Project 3422: G. D. de Lavigerie, M. Bosselaers, S. Goolaerts, T. Park, O. Lambert, F. G. Marx. 2020. New Pliocene right whale from Belgium informs balaenid phylogeny and function. Journal of Systematic Palaeontology. -:1-26.

Specimen: † Antwerpibalaena liberatlas (IRSNB 12345/IRSNB 12345)
View: Articulated left ear bones of the holotype of Antwerpibalaena liberatlas (IRSNB M2325). Photographs in A, anteroventral and B, ventral views.
View: Articulated left ear bones of the holotype of Antwerpibalaena liberatlas (IRSNB M2325). Photographs in A, anteroventral and B, ventral views.
Abstract
Right whales (Balaenidae) are the most distinctive family of extant baleen whales, thanks to their highly arched rostrum, tall lips and robust body shape. They are also the oldest, originating as much as 20 million years ago (Ma). Nevertheless, their fossil record is patchy and frequently understudied, obscuring their evolution. Here, we describe a new stem balaenid, Antwerpibalaena liberatlas, from northern Belgium, adding to the rich but historically problematic baleen whale assemblage of the Pliocene North Sea. Within right whales, Antwerpibalaena forms a clade with two previously described extinct genera, Balaenella and Balaenula. The holotype preserves much of the postcranial skeleton, and informs the emergence of typical balaenid traits like fused neck vertebrae and paddle-shaped flippers. Its size is intermediate between that of extant right whales and most of their extinct forebears revealing a more complex pattern of balaenid size evolution than previously thought.Read the article »
Article DOI: 10.1080/14772019.2020.1746422
Project DOI: 10.7934/P3422, http://dx.doi.org/10.7934/P3422
This project contains | Matrices |
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Download Project SDD File ![]() | Total scored cells: 18151 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 278 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 651 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:240/38 |
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MorphoBank Project 3422
MorphoBank Project 3422
- Creation Date:
13 March 2019 - Publication Date:
23 April 2020 - Project views: 12167
- Matrix downloads: 17
Authors' Institutions
- Natural History Museum, London
- Museum of New Zealand Te Papa Tongarewa, Wellington
- University of Oxford
- Université de Liège
- Royal Belgian Institute of Natural Sciences
- Royal Zeeland Scientific Society (Koninklijk Zeeuwsch Genootschap der Wetenschappen)
Members
member name | taxa ![]() |
specimens ![]() |
media ![]() | chars ![]() | character
| cell scorings (scored, NPA, "-") | cell
| rules ![]() | ||||||
Guillaume Duboys de Lavigerie Project Administrator | 110 | 1 | 1 | 278 | 0 | 0 | 18148 (16664, 157, 1327) | 0 | 0 | 0 | ||||
Felix Marx Full membership ![]() | 0 | 0 | 0 | 0 | 0 | 0 | 3 (2, 0, 1) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M25929' (110 Taxa)
