Project 3172: S. P. Zack. 2018. A skeleton of a Uintan machaeroidine ‘creodont’ and the phylogeny of carnivorous eutherian mammals. Journal of Systematic Palaeontology:1-37.
Specimen: † CM 2386 (CM:2386)
View: Dorsal

Abstract

An edentulous partial skeleton of a carnivorous mammal from the Uinta Formation (middle Eocene) of Utah is referred to the rare and enigmatic sabre-tooth clade Machaeroidinae primarily on the basis of alveolar patterns and cranial morphology. The newly recognized skeleton includes portions of both girdles, all long bones, and the first known tarsal and phalangeal material of a machaeroidine. The specimen permits a preliminary reconstruction of the locomotor habits of machaeroidines, which appear to have been adapted to scansorial or arboreal rather than terrestrial locomotion. The new material also prompts a review of machaeroidine affinities, which have been unresolved for over a century, with consensus opinion favouring a link to either oxyaenid or limnocyonine hyaenodontid ?creodonts?. Postcranial evidence favours a link to oxyaenids, as machaeroidines share numerous features with oxyaenids that are lacking in hyaenodontids. To test this relationship machaeroidines were included in a phylogenetic analysis broadly sampling early carnivorous eutherians, including members of both ?Creodonta? and Carnivoramorpha. Results place Machaeroidinae within Oxyaenidae, but fail to support either ?creodont? or carnivoramorphan monophyly. Instead, Oxyaenidae is linked with Carnivoraformes, while Viverravidae is basally positioned among carnivorous eutherians. Reconsideration of the character evidence cited in support of Carnivoramorpha indicates that many features are ambiguous in the context of a broad sample of ?creodonts? and early carnivoramorphans. Hyaenodontid monophyly is also not recovered but this likely reflects the influence of one morphologically divergent genus, Arfia.


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Article DOI: 10.1080/14772019.2018.1466374

Project DOI: 10.7934/P3172, http://dx.doi.org/10.7934/P3172
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 43 Taxa
  • 1 Specimen
  • 227 Characters
Total size of project's media files: 1 MB

Download Project SDD File
Total scored cells: 4952
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 227
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 534
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:225/2
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MorphoBank Project 3172

    Authors' Institutions

    • University of Arizona



    Members

    member name taxa specimens media media
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    Shawn Zack
    Project Administrator
    Last logged in 05/18/18
    4311022702004952
    (4952, 0, 0)
    00000


    Taxonomic Overview for Matrix 'M25422' (43 Taxa)

    taxon unscored
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    [1] OG dummy
    Last Modified in 05/18/18
    2234400001
    [2] † Protungulatum donnae
    Last Modified in 05/18/18
    134939300001
    [3] † Mithrandir gillianus
    Last Modified in 05/18/18
    9113713600001
    [4] † Altacreodus magnus
    Last Modified in 05/18/18
    180474700001
    [5] † Arfia spp.
    Last Modified in 05/18/18
    8814013900001
    [6] † Prolimnocyon spp.
    Last Modified in 05/18/18
    5717217000001
    [7] † Thinocyon spp.
    Last Modified in 05/18/18
    3019719700001
    [8] † Limnocyon spp.
    Last Modified in 05/18/18
    3219719500001
    [9] † Galecyon spp.
    Last Modified in 05/18/18
    7415315300001
    [10] † Prototomus martis
    Last Modified in 05/18/18
    11711011000001
    [11] † Pyrocyon strenuus
    Last Modified in 05/18/18
    7215515500001
    [12] † Prototomus phobos
    Last Modified in 05/18/18
    10412312300001
    [13] † Tritemnodon agilis
    Last Modified in 05/18/18
    2220720500001
    [14] † Sinopa spp.
    Last Modified in 05/18/18
    4019118700001
    [15] † Tytthaena parrisi
    Last Modified in 05/18/18
    185424200001
    [16] † Palaeonictis occidentalis
    Last Modified in 05/18/18
    9113913600001
    [17] † Dipsalidictis krausei
    Last Modified in 05/18/18
    7215515500001
    [18] † Oxyaena spp.
    Last Modified in 05/18/18
    5817116900001
    [19] † Machaeroides spp.
    Last Modified in 05/18/18
    8814113900001
    [20] † CM 2386
    Last Modified in 05/18/18
    142858500001
    [21] † Ictidopappus mustelinus
    Last Modified in 05/18/18
    186414100001
    [22] † Protictis haydenianus
    Last Modified in 05/18/18
    173555400001
    [23] † Bryanictis microlestes
    Last Modified in 05/18/18
    176545100001
    [24] † Simpsonictis spp.
    Last Modified in 05/18/18
    179564800001
    [25] † Viverravus schaffi
    Last Modified in 05/18/18
    129989800001
    [26] † Viverravus minutus
    Last Modified in 05/18/18
    144838300001
    [27] † Viverravus gracilis
    Last Modified in 05/18/18
    8913813800001
    [28] † Viverravus acutus
    Last Modified in 05/18/18
    5717017000001
    [29] † Didymictis spp.
    Last Modified in 05/18/18
    2120720600001
    [30] † Uintacyon gingerichi
    Last Modified in 05/18/18
    181464600001
    [31] † Uintacyon massetericus
    Last Modified in 05/18/18
    9013913700001
    [32] † Vassacyon promicrodon
    Last Modified in 05/18/18
    10013212700001
    [33] † Gracilocyon spp.
    Last Modified in 05/18/18
    186414100001
    [34] † Dormaalocyon latouri
    Last Modified in 05/18/18
    145838200001
    [35] † Vulpavus australis
    Last Modified in 05/18/18
    7116215600001
    [36] † Vulpavus ovatus
    Last Modified in 05/18/18
    132959500001
    [37] † Vulpavus profectus
    Last Modified in 05/18/18
    4618318100001
    [38] † Vulpavus palustris
    Last Modified in 05/18/18
    188403900001
    [39] † Oodectes jepseni
    Last Modified in 05/18/18
    176515100001
    [40] † Oodectes herpestoides
    Last Modified in 05/18/18
    7315415400001
    [41] † Miacis exiguus
    Last Modified in 05/18/18
    173575400001
    [42] † Miacis petilus
    Last Modified in 05/18/18
    11611311100001
    [43] † Miacis parvivorus
    Last Modified in 05/18/18
    137959000001


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    Project overview363
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    Media views486Media search (418 views); M565503 (68 views);
    Taxon list1495
    Specimen list213
    Views for media list153
    Bibliography99




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    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project43
    Matrix downloads7Carnivorous eutherian phylogeny (7 downloads);
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