a phylogenetic analysis are consistent with a relationship either to Solenodon or to Nyctitheriidae. When geolabidids and nyctitheres are allied, the latter group shifts from the euarchontan stem to the lipotyphlan stem, potentially reconciling two divergent views on the relationships of nyctitheres. Batodonoides provides an additional example of the impact of nondental material in challenging established views of the affinities of early Cenozoic ‘insectivores.’


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Article DOI: 10.1080/02724634.2019.1672076

Project DOI: 10.7934/P3168, http://dx.doi.org/10.7934/P3168
This project containsMatrices
  • 1 Media
  • 2 Matrices
  • 1 Document
  • 139 Taxa
  • 1 Specimen
  • 726 Characters


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Total scored cells: 32672
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 726
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 1952
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:726/0
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MorphoBank Project 3168


    Members

    member name taxa specimens media media
    notes
    chars character
    comms
       
    notes  media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
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    rules
    Shawn Zack
    Project Administrator
    Last logged in 11/07/19
    139110726000032672
    (32205, 0, 467)
    00000
    Tonya Penkrot
    Full membership
    Last logged in Never
    0000000000
    (0, 0, 0)
    00000


    Taxonomic Overview for Matrix 'M25416' (66 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] † Outgroup
    Last Modified in 11/07/19
    031131100002
    [2] † Ukhaatherium
    Last Modified in 11/07/19
    6224924900002
    [3] † Maelestes
    Last Modified in 11/07/19
    11020120100002
    [4] † Asioryctes
    Last Modified in 11/07/19
    8722422400002
    [5] † Zhelestidae
    Last Modified in 11/07/19
    13817317300002
    [6] † Zalambdalestidae
    Last Modified in 11/07/19
    1429729700002
    [7] † Cimolestes magnus
    Last Modified in 11/07/19
    230818100002
    [8] † Batodon
    Last Modified in 11/07/19
    240717100002
    [9] † Gypsonictops
    Last Modified in 11/07/19
    228838300002
    [10] † Deccanolestes
    Last Modified in 11/07/19
    21010110100002
    [11] † Afrodon bustylus
    Last Modified in 11/07/19
    18312812800002
    [12] † Fordonia
    Last Modified in 11/07/19
    231808000002
    [13] † Leptictidium
    Last Modified in 11/07/19
    16214914900002
    [14] † Prodiacodon palaeictops
    Last Modified in 11/07/19
    10820320300002
    [15] † Leptictis
    Last Modified in 11/07/19
    4426726700002
    [16] † Aphronorus
    Last Modified in 11/07/19
    19511611600002
    [17] † Coriphagus
    Last Modified in 11/07/19
    232797900002
    [18] † Palaeosinopa
    Last Modified in 11/07/19
    7423723700002
    [19] † Escavadodon
    Last Modified in 11/07/19
    16514614600002
    [20] † Miacis vulpavus
    Last Modified in 11/07/19
    2828328300002
    [21] † Viverravus
    Last Modified in 11/07/19
    13118018000002
    [22] † Protungulatum
    Last Modified in 11/07/19
    15016116100002
    [23] † Loxolophus
    Last Modified in 11/07/19
    14316816800002
    [24] † Arctocyon
    Last Modified in 11/07/19
    5325825800002
    [25] † Pseudictops
    Last Modified in 11/07/19
    16914214200002
    [26] † Gomphos
    Last Modified in 11/07/19
    8622522500002
    [27] † Rhombomylus
    Last Modified in 11/07/19
    1729429400002
    [28] † Macrocranion
    Last Modified in 11/07/19
    10520620600002
    [29] † Apheliscus
    Last Modified in 11/07/19
    15215915900002
    [30] † Haplomylus
    Last Modified in 11/07/19
    16714414400002
    [31] † Paschatherium
    Last Modified in 11/07/19
    18812312300002
    [32] Rhynchocyon
    Last Modified in 06/27/18
    730430400002
    [33] Soricidae
    Last Modified in 05/09/18
    1130030000002
    [34] Echinosorex
    Last Modified in 05/09/18
    131031000002
    [35] Erinaceus
    Last Modified in 06/27/18
    230930900002
    [36] † Onychonycteris
    Last Modified in 11/07/19
    214979700002
    [37] † Icaronycteris
    Last Modified in 11/07/19
    19311811800002
    [38] Rhinolophus
    Last Modified in 05/09/18
    2728428400002
    [39] † Plagioctenodon spp.
    Last Modified in 11/07/19
    13018418100002
    [40] † Cryptotopos
    Last Modified in 11/07/19
    12718418400002
    [41] Tupaia
    Last Modified in 05/09/18
    031131100002
    [42] Ptilocercus
    Last Modified in 06/27/18
    031131100002
    [43] † Mixodectes
    Last Modified in 11/07/19
    16914214200002
    [44] † Eudaemonema
    Last Modified in 11/07/19
    238737300002
    [45] † Elpidophorus
    Last Modified in 11/07/19
    229828200002
    [46] † Plagiomene
    Last Modified in 11/07/19
    20310810800002
    [47] Cynocephalus
    Last Modified in 05/09/18
    131031000002
    [48] † Purgatorius
    Last Modified in 11/07/19
    20211010900002
    [49] † Micromomyidae
    Last Modified in 11/07/19
    13517617600002
    [50] † Ignacius
    Last Modified in 11/07/19
    6824324300002
    [51] † Plesiadapis
    Last Modified in 11/07/19
    2428728700002
    [52] † Cantius notharctus
    Last Modified in 11/07/19
    2029129100002
    [53] † Butselia biveri
    Last Modified in 11/07/19
    19811411300002
    [54] † Proterixoides davisi
    Last Modified in 11/07/19
    19312011800002
    [55] † Crypholestes vaughni
    Last Modified in 11/07/19
    19711811400002
    [56] † Sespedectes spp.
    Last Modified in 11/07/19
    19211911900002
    [57] Solenodon
    Last Modified in 06/27/18
    1230029900002
    [58] Talpidae
    Last Modified in 05/09/18
    4727126400002
    [59] Zionodon
    Last Modified in 05/09/18
    231808000002
    [60] † Litocherus spp.
    Last Modified in 11/07/19
    244676700002
    [61] † Scenopagus spp.
    Last Modified in 11/07/19
    243686800002
    [62] † Patriolestes novaceki
    Last Modified in 11/07/19
    236767500002
    [63] † Eotalpa spp.
    Last Modified in 11/07/19
    19911211200002
    [64] † Centetodon spp.
    Last Modified in 11/07/19
    16914414200002
    [65] † Batodonoides spp. uintan
    Last Modified in 11/07/19
    221919000002
    [66] † Asionyctia guoi
    Last Modified in 11/07/19
    247646400002


