Project 3384: A. D. Marsh, W. G. Parker, M. C. Langer, S. J. Nesbitt. 2019. Redescription of the holotype specimen of Chindesaurus bryansmalli Long and Murry, 1995 (Dinosauria, Theropoda), from Petrified Forest National Park, Arizona. Journal of Vertebrate Paleontology. 39:e1645682.
Abstract
Chindesaurus bryansmalli is an early dinosaur of uncertain affinities from the Late Triassic Chinle Formation at Petrified Forest National Park, Arizona. Since its first description in 1995, the taxon has been considered a plateosaurid, a non-eusaurischian saurischian, a herrerasaurid, and/or a non-neotheropod member of Theropoda. Chindesaurus bryansmalli is usually scored for about 25% of the characters in a given phylogenetic analysis and many characters have been scored second-hand from misidentified elements. Here we provide a redescription of the holotype specimen of Chindesaurus bryansmalli, correct misidentifications, introduce previously unknown elements, and discuss novel morphological character observations. Chindesaurus bryansmalli is supported as the sister taxon to the non-neotheropod theropod Tawa hallae from the Chinle Formation at Ghost Ranch, New Mexico. The same two most parsimonious trees, with increasing node support, result from iteratively removing the three most incomplete taxa in the employed dataset, suggesting that the relationships of stem-averostran theropods is not highly affected by the inclusion of fragmentary specimens. The Chindesaurus + Tawa clade recovered here may represent a potentially diverse group of early theropods prior to the end-Triassic mass extinction.Read the article »
Article DOI: 10.1080/02724634.2019.1645682
Project DOI: 10.7934/P3384, http://dx.doi.org/10.7934/P3384
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 9643 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 352 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 789 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:352/0 |
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MorphoBank Project 3384
MorphoBank Project 3384
- Creation Date:
08 February 2019 - Publication Date:
04 September 2019 - Media downloads: 2
- Matrix downloads: 11
Authors' Institutions
- Museum of Northern Arizona, Flagstaff
- Petrified Forest National Park, Navajo County
- University of Texas at Austin
- Virginia Tech (Virginia Polytechnic Institute and State University)
- Universidade de Sao Paulo (USP, University of Sao Paulo)
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Adam Marsh Project Administrator | 1 | 2 | 1 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
MorphoBank Curator Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
MorphoBank Curator Full membership | 44 | 0 | 0 | 352 | 0 | 0 | 9643 (9643, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M26561' (45 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Erythrosuchus africanus Taxon name last Modified on 04/01/20 | 43 | 309 | 309 | 0 | 0 | 0 | 0 | 3 |
[2] † Euparkeria capensis Taxon name last Modified on 04/01/20 | 19 | 334 | 333 | 0 | 0 | 0 | 0 | 3 |
[3] † Revueltosaurus callenderi Taxon name last Modified on 04/01/20 | 48 | 304 | 304 | 0 | 0 | 0 | 0 | 3 |
[4] † Aetosaurus ferratus Taxon name last Modified on 04/01/20 | 120 | 233 | 232 | 0 | 0 | 0 | 0 | 3 |
[5] † Arizonasaurus babbitti Taxon name last Modified on 04/01/20 | 155 | 197 | 197 | 0 | 0 | 0 | 0 | 3 |
[6] † Effigia okeeffeae Taxon name last Modified on 04/01/20 | 88 | 264 | 264 | 0 | 0 | 0 | 0 | 3 |
[7] † Batrachotomus kuperferzellensis Taxon name last Modified on 04/01/20 | 57 | 295 | 295 | 0 | 0 | 0 | 0 | 3 |
[8] † Postosuchus kirkpatricki Taxon name last Modified on 04/01/20 | 62 | 290 | 290 | 0 | 0 | 0 | 0 | 3 |
[9] † Dromicosuchus grallator Taxon name last Modified on 04/01/20 | 122 | 230 | 230 | 0 | 0 | 0 | 0 | 3 |
[10] † Eudimorphodon ranzii Taxon name last Modified on 04/01/20 | 177 | 175 | 175 | 0 | 0 | 0 | 0 | 3 |
[11] † Dimorphodon macronyx Taxon name last Modified on 04/01/20 | 156 | 196 | 196 | 0 | 0 | 0 | 0 | 3 |
[12] † Lagerpeton chanarensis Taxon name last Modified on 04/01/20 | 213 | 139 | 139 | 0 | 0 | 0 | 0 | 3 |
[13] † Dromomeron romeri Taxon name last Modified on 04/01/20 | 281 | 71 | 71 | 0 | 0 | 0 | 0 | 3 |
[14] † Dromomeron gregorii Taxon name last Modified on 04/01/20 | 304 | 48 | 48 | 0 | 0 | 0 | 0 | 3 |
