Morphobank

Project 3422: G. D. de Lavigerie, M. Bosselaers, S. Goolaerts, T. Park, O. Lambert, F. G. Marx. 2020. New Pliocene right whale from Belgium informs balaenid phylogeny and function. Journal of Systematic Palaeontology. -:1-26.

Specimen: † Antwerpibalaena liberatlas (IRSNB 12345/IRSNB 12345)
View: Articulated left ear bones of the holotype of Antwerpibalaena liberatlas (IRSNB M2325). Photographs in A, anteroventral and B, ventral views.

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Abstract

Right whales (Balaenidae) are the most distinctive family of extant baleen whales, thanks to their highly arched rostrum, tall lips and robust body shape. They are also the oldest, originating as much as 20 million years ago (Ma). Nevertheless, their fossil record is patchy and frequently understudied, obscuring their evolution. Here, we describe a new stem balaenid, Antwerpibalaena liberatlas, from northern Belgium, adding to the rich but historically problematic baleen whale assemblage of the Pliocene North Sea. Within right whales, Antwerpibalaena forms a clade with two previously described extinct genera, Balaenella and Balaenula. The holotype preserves much of the postcranial skeleton, and informs the emergence of typical balaenid traits like fused neck vertebrae and paddle-shaped flippers. Its size is intermediate between that of extant right whales and most of their extinct forebears revealing a more complex pattern of balaenid size evolution than previously thought.

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Article DOI: 10.1080/14772019.2020.1746422

Project DOI: 10.7934/P3422, http://dx.doi.org/10.7934/P3422

This project contains	Matrices
1 Media 1 Matrix 1 Document 110 Taxa 1 Specimen 278 Characters Total size of project's media files: 1.9M Download Project SDD File	Total scored cells: 18151 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 278 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 651 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:240/38

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MorphoBank Project 3422

Creation Date:
13 March 2019
Publication Date:
23 April 2020
Matrix downloads: 21

Authors' Institutions

Natural History Museum, London

Museum of New Zealand Te Papa Tongarewa, Wellington

University of Oxford

Université de Liège

Royal Belgian Institute of Natural Sciences

Royal Zeeland Scientific Society (Koninklijk Zeeuwsch Genootschap der Wetenschappen)

Members

Guillaume Duboys de Lavigerie
Project Administrator

110

278

18148
(16664, 157, 1327)

Felix Marx
Full membership

3
(2, 0, 1)

