Morphobank

Project 3422: G. D. de Lavigerie, M. Bosselaers, S. Goolaerts, T. Park, O. Lambert, F. G. Marx. 2020. New Pliocene right whale from Belgium informs balaenid phylogeny and function. Journal of Systematic Palaeontology. -:1-26.

Specimen: † Antwerpibalaena liberatlas (IRSNB 12345/IRSNB 12345)
View: Articulated left ear bones of the holotype of Antwerpibalaena liberatlas (IRSNB M2325). Photographs in A, anteroventral and B, ventral views.

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Abstract

Right whales (Balaenidae) are the most distinctive family of extant baleen whales, thanks to their highly arched rostrum, tall lips and robust body shape. They are also the oldest, originating as much as 20 million years ago (Ma). Nevertheless, their fossil record is patchy and frequently understudied, obscuring their evolution. Here, we describe a new stem balaenid, Antwerpibalaena liberatlas, from northern Belgium, adding to the rich but historically problematic baleen whale assemblage of the Pliocene North Sea. Within right whales, Antwerpibalaena forms a clade with two previously described extinct genera, Balaenella and Balaenula. The holotype preserves much of the postcranial skeleton, and informs the emergence of typical balaenid traits like fused neck vertebrae and paddle-shaped flippers. Its size is intermediate between that of extant right whales and most of their extinct forebears revealing a more complex pattern of balaenid size evolution than previously thought.

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Article DOI: 10.1080/14772019.2020.1746422

Project DOI: 10.7934/P3422, http://dx.doi.org/10.7934/P3422

This project contains	Matrices
1 Media 1 Matrix 1 Document 110 Taxa 1 Specimen 278 Characters Total size of project's media files: 1.9M Download Project SDD File	Total scored cells: 18151 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 278 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 651 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:240/38

Currently Viewing:
MorphoBank Project 3422

Creation Date:
13 March 2019
Publication Date:
23 April 2020
Project views: 15466
Matrix downloads: 18

Authors' Institutions

Natural History Museum, London

Museum of New Zealand Te Papa Tongarewa, Wellington

University of Oxford

Université de Liège

Royal Belgian Institute of Natural Sciences

Royal Zeeland Scientific Society (Koninklijk Zeeuwsch Genootschap der Wetenschappen)

Members

Guillaume Duboys de Lavigerie
Project Administrator

110

278

18148
(16664, 157, 1327)

Felix Marx
Full membership

3
(2, 0, 1)