taxon | unscored cells |
scored cells ![]() |
no cell support ![]() |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Zygorhiza kochii Last Modified in 04/30/20 | 18 | 253 | 253 | 0 | 7 | 0 | 0 | 2 ![]() |
[2] † Albertocetus meffordorum Last Modified in 04/30/20 | 77 | 189 | 189 | 0 | 12 | 0 | 0 | 2 ![]() |
[3] † Archaeodelphis patrius Last Modified in 04/30/20 | 167 | 102 | 102 | 0 | 9 | 0 | 0 | 2 ![]() |
[4] † Olympicetus avitus Last Modified in 04/30/20 | 131 | 138 | 138 | 0 | 9 | 0 | 0 | 2 ![]() |
[5] † Waipatia maerewhenua Last Modified in 04/30/20 | 45 | 220 | 220 | 0 | 13 | 0 | 0 | 2 ![]() |
[6] Physeter macrocephalus Last Modified in 04/06/20 | 11 | 229 | 227 | 0 | 40 | 0 | 0 | 2 ![]() |
[7] † Aetiocetus cotylalveus Last Modified in 04/30/20 | 111 | 165 | 165 | 0 | 2 | 0 | 0 | 2 ![]() |
[8] † Aetiocetus polydentatus Last Modified in 04/30/20 | 97 | 178 | 178 | 0 | 3 | 0 | 0 | 2 ![]() |
[9] † Aetiocetus weltoni Last Modified in 04/30/20 | 84 | 191 | 191 | 0 | 3 | 0 | 0 | 2 ![]() |
[10] † Aglaocetus moreni Last Modified in 04/30/20 | 123 | 142 | 142 | 0 | 13 | 0 | 0 | 2 ![]() |
[11] † Aglaocetus patulus Last Modified in 04/30/20 | 79 | 187 | 185 | 0 | 14 | 0 | 0 | 2 ![]() |
[12] † Antwerpibalaena liberatlas Last Modified in 04/06/20 | 0 | 112 | 112 | 152 | 14 | 0 | 0 | 2 ![]() |
[13] † Archaebalaenoptera castriarquati Last Modified in 04/30/20 | 156 | 99 | 99 | 5 | 18 | 0 | 0 | 2 ![]() |
[14] † Aulocetus latus Last Modified in 04/30/20 | 172 | 100 | 100 | 0 | 6 | 0 | 0 | 2 ![]() |
[15] † Balaena montalionis Last Modified in 04/30/20 | 188 | 71 | 71 | 0 | 19 | 0 | 0 | 2 ![]() |
[16] Balaena mysticetus Last Modified in 04/06/20 | 2 | 255 | 252 | 0 | 24 | 0 | 0 | 2 ![]() |
[17] † Balaena ricei Last Modified in 04/30/20 | 163 | 96 | 96 | 0 | 19 | 0 | 0 | 2 ![]() |
[18] † Balaenella brachyrhynus Last Modified in 04/30/20 | 123 | 134 | 134 | 0 | 21 | 0 | 0 | 2 ![]() |
[19] Balaenoptera acutorostrata Last Modified in 04/06/20 | 0 | 261 | 260 | 0 | 18 | 0 | 0 | 2 ![]() |
[20] † Balaenoptera bertae Last Modified in 04/30/20 | 201 | 70 | 70 | 0 | 7 | 0 | 0 | 2 ![]() |
[21] Balaenoptera bonaerensis Last Modified in 04/06/20 | 5 | 256 | 256 | 0 | 17 | 0 | 0 | 2 ![]() |
[22] Balaenoptera borealis Last Modified in 04/06/20 | 1 | 260 | 259 | 0 | 18 | 0 | 0 | 2 ![]() |
[23] Balaenoptera musculus Last Modified in 04/06/20 | 3 | 258 | 257 | 0 | 18 | 0 | 0 | 2 ![]() |
[24] Balaenoptera omurai Last Modified in 04/06/20 | 7 | 254 | 254 | 0 | 17 | 0 | 0 | 2 ![]() |
[25] Balaenoptera physalus Last Modified in 04/06/20 | 1 | 261 | 259 | 0 | 18 | 0 | 0 | 2 ![]() |
[26] † Balaenoptera portisi Last Modified in 04/30/20 | 83 | 182 | 182 | 0 | 13 | 0 | 0 | 2 ![]() |
[27] † Balaenoptera ryani Last Modified in 04/30/20 | 193 | 81 | 80 | 0 | 5 | 0 | 0 | 2 ![]() |
[28] † Balaenoptera siberi Last Modified in 04/30/20 | 104 | 159 | 157 | 0 | 17 | 0 | 0 | 2 ![]() |