Taxonomic Overview for Matrix 'M25929' (110 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Zygorhiza kochii Taxon name last Modified on 04/30/20	18	253	253	0	7	0	0	2
[2] † Albertocetus meffordorum Taxon name last Modified on 04/30/20	77	189	189	0	12	0	0	2
[3] † Archaeodelphis patrius Taxon name last Modified on 04/30/20	167	102	102	0	9	0	0	2
[4] † Olympicetus avitus Taxon name last Modified on 04/30/20	131	138	138	0	9	0	0	2
[5] † Waipatia maerewhenua Taxon name last Modified on 04/30/20	45	220	220	0	13	0	0	2
[6] Physeter macrocephalus Taxon name last Modified on 04/06/20	11	229	227	0	40	0	0	2
[7] † Aetiocetus cotylalveus Taxon name last Modified on 04/30/20	111	165	165	0	2	0	0	2
[8] † Aetiocetus polydentatus Taxon name last Modified on 04/30/20	97	178	178	0	3	0	0	2
[9] † Aetiocetus weltoni Taxon name last Modified on 04/30/20	84	191	191	0	3	0	0	2
[10] † Aglaocetus moreni Taxon name last Modified on 04/30/20	123	142	142	0	13	0	0	2
[11] † Aglaocetus patulus Taxon name last Modified on 04/30/20	79	187	185	0	14	0	0	2
[12] † Antwerpibalaena liberatlas Taxon name last Modified on 04/06/20	0	112	112	152	14	0	0	2
[13] † Archaebalaenoptera castriarquati Taxon name last Modified on 04/30/20	156	99	99	5	18	0	0	2
[14] † Aulocetus latus Taxon name last Modified on 04/30/20	172	100	100	0	6	0	0	2
[15] † Balaena montalionis Taxon name last Modified on 04/30/20	188	71	71	0	19	0	0	2
[16] Balaena mysticetus Taxon name last Modified on 04/06/20	2	255	252	0	24	0	0	2
[17] † Balaena ricei Taxon name last Modified on 04/30/20	163	96	96	0	19	0	0	2
[18] † Balaenella brachyrhynus Taxon name last Modified on 04/30/20	123	134	134	0	21	0	0	2
[19] Balaenoptera acutorostrata Taxon name last Modified on 04/06/20	0	261	260	0	18	0	0	2
[20] † Balaenoptera bertae Taxon name last Modified on 04/30/20	201	70	70	0	7	0	0	2
[21] Balaenoptera bonaerensis Taxon name last Modified on 04/06/20	5	256	256	0	17	0	0	2
[22] Balaenoptera borealis Taxon name last Modified on 04/06/20	1	260	259	0	18	0	0	2
[23] Balaenoptera musculus Taxon name last Modified on 04/06/20	3	258	257	0	18	0	0	2
[24] Balaenoptera omurai Taxon name last Modified on 04/06/20	7	254	254	0	17	0	0	2
[25] Balaenoptera physalus Taxon name last Modified on 04/06/20	1	261	259	0	18	0	0	2
[26] † Balaenoptera portisi Taxon name last Modified on 04/30/20	83	182	182	0	13	0	0	2
[27] † Balaenoptera ryani Taxon name last Modified on 04/30/20	193	81	80	0	5	0	0	2
[28] † Balaenoptera siberi Taxon name last Modified on 04/30/20	104	159	157	0	17	0	0	2
[29] † Balaenula astensis Taxon name last Modified on 04/30/20	110	148	148	0	20	0	0	2
[30] † Balaenula sp. Taxon name last Modified on 04/30/20	134	125	125	0	19	0	0	2
[31] Brandtocetus chongulek Taxon name last Modified on 04/06/20	151	122	121	0	6	0	0	2
[32] Caperea marginata Taxon name last Modified on 04/30/20	3	254	248	0	27	0	0	2
[33] † Cephalotropis coronatus Taxon name last Modified on 04/30/20	127	138	138	0	13	0	0	2
[34] † Cetotherium megalophysum Taxon name last Modified on 04/30/20	144	117	116	0	18	0	0	2
[35] † Cetotherium rathkii Taxon name last Modified on 04/30/20	120	139	139	0	19	0	0	2
[36] † Cetotherium riabinini Taxon name last Modified on 04/30/20	73	187	187	0	18	0	0	2
[37] † Chonecetus sookensis Taxon name last Modified on 04/30/20	197	79	79	0	2	0	0	2