[15] † Marasuchus lilloensis Taxon name last Modified on 04/01/20 | 157 | 196 | 195 | 0 | 0 | 0 | 0 | 3 |
[16] † Eucoelophysis baldwini Taxon name last Modified on 04/01/20 | 323 | 29 | 29 | 0 | 0 | 0 | 0 | 3 |
[17] † Sacisaurus agudoensis Taxon name last Modified on 04/01/20 | 254 | 100 | 98 | 0 | 0 | 0 | 0 | 3 |
[18] † Silesaurus opolensis Taxon name last Modified on 04/01/20 | 59 | 294 | 293 | 0 | 0 | 0 | 0 | 3 |
[19] † Eocursor parvus Taxon name last Modified on 04/01/20 | 218 | 134 | 134 | 0 | 0 | 0 | 0 | 3 |
[20] † Pisanosaurus mertii Taxon name last Modified on 04/01/20 | 257 | 95 | 95 | 0 | 0 | 0 | 0 | 3 |
[21] † Heterodontosaurus tucki Taxon name last Modified on 04/01/20 | 62 | 292 | 290 | 0 | 0 | 0 | 0 | 3 |
[22] † Lesothosaurus dianosticus Taxon name last Modified on 04/01/20 | 72 | 280 | 280 | 0 | 0 | 0 | 0 | 3 |
[23] † Scutellosaurus lawleri Taxon name last Modified on 04/01/20 | 178 | 175 | 174 | 0 | 0 | 0 | 0 | 3 |
[24] † Saturnalia tupiniquim Taxon name last Modified on 04/01/20 | 156 | 200 | 196 | 0 | 0 | 0 | 0 | 3 |
[25] † Plateosaurus engelhardti Taxon name last Modified on 04/01/20 | 18 | 335 | 334 | 0 | 0 | 0 | 0 | 3 |
[26] † Efraasia minor Taxon name last Modified on 04/01/20 | 64 | 288 | 288 | 0 | 0 | 0 | 0 | 3 |
[27] † Herrerasaurus ischigualastensis Taxon name last Modified on 04/01/20 | 39 | 314 | 313 | 0 | 0 | 0 | 0 | 3 |
[28] † Staurikosaurus pricei Taxon name last Modified on 04/01/20 | 222 | 130 | 130 | 0 | 0 | 0 | 0 | 3 |
[29] † Chindesaurus bryansmalli Taxon name last Modified on 04/01/20 | 264 | 89 | 88 | 0 | 0 | 0 | 0 | 3 |
[30] † Eoraptor lunensis Taxon name last Modified on 04/01/20 | 95 | 258 | 257 | 0 | 0 | 0 | 0 | 3 |
[31] † Tawa hallae Taxon name last Modified on 04/01/20 | 47 | 305 | 305 | 0 | 0 | 0 | 0 | 3 |
[32] † Coelophysis bauri Taxon name last Modified on 04/01/20 | 27 | 326 | 325 | 0 | 0 | 0 | 0 | 3 |
[33] † Syntarsus kayentakatae Taxon name last Modified on 04/01/20 | 96 | 256 | 256 | 0 | 0 | 0 | 0 | 3 |
[34] † Liliensternus liliensterni Taxon name last Modified on 04/01/20 | 148 | 206 | 204 | 0 | 0 | 0 | 0 | 3 |
[35] † Zupaysaurus rougieri Taxon name last Modified on 04/01/20 | 240 | 112 | 112 | 0 | 0 | 0 | 0 | 3 |
[36] † Dilophosaurus wetherelli Taxon name last Modified on 04/01/20 | 49 | 303 | 303 | 0 | 0 | 0 | 0 | 3 |
[37] † Cryolophosaurus ellioti Taxon name last Modified on 04/01/20 | 215 | 137 | 137 | 0 | 0 | 0 | 0 | 3 |
[38] † Ceratosaurus nasicornis Taxon name last Modified on 04/01/20 | 74 | 278 | 278 | 0 | 0 | 0 | 0 | 3 |
[39] † Piatnitzkysaurus floresi Taxon name last Modified on 04/01/20 | 177 | 175 | 175 | 0 | 0 | 0 | 0 | 3 |
[40] † Allosaurus fragilis Taxon name last Modified on 04/01/20 | 21 | 331 | 331 | 0 | 0 | 0 | 0 | 3 |
[41] † Velociraptor mongoliensis Taxon name last Modified on 04/01/20 | 63 | 289 | 289 | 0 | 0 | 0 | 0 | 3 |
[42] † Camposaurus arizonensis Taxon name last Modified on 04/01/20 | 323 | 29 | 29 | 0 | 0 | 0 | 0 | 3 |
[43] † Coelophysis rhodesiensis Taxon name last Modified on 04/01/20 | 25 | 327 | 327 | 0 | 0 | 0 | 0 | 3 |
[44] † Lepidus praecisio holotpye Taxon name last Modified on 04/01/20 | 302 | 51 | 50 | 0 | 0 | 0 | 0 | 3 |
[45] † Eodromaeus murphi Taxon name last Modified on 04/01/20 | 134 | 224 | 218 | 0 | 0 | 0 | 0 | 3 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 223 | |
Document downloads | 56 | Supplementary Data 2 (9 downloads); Marsh et al Chindesaurus matrix (5 downloads); PEFO 10395 proximal left femur (5 downloads); PEFO 10395 right femur (8 downloads); PEFO 10395 cervical centrum (2 downloads); PEFO 10395 anterior caudal vertebra (5 downloads); Morphology matrix in Excel format (1 download); PEFO 10395 astragalus (4 downloads); PEFO 10395 caudal vertebra A (1 download); PEFO 10395 caudal vertebra B (1 download); PEFO 10395 dorsal centrum (1 download); PEFO 10395 proximal right tibia (4 downloads); PEFO 10395 right tibia distal (4 downloads); PEFO 10395 sacral vertebra 1 (1 download); PEFO 10395 sacral vertebra 2 (2 downloads); MPTs (3 downloads); |
Project downloads | 154 | |
Media downloads | 2 | M657130 (2 downloads); |
Matrix downloads | 11 | Matrix from the publication of Marsh et al. 2019 (prepared by MorphoBank curators from original submission) (11 downloads); |