    Taxonomic Overview for Matrix 'M25527' (90 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] † Nanolestes
    Last Modified in 11/07/19
    30411111100002
    [2] † Peramus
    Last Modified in 11/07/19
    29212712300002
    [3] † Vincelestes
    Last Modified in 11/07/19
    4237437300002
    [4] † Kielantherium
    Last Modified in 11/07/19
    340767500002
    [5] † Deltatheridium
    Last Modified in 11/07/19
    16525225000002
    [6] † Mayulestes
    Last Modified in 11/07/19
    8732832800002
    [7] † Pucadelphys
    Last Modified in 11/07/19
    5735835800002
    [8] † Acristatherium
    Last Modified in 11/07/19
    23718017800002
    [9] † Eomaia
    Last Modified in 11/07/19
    23817717700002
    [10] † Prokennalestes
    Last Modified in 11/07/19
    22319419200002
    [11] † Murtoilestes
    Last Modified in 11/07/19
    355606000002
    [12] † Bobolestes
    Last Modified in 11/07/19
    30011611500002
    [13] † Montanalestes
    Last Modified in 11/07/19
    348696700002
    [14] † Sheikhdzheilia
    Last Modified in 11/07/19
    332838300002
    [15] † Gallolestes
    Last Modified in 11/07/19
    375414000002
    [16] † Parazhelestes
    Last Modified in 11/07/19
    21620519900002
    [17] † Aspanlestes
    Last Modified in 11/07/19
    22419619100002
    [18] † Zhelestes
    Last Modified in 11/07/19
    21220720300002
    [19] † Paranyctoides
    Last Modified in 11/07/19
    29911911600002
    [20] † Eozhelestes
    Last Modified in 11/07/19
    362535300002
    [21] † Maelestes
    Last Modified in 11/07/19
    10332031200002
    [22] † Batodon
    Last Modified in 11/07/19
    31210510300002
    [23] † Bulaklestes
    Last Modified in 11/07/19
    31510010000002
    [24] † Daulestes
    Last Modified in 11/07/19
    31410310100002
    [25] † Uchkudukodon
    Last Modified in 11/07/19
    17125124400002
    [26] † Kennalestes
    Last Modified in 11/07/19
    12029629500002
    [27] † Asioryctes
    Last Modified in 11/07/19
    8233533300002
    [28] † Ukhaatherium
    Last Modified in 11/07/19
    6635134900002
    [29] † Deccanolestes hislopi
    Last Modified in 11/07/19
    26915514600002
    [30] † Afrodon germanicus
    Last Modified in 11/07/19
    31410410100002
    [31] † Kulbeckia
    Last Modified in 11/07/19
    22020519500002
    [32] † Zhangolestes
    Last Modified in 11/07/19
    364525100002
    [33] † Zalambdalestes
    Last Modified in 11/07/19
    4937836600002
    [34] † Barunlestes
    Last Modified in 11/07/19
    13428328100002
    [35] † Gypsonictops
    Last Modified in 11/07/19
    29013112500002
    [36] † Leptictis
    Last Modified in 11/07/19
    2139539400002
    [37] † Purgatorius
    Last Modified in 11/07/19
    28513313000002
    [38] † Protungulatum
    Last Modified in 11/07/19
    20421821100002
    [39] † Oxyprimus
    Last Modified in 11/07/19
    319989600002
    [40] † Vulpavus
    Last Modified in 11/07/19
    8634332900002
    [41] † Miacis
    Last Modified in 11/07/19
    13229128300002
    [42] † Gujaratia
    Last Modified in 11/07/19
    9232632300002
    [43] † Hyopsodus
    Last Modified in 11/07/19
    4338237200002
    [44] † Meniscotherium
    Last Modified in 11/07/19
    4038037500002
    [45] † Phenacodus
    Last Modified in 11/07/19
    5936635600002
    [46] Ptilocercus
    Last Modified in 06/27/18
    1340440200002
    [47] † Plesiadapis
    Last Modified in 11/07/19
    29386369017002
    [48] † Notharctus
    Last Modified in 11/07/19
    4138237400002
    [49] † Adapis
    Last Modified in 11/07/19
    6635834900002
    [50] † Tribosphenomys
    Last Modified in 11/07/19
    26614914900002
    [51] † Paramys