[38] † Ciuciulea davidi Taxon name last Modified on 04/30/20	189	86	86	0	3	0	0	2
[39] † Coronodon havensteini Taxon name last Modified on 04/30/20	70	204	203	0	5	0	0	2
[40] † Diorocetus chichibuensis Taxon name last Modified on 04/30/20	87	177	177	0	14	0	0	2
[41] † Diorocetus hiatus Taxon name last Modified on 04/30/20	48	216	216	0	14	0	0	2
[42] † Diorocetus shobarensis Taxon name last Modified on 04/30/20	138	127	126	0	14	0	0	2
[43] † Diunatans luctoretemergo Taxon name last Modified on 04/30/20	152	123	122	0	4	0	0	2
[44] † Eomysticetus whitmorei Taxon name last Modified on 04/30/20	81	181	181	0	16	0	0	2
[45] † Eschrichtioides gastaldii Taxon name last Modified on 04/30/20	113	146	146	0	19	0	0	2
[46] Eschrichtius robustus Taxon name last Modified on 04/06/20	0	265	260	0	18	0	0	2
[47] † Eubalaena ianitrix Taxon name last Modified on 04/30/20	203	69	69	0	6	0	0	2
[48] † Eubalaena shinshuensis Taxon name last Modified on 04/30/20	204	55	55	0	19	0	0	2
[49] Eubalaena spp. Taxon name last Modified on 04/06/20	0	260	254	0	24	0	0	2
[50] † Fragilicetus velponi Taxon name last Modified on 04/30/20	188	87	86	0	4	0	0	2
[51] † Fucaia buelli Taxon name last Modified on 04/30/20	126	149	149	0	3	0	0	2
[52] † Fucaia goedertorum Taxon name last Modified on 04/30/20	82	192	192	0	4	0	0	2
[53] † Gricetoides aurorae Taxon name last Modified on 04/30/20	195	82	82	0	1	0	0	2
[54] † Herentalia nigra Taxon name last Modified on 04/30/20	204	67	67	0	7	0	0	2
[55] † Herpetocetus bramblei Taxon name last Modified on 04/30/20	113	147	147	0	18	0	0	2
[56] † Herpetocetus morrowi Taxon name last Modified on 04/30/20	41	219	217	0	20	0	0	2
[57] † Herpetocetus sp. Taxon name last Modified on 04/30/20	207	64	64	0	7	0	0	2
[58] † Herpetocetus transatlanticus Taxon name last Modified on 04/30/20	155	116	116	0	7	0	0	2
[59] † Horopeta umarere Taxon name last Modified on 04/30/20	147	118	118	0	13	0	0	2
[60] † Incakujira anillodefuego Taxon name last Modified on 04/30/20	59	204	202	0	17	0	0	2
[61] † Isanacetus laticephalus Taxon name last Modified on 04/30/20	66	197	196	0	16	0	0	2
[62] † Janjucetus hunderi Taxon name last Modified on 04/30/20	90	182	182	0	6	0	0	2
[63] † Joumocetus shimizui Taxon name last Modified on 04/30/20	146	116	116	0	16	0	0	2
[64] † Kurdalagonus mchedlidzei Taxon name last Modified on 04/30/20	132	129	129	0	17	0	0	2
[65] † Llanocetus denticrenatus Taxon name last Modified on 04/30/20	92	181	181	0	5	0	0	2
[66] † Mammalodon colliveri Taxon name last Modified on 04/30/20	93	178	178	0	7	0	0	2
[67] † Mammalodon hakataramea Taxon name last Modified on 04/30/20	242	37	36	0	0	0	0	2
[68] † Matapanui waihao Taxon name last Modified on 04/30/20	141	121	121	0	16	0	0	2
[69] † Mauicetus parki Taxon name last Modified on 04/30/20	81	184	183	0	14	0	0	2
[70] † Megaptera hubachi Taxon name last Modified on 04/30/20	75	185	185	0	18	0	0	2
[71] † Megaptera miocaena Taxon name last Modified on 04/30/20	109	150	150	0	19	0	0	2
[72] Megaptera novaeangliae Taxon name last Modified on 04/06/20	1	263	261	0	16	0	0	2
[73] † Metopocetus durinasus Taxon name last Modified on 04/30/20	191	81	81	0	6	0	0	2
[74] † Metopocetus hunteri Taxon name last Modified on 04/30/20	165	107	106	0	7	0	0	2