[29] † Balaenula astensis Last Modified in 04/30/20 | 110 | 148 | 148 | 0 | 20 | 0 | 0 | 2 ![]() |
[30] † Balaenula sp. Last Modified in 04/30/20 | 134 | 125 | 125 | 0 | 19 | 0 | 0 | 2 ![]() |
[31] Brandtocetus chongulek Last Modified in 04/06/20 | 151 | 122 | 121 | 0 | 6 | 0 | 0 | 2 ![]() |
[32] Caperea marginata Last Modified in 04/30/20 | 3 | 254 | 248 | 0 | 27 | 0 | 0 | 2 ![]() |
[33] † Cephalotropis coronatus Last Modified in 04/30/20 | 127 | 138 | 138 | 0 | 13 | 0 | 0 | 2 ![]() |
[34] † Cetotherium megalophysum Last Modified in 04/30/20 | 144 | 117 | 116 | 0 | 18 | 0 | 0 | 2 ![]() |
[35] † Cetotherium rathkii Last Modified in 04/30/20 | 120 | 139 | 139 | 0 | 19 | 0 | 0 | 2 ![]() |
[36] † Cetotherium riabinini Last Modified in 04/30/20 | 73 | 187 | 187 | 0 | 18 | 0 | 0 | 2 ![]() |
[37] † Chonecetus sookensis Last Modified in 04/30/20 | 197 | 79 | 79 | 0 | 2 | 0 | 0 | 2 ![]() |
[38] † Ciuciulea davidi Last Modified in 04/30/20 | 189 | 86 | 86 | 0 | 3 | 0 | 0 | 2 ![]() |
[39] † Coronodon havensteini Last Modified in 04/30/20 | 70 | 204 | 203 | 0 | 5 | 0 | 0 | 2 ![]() |
[40] † Diorocetus chichibuensis Last Modified in 04/30/20 | 87 | 177 | 177 | 0 | 14 | 0 | 0 | 2 ![]() |
[41] † Diorocetus hiatus Last Modified in 04/30/20 | 48 | 216 | 216 | 0 | 14 | 0 | 0 | 2 ![]() |
[42] † Diorocetus shobarensis Last Modified in 04/30/20 | 138 | 127 | 126 | 0 | 14 | 0 | 0 | 2 ![]() |
[43] † Diunatans luctoretemergo Last Modified in 04/30/20 | 152 | 123 | 122 | 0 | 4 | 0 | 0 | 2 ![]() |
[44] † Eomysticetus whitmorei Last Modified in 04/30/20 | 81 | 181 | 181 | 0 | 16 | 0 | 0 | 2 ![]() |
[45] † Eschrichtioides gastaldii Last Modified in 04/30/20 | 113 | 146 | 146 | 0 | 19 | 0 | 0 | 2 ![]() |
[46] Eschrichtius robustus Last Modified in 04/06/20 | 0 | 265 | 260 | 0 | 18 | 0 | 0 | 2 ![]() |
[47] † Eubalaena ianitrix Last Modified in 04/30/20 | 203 | 69 | 69 | 0 | 6 | 0 | 0 | 2 ![]() |
[48] † Eubalaena shinshuensis Last Modified in 04/30/20 | 204 | 55 | 55 | 0 | 19 | 0 | 0 | 2 ![]() |
[49] Eubalaena spp. Last Modified in 04/06/20 | 0 | 260 | 254 | 0 | 24 | 0 | 0 | 2 ![]() |
[50] † Fragilicetus velponi Last Modified in 04/30/20 | 188 | 87 | 86 | 0 | 4 | 0 | 0 | 2 ![]() |
[51] † Fucaia buelli Last Modified in 04/30/20 | 126 | 149 | 149 | 0 | 3 | 0 | 0 | 2 ![]() |
[52] † Fucaia goedertorum Last Modified in 04/30/20 | 82 | 192 | 192 | 0 | 4 | 0 | 0 | 2 ![]() |
[53] † Gricetoides aurorae Last Modified in 04/30/20 | 195 | 82 | 82 | 0 | 1 | 0 | 0 | 2 ![]() |
[54] † Herentalia nigra Last Modified in 04/30/20 | 204 | 67 | 67 | 0 | 7 | 0 | 0 | 2 ![]() |
[55] † Herpetocetus bramblei Last Modified in 04/30/20 | 113 | 147 | 147 | 0 | 18 | 0 | 0 | 2 ![]() |
[56] † Herpetocetus morrowi Last Modified in 04/30/20 | 41 | 219 | 217 | 0 | 20 | 0 | 0 | 2 ![]() |
[57] † Herpetocetus sp. Last Modified in 04/30/20 | 207 | 64 | 64 | 0 | 7 | 0 | 0 | 2 ![]() |
[58] † Herpetocetus transatlanticus Last Modified in 04/30/20 | 155 | 116 | 116 | 0 | 7 | 0 | 0 | 2 ![]() |