Taxonomic Overview for Matrix 'M25929' (110 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Zygorhiza kochii Last Modified in 04/30/20	18	253	253	0	7	0	0	2
[2] † Albertocetus meffordorum Last Modified in 04/30/20	77	189	189	0	12	0	0	2
[3] † Archaeodelphis patrius Last Modified in 04/30/20	167	102	102	0	9	0	0	2
[4] † Olympicetus avitus Last Modified in 04/30/20	131	138	138	0	9	0	0	2
[5] † Waipatia maerewhenua Last Modified in 04/30/20	45	220	220	0	13	0	0	2
[6] Physeter macrocephalus Last Modified in 04/06/20	11	229	227	0	40	0	0	2
[7] † Aetiocetus cotylalveus Last Modified in 04/30/20	111	165	165	0	2	0	0	2
[8] † Aetiocetus polydentatus Last Modified in 04/30/20	97	178	178	0	3	0	0	2
[9] † Aetiocetus weltoni Last Modified in 04/30/20	84	191	191	0	3	0	0	2
[10] † Aglaocetus moreni Last Modified in 04/30/20	123	142	142	0	13	0	0	2
[11] † Aglaocetus patulus Last Modified in 04/30/20	79	187	185	0	14	0	0	2
[12] † Antwerpibalaena liberatlas Last Modified in 04/06/20	0	112	112	152	14	0	0	2
[13] † Archaebalaenoptera castriarquati Last Modified in 04/30/20	156	99	99	5	18	0	0	2
[14] † Aulocetus latus Last Modified in 04/30/20	172	100	100	0	6	0	0	2
[15] † Balaena montalionis Last Modified in 04/30/20	188	71	71	0	19	0	0	2
[16] Balaena mysticetus Last Modified in 04/06/20	2	255	252	0	24	0	0	2
[17] † Balaena ricei Last Modified in 04/30/20	163	96	96	0	19	0	0	2
[18] † Balaenella brachyrhynus Last Modified in 04/30/20	123	134	134	0	21	0	0	2
[19] Balaenoptera acutorostrata Last Modified in 04/06/20	0	261	260	0	18	0	0	2
[20] † Balaenoptera bertae Last Modified in 04/30/20	201	70	70	0	7	0	0	2
[21] Balaenoptera bonaerensis Last Modified in 04/06/20	5	256	256	0	17	0	0	2
[22] Balaenoptera borealis Last Modified in 04/06/20	1	260	259	0	18	0	0	2
[23] Balaenoptera musculus Last Modified in 04/06/20	3	258	257	0	18	0	0	2
[24] Balaenoptera omurai Last Modified in 04/06/20	7	254	254	0	17	0	0	2
[25] Balaenoptera physalus Last Modified in 04/06/20	1	261	259	0	18	0	0	2
[26] † Balaenoptera portisi Last Modified in 04/30/20	83	182	182	0	13	0	0	2
[27] † Balaenoptera ryani Last Modified in 04/30/20	193	81	80	0	5	0	0	2
[28] † Balaenoptera siberi Last Modified in 04/30/20	104	159	157	0	17	0	0	2
[29] † Balaenula astensis Last Modified in 04/30/20	110	148	148	0	20	0	0	2
[30] † Balaenula sp. Last Modified in 04/30/20	134	125	125	0	19	0	0	2
[31] Brandtocetus chongulek Last Modified in 04/06/20	151	122	121	0	6	0	0	2
[32] Caperea marginata Last Modified in 04/30/20	3	254	248	0	27	0	0	2
[33] † Cephalotropis coronatus Last Modified in 04/30/20	127	138	138	0	13	0	0	2
[34] † Cetotherium megalophysum Last Modified in 04/30/20	144	117	116	0	18	0	0	2
[35] † Cetotherium rathkii Last Modified in 04/30/20	120	139	139	0	19	0	0	2
[36] † Cetotherium riabinini Last Modified in 04/30/20	73	187	187	0	18	0	0	2
[37] † Chonecetus sookensis Last Modified in 04/30/20	197	79	79	0	2	0	0	2
[38] † Ciuciulea davidi Last Modified in 04/30/20	189	86	86	0	3	0	0	2
[39] † Coronodon havensteini Last Modified in 04/30/20	70	204	203	0	5	0	0	2
[40] † Diorocetus chichibuensis Last Modified in 04/30/20	87	177	177	0	14	0	0	2
[41] † Diorocetus hiatus Last Modified in 04/30/20	48	216	216	0	14	0	0	2
[42] † Diorocetus shobarensis Last Modified in 04/30/20	138	127	126	0	14	0	0	2
[43] † Diunatans luctoretemergo Last Modified in 04/30/20	152	123	122	0	4	0	0	2
[44] † Eomysticetus whitmorei Last Modified in 04/30/20	81	181	181	0	16	0	0	2
[45] † Eschrichtioides gastaldii Last Modified in 04/30/20	113	146	146	0	19	0	0	2
[46] Eschrichtius robustus Last Modified in 04/06/20	0	265	260	0	18	0	0	2
[47] † Eubalaena ianitrix Last Modified in 04/30/20	203	69	69	0	6	0	0	2
[48] † Eubalaena shinshuensis Last Modified in 04/30/20	204	55	55	0	19	0	0	2
[49] Eubalaena spp. Last Modified in 04/06/20	0	260	254	0	24	0	0	2
[50] † Fragilicetus velponi Last Modified in 04/30/20	188	87	86	0	4	0	0	2
[51] † Fucaia buelli Last Modified in 04/30/20	126	149	149	0	3	0	0	2
[52] † Fucaia goedertorum Last Modified in 04/30/20	82	192	192	0	4	0	0	2
[53] † Gricetoides aurorae Last Modified in 04/30/20	195	82	82	0	1	0	0	2
[54] † Herentalia nigra Last Modified in 04/30/20	204	67	67	0	7	0	0	2
[55] † Herpetocetus bramblei Last Modified in 04/30/20	113	147	147	0	18	0	0	2
[56] † Herpetocetus morrowi Last Modified in 04/30/20	41	219	217	0	20	0	0	2
[57] † Herpetocetus sp. Last Modified in 04/30/20	207	64	64	0	7	0	0	2
[58] † Herpetocetus transatlanticus Last Modified in 04/30/20	155	116	116	0	7	0	0	2
[59] † Horopeta umarere Last Modified in 04/30/20	147	118	118	0	13	0	0	2