[75] † Metopocetus vandelli Taxon name last Modified on 04/30/20	172	101	100	0	6	0	0	2
[76] † Micromysticetus rothauseni Taxon name last Modified on 04/30/20	186	90	90	0	2	0	0	2
[77] † Miocaperea pulchra Taxon name last Modified on 04/30/20	120	137	136	0	22	0	0	2
[78] † Morawanocetus yabukii Taxon name last Modified on 04/30/20	130	143	143	0	5	0	0	2
[79] † Morenocetus parvus Taxon name last Modified on 04/30/20	160	113	113	0	5	0	0	2
[80] † Mystacodon selenensis Taxon name last Modified on 04/30/20	146	127	127	0	5	0	0	2
[81] † Nannocetus eremus Taxon name last Modified on 04/30/20	187	89	89	0	2	0	0	2
[82] † NMNS pv19540 Taxon name last Modified on 04/30/20	90	170	169	0	19	0	0	2
[83] † NMNZ mm001630 Taxon name last Modified on 04/30/20	64	195	195	0	19	0	0	2
[84] † OCPC 1178 Taxon name last Modified on 04/30/20	185	91	91	0	2	0	0	2
[85] † OU 22224 Taxon name last Modified on 04/30/20	169	107	107	0	2	0	0	2
[86] † OU 22705 Taxon name last Modified on 04/30/20	110	153	153	0	15	0	0	2
[87] † OU gs10897 Taxon name last Modified on 04/30/20	219	57	56	0	3	0	0	2
[88] † Parabalaenoptera baulinensis Taxon name last Modified on 04/30/20	98	164	164	0	16	0	0	2
[89] † Parietobalaena campiniana Taxon name last Modified on 04/30/20	109	154	154	0	15	0	0	2
[90] † Parietobalaena palmeri Taxon name last Modified on 04/30/20	50	219	214	0	14	0	0	2
[91] † Parietobalaena sp. Taxon name last Modified on 04/30/20	106	156	156	0	16	0	0	2
[92] † Parietobalaena yamaokai Taxon name last Modified on 04/30/20	91	176	173	0	14	0	0	2
[93] † Pelocetus calvertensis Taxon name last Modified on 04/30/20	41	222	222	0	15	0	0	2
[94] † Peripolocetus vexillifer Taxon name last Modified on 04/30/20	217	60	60	0	1	0	0	2
[95] † Pinocetus polonicus Taxon name last Modified on 04/30/20	150	115	115	0	13	0	0	2
[96] † Piscobalaena nana Taxon name last Modified on 04/30/20	15	244	243	0	20	0	0	2
[97] † Plesiobalaenoptera quarantellii Taxon name last Modified on 04/30/20	122	138	138	0	18	0	0	2
[98] † Thinocetus arthritus Taxon name last Modified on 04/30/20	120	145	145	0	13	0	0	2
[99] † Tiphyocetus temblorensis Taxon name last Modified on 04/30/20	207	71	71	0	0	0	0	2
[100] † Titanocetus sammarinensis Taxon name last Modified on 04/30/20	125	141	141	0	12	0	0	2
[101] † Tiucetus rosae Taxon name last Modified on 04/30/20	138	139	139	0	1	0	0	2
[102] † Tohoraata raekohao Taxon name last Modified on 04/30/20	161	115	115	0	2	0	0	2
[103] † Tokarahia kauaeroa Taxon name last Modified on 04/30/20	115	149	149	0	14	0	0	2
[104] † Tranatocetus argillarius Taxon name last Modified on 04/30/20	163	107	107	0	8	0	0	2
[105] † Tranatocetus maregermanicum Taxon name last Modified on 04/30/20	156	115	114	0	8	0	0	2
[106] † Uranocetus gramensis Taxon name last Modified on 04/30/20	86	175	175	0	17	0	0	2
[107] † Waharoa ruwhenua Taxon name last Modified on 04/30/20	63	199	197	0	18	0	0	2
[108] † Whakakai waipata Taxon name last Modified on 04/30/20	197	80	80	0	1	0	0	2
[109] † Yamatocetus canaliculatus Taxon name last Modified on 04/30/20	71	192	192	0	15	0	0	2
[110] † ZMT 67 Taxon name last Modified on 04/30/20	124	140	140	0	14	0	0	2

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	140
Matrix downloads	21	Mysticete phylogeny with a new fossil balaenids taxon, Antwerpibalaena liberatlas (21 downloads);
Project downloads	119