[59] † Horopeta umarere Last Modified in 04/30/20 | 147 | 118 | 118 | 0 | 13 | 0 | 0 | 2 ![]() |
[60] † Incakujira anillodefuego Last Modified in 04/30/20 | 59 | 204 | 202 | 0 | 17 | 0 | 0 | 2 ![]() |
[61] † Isanacetus laticephalus Last Modified in 04/30/20 | 66 | 197 | 196 | 0 | 16 | 0 | 0 | 2 ![]() |
[62] † Janjucetus hunderi Last Modified in 04/30/20 | 90 | 182 | 182 | 0 | 6 | 0 | 0 | 2 ![]() |
[63] † Joumocetus shimizui Last Modified in 04/30/20 | 146 | 116 | 116 | 0 | 16 | 0 | 0 | 2 ![]() |
[64] † Kurdalagonus mchedlidzei Last Modified in 04/30/20 | 132 | 129 | 129 | 0 | 17 | 0 | 0 | 2 ![]() |
[65] † Llanocetus denticrenatus Last Modified in 04/30/20 | 92 | 181 | 181 | 0 | 5 | 0 | 0 | 2 ![]() |
[66] † Mammalodon colliveri Last Modified in 04/30/20 | 93 | 178 | 178 | 0 | 7 | 0 | 0 | 2 ![]() |
[67] † Mammalodon hakataramea Last Modified in 04/30/20 | 242 | 37 | 36 | 0 | 0 | 0 | 0 | 2 ![]() |
[68] † Matapanui waihao Last Modified in 04/30/20 | 141 | 121 | 121 | 0 | 16 | 0 | 0 | 2 ![]() |
[69] † Mauicetus parki Last Modified in 04/30/20 | 81 | 184 | 183 | 0 | 14 | 0 | 0 | 2 ![]() |
[70] † Megaptera hubachi Last Modified in 04/30/20 | 75 | 185 | 185 | 0 | 18 | 0 | 0 | 2 ![]() |
[71] † Megaptera miocaena Last Modified in 04/30/20 | 109 | 150 | 150 | 0 | 19 | 0 | 0 | 2 ![]() |
[72] Megaptera novaeangliae Last Modified in 04/06/20 | 1 | 263 | 261 | 0 | 16 | 0 | 0 | 2 ![]() |
[73] † Metopocetus durinasus Last Modified in 04/30/20 | 191 | 81 | 81 | 0 | 6 | 0 | 0 | 2 ![]() |
[74] † Metopocetus hunteri Last Modified in 04/30/20 | 165 | 107 | 106 | 0 | 7 | 0 | 0 | 2 ![]() |
[75] † Metopocetus vandelli Last Modified in 04/30/20 | 172 | 101 | 100 | 0 | 6 | 0 | 0 | 2 ![]() |
[76] † Micromysticetus rothauseni Last Modified in 04/30/20 | 186 | 90 | 90 | 0 | 2 | 0 | 0 | 2 ![]() |
[77] † Miocaperea pulchra Last Modified in 04/30/20 | 120 | 137 | 136 | 0 | 22 | 0 | 0 | 2 ![]() |
[78] † Morawanocetus yabukii Last Modified in 04/30/20 | 130 | 143 | 143 | 0 | 5 | 0 | 0 | 2 ![]() |
[79] † Morenocetus parvus Last Modified in 04/30/20 | 160 | 113 | 113 | 0 | 5 | 0 | 0 | 2 ![]() |
[80] † Mystacodon selenensis Last Modified in 04/30/20 | 146 | 127 | 127 | 0 | 5 | 0 | 0 | 2 ![]() |
[81] † Nannocetus eremus Last Modified in 04/30/20 | 187 | 89 | 89 | 0 | 2 | 0 | 0 | 2 ![]() |
[82] † NMNS pv19540 Last Modified in 04/30/20 | 90 | 170 | 169 | 0 | 19 | 0 | 0 | 2 ![]() |
[83] † NMNZ mm001630 Last Modified in 04/30/20 | 64 | 195 | 195 | 0 | 19 | 0 | 0 | 2 ![]() |
[84] † OCPC 1178 Last Modified in 04/30/20 | 185 | 91 | 91 | 0 | 2 | 0 | 0 | 2 ![]() |
[85] † OU 22224 Last Modified in 04/30/20 | 169 | 107 | 107 | 0 | 2 | 0 | 0 | 2 ![]() |
[86] † OU 22705 Last Modified in 04/30/20 | 110 | 153 | 153 | 0 | 15 | 0 | 0 | 2 ![]() |
[87] † OU gs10897 Last Modified in 04/30/20 | 219 | 57 | 56 | 0 | 3 | 0 | 0 | 2 ![]() |
[88] † Parabalaenoptera baulinensis Last Modified in 04/30/20 | 98 | 164 | 164 | 0 | 16 | 0 | 0 | 2 ![]() |