    Last Modified in 11/07/19
    6835234700002
    [52] † Rhombomylus
    Last Modified in 11/07/19
    5936335600002
    [53] † Gomphos
    Last Modified in 11/07/19
    13927827600002
    [54] † Mimotona
    Last Modified in 11/07/19
    29212712300002
    [55] Blarina
    Last Modified in 11/07/19
    6735034800002
    [56] Erinaceus
    Last Modified in 06/27/18
    2340139200002
    [57] Solenodon
    Last Modified in 06/27/18
    3638037900002
    [58] † Eoryctes
    Last Modified in 11/07/19
    21320520200002
    [59] Potamogale
    Last Modified in 06/27/18
    3338238200002
    [60] Orycteropus
    Last Modified in 06/27/18
    13828427700002
    [61] Rhynchocyon
    Last Modified in 06/27/18
    3937837600002
    [62] Procavia
    Last Modified in 06/27/18
    3039238500002
    [63] † Moeritherium
    Last Modified in 11/07/19
    14826926700002
    [64] Chaetophractus
    Last Modified in 06/27/18
    13428128100002
    [65] Bradypus
    Last Modified in 06/27/18
    13428428100002
    [66] Tamandua
    Last Modified in 06/27/18
    15925825600002
    [67] † Dilambdogale
    Last Modified in 11/07/19
    31210710300002
    [68] † Widanelfarasia
    Last Modified in 11/07/19
    29312312200002
    [69] † Todralestes
    Last Modified in 11/07/19
    28712712305002
    [70] Microgale
    Last Modified in 06/27/18
    12383379024002
    [71] † Foxomomys fremdi
    Last Modified in 11/07/19
    29411711407002
    [72] † Dryomomys szalayi
    Last Modified in 11/07/19
    149255255011002
    [73] † Tinimomys graybulliensis
    Last Modified in 11/07/19
    164244241010002
    [74] † Carpolestes simpsoni
    Last Modified in 11/07/19
    66328326023002
    [75] † Ignacius
    Last Modified in 11/07/19
    62332332021002
    [76] Cynocephalus volans
    Last Modified in 06/27/18
    5388385025002
    [77] † Plagioctenodon rosei
    Last Modified in 11/07/19
    126285279010002
    [78] † Leptacodon tener
    Last Modified in 11/07/19
    31010510302002
    [79] † Plagioctenodon thewisseni
    Last Modified in 11/07/19
    21020119906002
    [80] † Adunator minutus
    Last Modified in 11/07/19
    30210910904002
    [81] † Macrocranion junnei
    Last Modified in 11/07/19
    28212812805002
    [82] † Macrocranion nitens
    Last Modified in 11/07/19
    24216816706002
    [83] † Zhalmouzia bazhanovi
    Last Modified in 11/07/19
    370454500002
    [84] † Cimolestes
    Last Modified in 11/07/19
    26515314505002
    [85] † Puercolestes simpsoni
    Last Modified in 11/07/19
    28613312603002
    [86] † Chacopterygus minimus
    Last Modified in 11/07/19
    366454405002
    [87] † Betonnia tsosia
    Last Modified in 11/07/19
    345656505002
    [88] † Procerberus
    Last Modified in 11/07/19
    27813513205002
    [89] † Batodonoides powayensis
    Last Modified in 06/27/18
    551011010259002
    [90] † Centetodon spp.
    Last Modified in 11/07/19
    18522122109002


    Project views

    type number of views Individual items viewed (where applicable)
    Total project views138
    Project overview39
    Matrix views22Modified version of Hooker (2014) matrix *NO STEPMATRICES* (6 views); Modified version of Manz et al. (2015) matrix (6 views); Matrix landing page (10 views);
    Media views20Media search (17 views); M508411 (3 views);
    Views for media list4
    Specimen list7
    Taxon list37
    Documents list7
    Bibliography2




    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project3
    Matrix downloads2Modified version of Hooker (2014) matrix *NO STEPMATRICES* (1 download); Modified version of Manz et al. (2015) matrix (1 download);
    Project downloads1