[60] † Incakujira anillodefuego Last Modified in 04/30/20	59	204	202	0	17	0	0	2
[61] † Isanacetus laticephalus Last Modified in 04/30/20	66	197	196	0	16	0	0	2
[62] † Janjucetus hunderi Last Modified in 04/30/20	90	182	182	0	6	0	0	2
[63] † Joumocetus shimizui Last Modified in 04/30/20	146	116	116	0	16	0	0	2
[64] † Kurdalagonus mchedlidzei Last Modified in 04/30/20	132	129	129	0	17	0	0	2
[65] † Llanocetus denticrenatus Last Modified in 04/30/20	92	181	181	0	5	0	0	2
[66] † Mammalodon colliveri Last Modified in 04/30/20	93	178	178	0	7	0	0	2
[67] † Mammalodon hakataramea Last Modified in 04/30/20	242	37	36	0	0	0	0	2
[68] † Matapanui waihao Last Modified in 04/30/20	141	121	121	0	16	0	0	2
[69] † Mauicetus parki Last Modified in 04/30/20	81	184	183	0	14	0	0	2
[70] † Megaptera hubachi Last Modified in 04/30/20	75	185	185	0	18	0	0	2
[71] † Megaptera miocaena Last Modified in 04/30/20	109	150	150	0	19	0	0	2
[72] Megaptera novaeangliae Last Modified in 04/06/20	1	263	261	0	16	0	0	2
[73] † Metopocetus durinasus Last Modified in 04/30/20	191	81	81	0	6	0	0	2
[74] † Metopocetus hunteri Last Modified in 04/30/20	165	107	106	0	7	0	0	2
[75] † Metopocetus vandelli Last Modified in 04/30/20	172	101	100	0	6	0	0	2
[76] † Micromysticetus rothauseni Last Modified in 04/30/20	186	90	90	0	2	0	0	2
[77] † Miocaperea pulchra Last Modified in 04/30/20	120	137	136	0	22	0	0	2
[78] † Morawanocetus yabukii Last Modified in 04/30/20	130	143	143	0	5	0	0	2
[79] † Morenocetus parvus Last Modified in 04/30/20	160	113	113	0	5	0	0	2
[80] † Mystacodon selenensis Last Modified in 04/30/20	146	127	127	0	5	0	0	2
[81] † Nannocetus eremus Last Modified in 04/30/20	187	89	89	0	2	0	0	2
[82] † NMNS pv19540 Last Modified in 04/30/20	90	170	169	0	19	0	0	2
[83] † NMNZ mm001630 Last Modified in 04/30/20	64	195	195	0	19	0	0	2
[84] † OCPC 1178 Last Modified in 04/30/20	185	91	91	0	2	0	0	2
[85] † OU 22224 Last Modified in 04/30/20	169	107	107	0	2	0	0	2
[86] † OU 22705 Last Modified in 04/30/20	110	153	153	0	15	0	0	2
[87] † OU gs10897 Last Modified in 04/30/20	219	57	56	0	3	0	0	2
[88] † Parabalaenoptera baulinensis Last Modified in 04/30/20	98	164	164	0	16	0	0	2
[89] † Parietobalaena campiniana Last Modified in 04/30/20	109	154	154	0	15	0	0	2
[90] † Parietobalaena palmeri Last Modified in 04/30/20	50	219	214	0	14	0	0	2
[91] † Parietobalaena sp. Last Modified in 04/30/20	106	156	156	0	16	0	0	2
[92] † Parietobalaena yamaokai Last Modified in 04/30/20	91	176	173	0	14	0	0	2
[93] † Pelocetus calvertensis Last Modified in 04/30/20	41	222	222	0	15	0	0	2
[94] † Peripolocetus vexillifer Last Modified in 04/30/20	217	60	60	0	1	0	0	2
[95] † Pinocetus polonicus Last Modified in 04/30/20	150	115	115	0	13	0	0	2
[96] † Piscobalaena nana Last Modified in 04/30/20	15	244	243	0	20	0	0	2
[97] † Plesiobalaenoptera quarantellii Last Modified in 04/30/20	122	138	138	0	18	0	0	2
[98] † Thinocetus arthritus Last Modified in 04/30/20	120	145	145	0	13	0	0	2
[99] † Tiphyocetus temblorensis Last Modified in 04/30/20	207	71	71	0	0	0	0	2
[100] † Titanocetus sammarinensis Last Modified in 04/30/20	125	141	141	0	12	0	0	2
[101] † Tiucetus rosae Last Modified in 04/30/20	138	139	139	0	1	0	0	2
[102] † Tohoraata raekohao Last Modified in 04/30/20	161	115	115	0	2	0	0	2
[103] † Tokarahia kauaeroa Last Modified in 04/30/20	115	149	149	0	14	0	0	2
[104] † Tranatocetus argillarius Last Modified in 04/30/20	163	107	107	0	8	0	0	2
[105] † Tranatocetus maregermanicum Last Modified in 04/30/20	156	115	114	0	8	0	0	2
[106] † Uranocetus gramensis Last Modified in 04/30/20	86	175	175	0	17	0	0	2
[107] † Waharoa ruwhenua Last Modified in 04/30/20	63	199	197	0	18	0	0	2
[108] † Whakakai waipata Last Modified in 04/30/20	197	80	80	0	1	0	0	2
[109] † Yamatocetus canaliculatus Last Modified in 04/30/20	71	192	192	0	15	0	0	2
[110] † ZMT 67 Last Modified in 04/30/20	124	140	140	0	14	0	0	2

Project views

type	number of views	Individual items viewed (where applicable)
Total project views	15466
Project overview	1211
Matrix views	873	Matrix landing page (751 views); Mysticete phylogeny with a new fossil balaenids taxon, Antwerpibalaena liberatlas (122 views);
Taxon list	7970
Media views	2366	Media search (2008 views); M687533 (358 views);
Documents list	601
Views for media list	718
Specimen list	1246
Bibliography	481

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	138
Matrix downloads	18	Mysticete phylogeny with a new fossil balaenids taxon, Antwerpibalaena liberatlas (18 downloads);
Project downloads	120