[89] † Parietobalaena campiniana Last Modified in 04/30/20 | 109 | 154 | 154 | 0 | 15 | 0 | 0 | 2 ![]() |
[90] † Parietobalaena palmeri Last Modified in 04/30/20 | 50 | 219 | 214 | 0 | 14 | 0 | 0 | 2 ![]() |
[91] † Parietobalaena sp. Last Modified in 04/30/20 | 106 | 156 | 156 | 0 | 16 | 0 | 0 | 2 ![]() |
[92] † Parietobalaena yamaokai Last Modified in 04/30/20 | 91 | 176 | 173 | 0 | 14 | 0 | 0 | 2 ![]() |
[93] † Pelocetus calvertensis Last Modified in 04/30/20 | 41 | 222 | 222 | 0 | 15 | 0 | 0 | 2 ![]() |
[94] † Peripolocetus vexillifer Last Modified in 04/30/20 | 217 | 60 | 60 | 0 | 1 | 0 | 0 | 2 ![]() |
[95] † Pinocetus polonicus Last Modified in 04/30/20 | 150 | 115 | 115 | 0 | 13 | 0 | 0 | 2 ![]() |
[96] † Piscobalaena nana Last Modified in 04/30/20 | 15 | 244 | 243 | 0 | 20 | 0 | 0 | 2 ![]() |
[97] † Plesiobalaenoptera quarantellii Last Modified in 04/30/20 | 122 | 138 | 138 | 0 | 18 | 0 | 0 | 2 ![]() |
[98] † Thinocetus arthritus Last Modified in 04/30/20 | 120 | 145 | 145 | 0 | 13 | 0 | 0 | 2 ![]() |
[99] † Tiphyocetus temblorensis Last Modified in 04/30/20 | 207 | 71 | 71 | 0 | 0 | 0 | 0 | 2 ![]() |
[100] † Titanocetus sammarinensis Last Modified in 04/30/20 | 125 | 141 | 141 | 0 | 12 | 0 | 0 | 2 ![]() |
[101] † Tiucetus rosae Last Modified in 04/30/20 | 138 | 139 | 139 | 0 | 1 | 0 | 0 | 2 ![]() |
[102] † Tohoraata raekohao Last Modified in 04/30/20 | 161 | 115 | 115 | 0 | 2 | 0 | 0 | 2 ![]() |
[103] † Tokarahia kauaeroa Last Modified in 04/30/20 | 115 | 149 | 149 | 0 | 14 | 0 | 0 | 2 ![]() |
[104] † Tranatocetus argillarius Last Modified in 04/30/20 | 163 | 107 | 107 | 0 | 8 | 0 | 0 | 2 ![]() |
[105] † Tranatocetus maregermanicum Last Modified in 04/30/20 | 156 | 115 | 114 | 0 | 8 | 0 | 0 | 2 ![]() |
[106] † Uranocetus gramensis Last Modified in 04/30/20 | 86 | 175 | 175 | 0 | 17 | 0 | 0 | 2 ![]() |
[107] † Waharoa ruwhenua Last Modified in 04/30/20 | 63 | 199 | 197 | 0 | 18 | 0 | 0 | 2 ![]() |
[108] † Whakakai waipata Last Modified in 04/30/20 | 197 | 80 | 80 | 0 | 1 | 0 | 0 | 2 ![]() |
[109] † Yamatocetus canaliculatus Last Modified in 04/30/20 | 71 | 192 | 192 | 0 | 15 | 0 | 0 | 2 ![]() |
[110] † ZMT 67 Last Modified in 04/30/20 | 124 | 140 | 140 | 0 | 14 | 0 | 0 | 2 ![]() |
Project views 
type | number of views | Individual items viewed (where applicable) |
Total project views | 12167 | |
Project overview | 958 | |
Matrix views | 730 | Matrix landing page (626 views); Mysticete phylogeny with a new fossil balaenids taxon, Antwerpibalaena liberatlas (104 views); |
Taxon list | 6189 | |
Media views | 1890 | Media search (1603 views); M687533 (287 views); |
Documents list | 474 | |
Views for media list | 593 | |
Specimen list | 951 | |
Bibliography | 382 |
Project downloads 
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 135 | |
Matrix downloads | 17 | Mysticete phylogeny with a new fossil balaenids taxon, Antwerpibalaena liberatlas (17 downloads); |
Project downloads | 118 |