Project 3390: J.C.A. Marsola, J. S. Bittencourt, R. J. Butler, Á. A. S. Da Rosa, J. M. Sayão, M. C. Langer. 2019. A new dinosaur with theropod affinities from the Late Triassic Santa Maria Formation, South Brazil. Journal of Vertebrate Paleontology. 38 (5):e1531878.
Specimen: † Nhandumirim waldsangae (unvouchered)
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The Late Triassic (Carnian) upper Santa Maria Formation of south Brazil has yielded some of the oldest unequivocal records of dinosaurs. Here, we describe a new saurischian dinosaur from this formation, Nhandumirim waldsangae, gen. et sp. nov., based on a semiarticulated skeleton, including trunk, sacral, and caudal vertebrae, one chevron, right ilium, femur, partial tibia, fibula, and metatarsals II and IV, as well as ungual and non-ungual phalanges. The new taxon differs from all other Carnian dinosauromorphs through a unique combination of characters, some of which are autapomorphic: caudal centra with sharp longitudinal ventral keels; brevis fossa extending for less than three-quarters of the ventral surface of the postacetabular ala of the ilium; dorsolateral trochanter ending well distal to the level of the femoral head; distal part of the tibia with a mediolaterally extending tuberosity on its cranial surface and a tabular caudolateral flange; conspicuous, craniomedially oriented semicircular articular facet on the distal fibula; and a straight metatarsal IV. This clearly distinguishes Nhandumirim waldsangae from both Saturnalia tupiniquim and Staurikosaurus pricei, which were collected nearby and at a similar stratigraphic level. Despite not being fully grown, the differences between Nhandumirim waldsangae and those saurischians cannot be attributed to ontogeny. The phylogenetic position of Nhandumirim waldsangae suggests that it represents one of the earliest members of Theropoda. Nhandumirim waldsangae shows that some typical theropod characters were already present early in dinosaur evolution, and it represents possibly the oldest record of the group known in Brazil.

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Article DOI: 10.1080/02724634.2018.1531878

Project DOI: 10.7934/P3390,
This project contains
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MorphoBank Project 3390
  • Creation Date:
    14 February 2019
  • Publication Date:
    13 June 2019
  • Project views: 5853

    Authors' Institutions

    • University of Birmingham

    • Universidade Federal de Minas Gerais

    • Universidade de Sao Paulo (USP, University of Sao Paulo)

    • Universidade Federal de Santa Maria (UFSM)

    • Universidade Federal de Pernambuco


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    MorphoBank Curator
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    Julio Marsola
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